vertical structure in coastal waters-fresh water run-off and tidal mixing
TRANSCRIPT
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Crisencio M. Paner, M.Sc.
Ph.D. in Biological Science
(Student)
Vertical structure in coastal waters:freshwater run-off and tidal mixing
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Introduction Coastal waters( which extends from the edge of the
continental shelf to the high water mark) are subjected tothe same seasonal cycles of warming and cooling as the openocean, and in temperate climates the mixed layer mayalternate between being shallow and deep in the same way asin open water.
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Introduction However, the process is greatly complicated by factors
peculiar to the coastal zone. The first of these is the shallowness, which leads to a
situation in which a relatively shallow mixed layer may
extend to the bottom.
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Introduction Since dead biological material, detritus, tends to accumulate
and decompose on the bottom, the nutrients released by itmay be carried to the surface waters and rapidly used inphotosynthesis.
The second important factor is the presence of tidal currentsthat create turbulence in the water.
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Introduction If the depth is not too great in relation to the strength of a
tidal current, tidally induced mixing may extend all the wayto the surface.
The third peculiar feature is the barrier to advection posedby the coastline itself.
For example, if surface water is driven by wind action awayfrom the coast, the only way for it to be replaced is byupwelling from below.
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Introduction Since there is a good chance that the upwelled water has
been enriched in nutrients, an upwelling area is likely to be asite of enhanced biological production.
When stratification first sets in, some phytoplankton is
trapped in a well-lighted mixed layer and production isenhanced.
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Introduction In an area where stratification has been present for a
relatively long time, the nutrients in the mixed layer maybecome depleted and their renewal from below is inhibitedby the pycnocline, so that primary production tends to bedepressed.
Hence, stratification may act in a positive or a negative wayon primary production, but its effects are always important.
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Introduction In the open ocean, stratification is almost always induced by
temperature differences between the layers. In coastal waters a very important additional factor must be
considered- namely, the flow of fresh water from the land.
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Introduction Having salinity close to zero, it is much lighter than sea
water and by lying on top of the sea water creates astratification that can be independent of temperaturedifferences between the layers.
Furthermore this surface layer, being less dense rides higher
at the sea surface, creating a slope along which water flows.
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Introduction
Since these flows depend on the buoyancy, they are knownas buoyancy-driven currents.
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Coriolis Effect Coriolis effect is an inertial force described by the 19th-
century French engineer-mathematician Gustave-GaspardCoriolis in 1835.
The effect of the Coriolis force is an apparent deflection of
the path of an object that moves within a rotating coordinatesystem.
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Coriolis Effect The object does not actually deviate from its path, but it
appears to do so because of the motion of the coordinate
system. The Coriolis effect is most apparent in the path of an object
moving longitudinally.
On the Earth an object that moves along a north-south path,or longitudinal line, will undergo apparent deflection to theright in the Northern Hemisphere and to the left in theSouthern Hemisphere.
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Figure 1: In the inertial frame of reference (upper part of the picture), the black object moves ina straight line, without significant friction with the disc. However, the observer (red dot) who isstanding in the rotating (non-inertial) frame of reference (lower part of the picture) sees theobject as following a curved path due to the Coriolis and centrifugal forces present in this frame.
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Coriolis Effect There are two reasons for this phenomenon: first, the Earth
rotates eastward; and second, the tangential velocity of apoint on the Earth is a function of latitude (the velocity isessentially zero at the poles and it attains a maximum valueat the Equator).
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Coriolis Effect Thus, the Coriolis force figures prominently in studies of the
dynamics of the atmosphere, in which it affects prevailingwinds and the rotation of storms, and in the hydrosphere, inwhich it affects the rotation of the oceanic currents.
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Coriolis Effect and Ocean Currents The influence of the Coriolis force on ocean currents was
first noted by the remarkable observations of Norwegianscientist and explorer Fridtjof Nansen (18611930).
Nansen designed a vessel for the specific purpose of freezing
it into Arctic ice and allowing it to drift with the ice over ayear.
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Coriolis Effect and Ocean Currents In 1893, he positioned his 128-ft wooden ship, Fram, so that
it would freeze into Arctic pack ice 1000 km south of theNorth Pole.
Fram remained locked in the ice for 35 months.
Nansen observed that the direction of ice and ship
movement was not parallel to the wind but typically at anangle of 2040 to the right of it.
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Coriolis Effect and Ocean Currents Nansen related the problem to Vilhelm Bjerknes at the
University of Uppsala, who assigned working out the theoryto his student V. Wilfrid Ekman (1905).
Wind-driven currents deflected by the Coriolis force arenow known as Ekman currents.
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Coriolis Effect and Ocean Currents The Coriolis Effect is the force behind ocean currents and
thus affects marine life in a profound way. Many marine animals rely on currents for the dispersion and
upwelling of nutrients, both of which would be verydifferent without the Coriolis effect.
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Estuaries An estuary is a partly enclosed coastal body of water with one
or more rivers or streams flowing into it, and with a freeconnection to the open sea.
Estuaries form a transition zone between river environments andocean environments and are subject to both marine influences,
such as tides, waves, and the influx of saline water; and riverineinfluences, such as flows of fresh water and sediment.
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Estuaries The inflow of both seawater and freshwater provide high levels of
nutrients in both the water column and sediment, makingestuaries among the most productive natural habitats in theworld.
Most modern-day estuaries were formed during the Holocene
epoch by the flooding of river-eroded or glacially-scoured valleyswhen sea level began to rise about 10,000-12,000 years ago.
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Estuaries Estuaries are typically classified by their geomorphological
features or by water circulation patterns and can be referredto by many different names, such as bays, harbors, lagoons,or inlets, although sometimes these water bodies do notnecessarily meet the above criteria of an estuary and may be
fully saline.
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Estuaries Estuaries are amongst the most heavily populated areas
throughout the world, with about 60% of the worldspopulation living along estuaries and the coast.
As a result, estuaries are suffering degradation by manyfactors, including sedimentation from soil erosion from
deforestation; overgrazing and other poor farming practices;
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Estuaries overfishing; drainage and filling of wetlands; eutrophication
due to excessive nutrients from sewage and animal wastes;pollutants including heavy metals, PCBs, radionuclides andhydrocarbons from sewage inputs; and diking or dammingfor flood control or water diversion.
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Estuaries
River Exe estuary River Nith estuary
Estuary mouth located in Darwin, NorthernTerritory,Australia
http://en.wikipedia.org/wiki/River_Exehttp://en.wikipedia.org/wiki/River_Nithhttp://en.wikipedia.org/wiki/Darwin,_Northern_Territoryhttp://en.wikipedia.org/wiki/Northern_Territoryhttp://en.wikipedia.org/wiki/Northern_Territoryhttp://en.wikipedia.org/wiki/Australiahttp://en.wikipedia.org/wiki/Australiahttp://en.wikipedia.org/wiki/Australiahttp://en.wikipedia.org/wiki/Northern_Territoryhttp://en.wikipedia.org/wiki/Northern_Territoryhttp://en.wikipedia.org/wiki/Darwin,_Northern_Territoryhttp://en.wikipedia.org/wiki/File:Estuary_mouth.jpghttp://en.wikipedia.org/wiki/River_Nithhttp://en.wikipedia.org/wiki/File:River_Nith_estuary.jpghttp://en.wikipedia.org/wiki/River_Exehttp://en.wikipedia.org/wiki/File:Exe_estuary_from_balloon.jpg -
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Classification of Estuaries Based onWater Circulation
In this type of estuary, river output greatly exceeds marine inputand tidal effects have a minor importance.
Fresh water floats on top of the seawater in a layer that gradually
thins as it moves seaward. The denser seawater moves landward along the bottom of the
estuary, forming a wedge-shaped layer that is thinner as itapproaches land.
Salt Wedge
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Classification of Estuaries Based onWater Circulation
As a velocity difference develops between the two layers,shear forces generate internal waves at the interface, mixing
the seawater upward with the freshwater. An example of a salt wedge estuary is the Mississippi River.
Salt Wedge
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Classification of Estuaries Based onWater Circulation
As tidal forcing increases, river output becomes less than themarine input.
Here, current induced turbulence causes mixing of the wholewater column such that salinity varies more longitudinally ratherthan vertically, leading to a moderately stratified condition.
Examples include the Chesapeake BayandNarragansett Bay
Partially mixed
http://en.wikipedia.org/wiki/Chesapeake_Bayhttp://en.wikipedia.org/wiki/Narragansett_Bayhttp://en.wikipedia.org/wiki/Narragansett_Bayhttp://en.wikipedia.org/wiki/Chesapeake_Bay -
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Classification of Estuaries Based onWater Circulation
Tidal mixing forces exceed river output, resulting in a wellmixed water column and the disappearance of the verticalsalinity gradient.
The freshwater-seawater boundary is eliminated due to theintense turbulent mixing and eddy effects.
The lower reaches of the Delaware Bay and the Raritan River inNew Jersey are examples of vertically homogenous estuaries.
Vertically Homogenous
http://en.wikipedia.org/wiki/Delaware_Bayhttp://en.wikipedia.org/wiki/Raritan_Riverhttp://en.wikipedia.org/wiki/New_Jerseyhttp://en.wikipedia.org/wiki/New_Jerseyhttp://en.wikipedia.org/wiki/Raritan_Riverhttp://en.wikipedia.org/wiki/Delaware_Bay -
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Classification of Estuaries Based onWater Circulation
Inverse estuaries occur in dry climates where evaporationgreatly exceeds the inflow of fresh water.
A salinity maximum zone is formed, and both riverine and
oceanic water flow close to the surface towards this zone. This water is pushed downward and spreads along the bottom in
both the seaward and landward direction.
Inverse
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Classification of Estuaries Based onWater Circulation
An example of an inverse estuary is Spencer Gulf, SouthAustralia.
Inverse
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Classification of Estuaries Based onWater Circulation
Estuary type varies dramatically depending on freshwaterinput, and is capable of changing from a wholly marine
embayment to any of the other estuary types.
Intermittent
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Implications for Marine Life Estuaries provide habitats for a large number of organisms and
support very high productivity. Estuaries provide habitats for many fish nurseries, depending
upon their locations in the world, such as salmon and sea trout.
Also, migratory bird populations, such as the black-tailed godwit,
Limosa limosa islandica make essential use of estuaries.
http://en.wikipedia.org/wiki/Salmonhttp://en.wikipedia.org/wiki/Sea_trouthttp://en.wikipedia.org/wiki/Bird_migrationhttp://en.wikipedia.org/w/index.php?title=Limosa_limosa_islandica&action=edit&redlink=1http://en.wikipedia.org/w/index.php?title=Limosa_limosa_islandica&action=edit&redlink=1http://en.wikipedia.org/wiki/Bird_migrationhttp://en.wikipedia.org/wiki/Sea_trouthttp://en.wikipedia.org/wiki/Salmon -
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Implications for Marine Life Two of the main challenges of estuarine life are the variability in
salinity and sedimentation. Many species offish and invertebrates have various methods to
control or conform to the shifts in salt concentrations and aretermed osmoconformers and osmoregulators.
Many animals alsoburrow to avoid predation and to live in themore stable sedimental environment.
http://en.wikipedia.org/wiki/Salinityhttp://en.wikipedia.org/wiki/Sedimentationhttp://en.wikipedia.org/wiki/Fishhttp://en.wikipedia.org/wiki/Invertebratehttp://en.wikipedia.org/wiki/Osmoconformerhttp://en.wikipedia.org/wiki/Osmoregulatorhttp://en.wikipedia.org/wiki/Burrowinghttp://en.wikipedia.org/wiki/Predationhttp://en.wikipedia.org/wiki/Predationhttp://en.wikipedia.org/wiki/Burrowinghttp://en.wikipedia.org/wiki/Osmoregulatorhttp://en.wikipedia.org/wiki/Osmoconformerhttp://en.wikipedia.org/wiki/Invertebratehttp://en.wikipedia.org/wiki/Fishhttp://en.wikipedia.org/wiki/Sedimentationhttp://en.wikipedia.org/wiki/Salinity -
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Implications for Marine Life However, large numbers of bacteria are found within the
sediment which have a very high oxygen demand. This reduces the levels of oxygen within the sediment often
resulting in partially anoxic conditions, which can be furtherexacerbated by limited water flux.
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Implications for Marine Life Plankton are key primary producers in estuaries. They move with
the water bodies and can be flushed in and out with the tides.
Their productivity is largely dependant upon the turbidity of thewater.
The main plankton present are diatoms and dinoflagellates which
are abundant in the sediment.
http://en.wikipedia.org/wiki/Planktonhttp://en.wikipedia.org/wiki/Tidehttp://en.wikipedia.org/wiki/Turbidityhttp://en.wikipedia.org/wiki/Diatomshttp://en.wikipedia.org/wiki/Dinoflagellateshttp://en.wikipedia.org/wiki/Dinoflagellateshttp://en.wikipedia.org/wiki/Diatomshttp://en.wikipedia.org/wiki/Turbidityhttp://en.wikipedia.org/wiki/Tidehttp://en.wikipedia.org/wiki/Plankton -
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Human Impacts Of the 32 largest cities in the world, 22 are located on
estuaries.
For example, New York City is located at the orifice of theHudson River estuary.
As ecosystems, estuaries are under threat from humanactivities such as pollution and overfishing.
http://en.wikipedia.org/wiki/New_York_Cityhttp://en.wikipedia.org/wiki/Hudson_Riverhttp://en.wikipedia.org/wiki/Pollutionhttp://en.wikipedia.org/wiki/Overfishinghttp://en.wikipedia.org/wiki/Overfishinghttp://en.wikipedia.org/wiki/Pollutionhttp://en.wikipedia.org/wiki/Hudson_Riverhttp://en.wikipedia.org/wiki/New_York_City -
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Human Impacts They are also threatened by sewage, coastal settlement, land
clearance and much more. Estuaries are affected by events far upstream, and
concentrate materials such as pollutants and sediments.
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Human Impacts Land run-off and industrial, agricultural, and domestic
waste enter rivers and are discharged into estuaries. Contaminants can be introduced which do not disintegrate
rapidly in the marine environment, such as plastics,pesticides, furans, dioxins, phenols and heavy metals.
http://en.wikipedia.org/wiki/Marine_debrishttp://en.wikipedia.org/wiki/Pesticidehttp://en.wikipedia.org/wiki/Furanhttp://en.wikipedia.org/wiki/Polychlorinated_dibenzodioxinshttp://en.wikipedia.org/wiki/Phenolhttp://en.wikipedia.org/wiki/Heavy_metal_musichttp://en.wikipedia.org/wiki/Heavy_metal_musichttp://en.wikipedia.org/wiki/Phenolhttp://en.wikipedia.org/wiki/Polychlorinated_dibenzodioxinshttp://en.wikipedia.org/wiki/Furanhttp://en.wikipedia.org/wiki/Pesticidehttp://en.wikipedia.org/wiki/Marine_debris -
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Human Impacts Such toxins can accumulate in the tissues of many species of
aquatic life in a process calledbioaccumulation. They also accumulate inbenthic environments, such as
estuaries and bay muds: a geological record of humanactivities of the last century.
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Human Impacts For example, Chinese and Russian industrial pollution, such
as phenols and heavy metals, in the Amur River havedevastated fish stocks and damaged its estuary soil.
Estuaries tend to be naturally eutrophic because land runoffdischarges nutrients into estuaries.
http://en.wikipedia.org/wiki/Amur_Riverhttp://en.wikipedia.org/wiki/Eutrophichttp://en.wikipedia.org/wiki/Land_runoffhttp://en.wikipedia.org/wiki/Land_runoffhttp://en.wikipedia.org/wiki/Eutrophichttp://en.wikipedia.org/wiki/Amur_River -
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Human Impacts With human activities, land run-off also now includes the many
chemicals used as fertilizers in agriculture as well as waste fromlivestock and humans.
Excess oxygen depleting chemicals in the water can lead tohypoxia and the creation ofdead zones.
It can result in reductions in water quality, fish, and other animalpopulations.
http://en.wikipedia.org/wiki/Hypoxia_(environmental)http://en.wikipedia.org/wiki/Dead_zone_(ecology)http://en.wikipedia.org/wiki/Dead_zone_(ecology)http://en.wikipedia.org/wiki/Hypoxia_(environmental) -
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Human Impacts Overfishing also occurs. Chesapeake Bay once had a
flourishing oyster population which has been almost wipedout by overfishing.
Historically the oysters filtered the estuary's entire watervolume of excess nutrients every three or four days.
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Human Impacts Today that process takes almost a year, and sediment,
nutrients, and algae can cause problems in local waters.Oysters filter these pollutants, and either eat them or shapethem into small packets that are deposited on the bottomwhere they are harmless.
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Tidal mixing and phytoplanktonproduction
The idea that the turbulence generated by high tidalcurrents keeps the water over some shallow regions mixedall year while the quieter regions in the deeper watersbecome stratified in summer was first advanced more than70 years ago.
The lack of stratification would be expected to decreasephytoplankton productivity.
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Tidal mixing and phytoplanktonproduction
On the other hand, dead organic matter is continuallydecomposing on and in the sediments, providing a source ofnutrients that can be mixed back into the water column tostimulate phytoplankton production.
In the open ocean initiation of the spring growth in thezooplankton population is often triggered by the upwardmigration of a large biomass of adult and late-stage copepods thathave spent the winter at depths greater than 350 m.
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Tidal mixing and phytoplanktonproduction
Reproduction begins as soon as this population reaches surfacewaters, and grazing pressure on the phytoplankton developsrelatively early in the season.
In part this process is made possible by the rapid warming of the
surface waters after stratification.
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Tidal mixing and phytoplanktonproduction
In tidally mixed water, warming is delayed by the lack ofstratification, and there is no population from deep waterthat can ascend to the euphotic zone and begin to feed onthe phytoplankton.
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Introduction to marine biologyKarleskint, George.
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Castro, Peter.New York : McGraw-Hill Higher Education, c2007.
Introduction to marine biologyKarleskint, George.Southbank, Victoria ; Belmont, CA : Thomson, c2006.
Marine biology : an ecological approach
Nybakken, James Willard.San Francisco : Pearson Benjamin Cummings, c2005.
The Marine biology coloring bookNiesen, Thomas M.New York : Barnes & Noble, c1982.
Chronobiology of marine organismsNaylor, E. (Ernest), 1931-Cambridge ; New York : Cambridge University Press, 2010.
Laboratory and field investigations in marine lifeSumich, James L.Sudnury, MA : Jones and Bartlett, c2009.
Marine science : an illustrated guide to scienceDiagram Group.New York : Chelsea House Publishers, 2006.
The SeaAlderton, David.United Kingdom : Ladybird discovery, 1995.
BACKGROUND OF PROF. CRISENCIO M. PANER
http://ustlib.ust.edu.ph/search~S2?/Xdynamics+marine&searchscope=2&SORT=DZ/Xdynamics+marine&searchscope=2&SORT=DZ&extended=0&SUBKEY=dynamics%20marine/1,5,5,B/frameset&FF=Xdynamics+marine&searchscope=2&SORT=DZ&1,1,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&1,1,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&1,1,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&2,2,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&3,3,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&5,5,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&8,8,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&13,13,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&15,15,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&20,20,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&24,24,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&24,24,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&20,20,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&15,15,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&13,13,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&13,13,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/13,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&13,13,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&8,8,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&5,5,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&3,3,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&2,2,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&1,1,http://ustlib.ust.edu.ph/search~S2?/Xmarine+biology&SORT=D&searchscope=2/Xmarine+biology&SORT=D&searchscope=2&SUBKEY=marine%20biology/1,45,45,B/frameset&FF=Xmarine+biology&SORT=D&searchscope=2&1,1,http://ustlib.ust.edu.ph/search~S2?/Xdynamics+marine&searchscope=2&SORT=DZ/Xdynamics+marine&searchscope=2&SORT=DZ&extended=0&SUBKEY=dynamics%20marine/1,5,5,B/frameset&FF=Xdynamics+marine&searchscope=2&SORT=DZ&1,1,http://ustlib.ust.edu.ph/search~S2?/Xdynamics+marine&searchscope=2&SORT=DZ/Xdynamics+marine&searchscope=2&SORT=DZ&extended=0&SUBKEY=dynamics%20marine/1,5,5,B/frameset&FF=Xdynamics+marine&searchscope=2&SORT=DZ&1,1,http://ustlib.ust.edu.ph/search~S2?/Xdynamics+marine&searchscope=2&SORT=DZ/Xdynamics+marine&searchscope=2&SORT=DZ&extended=0&SUBKEY=dynamics%20marine/1,5,5,B/frameset&FF=Xdynamics+marine&searchscope=2&SORT=DZ&1,1, 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Ph.D. in Biological Science (Candidate), UST
M.S. in Microbiology, UST
B.S. Biochemistry, UST
Italian Government ScholarCollege Scholar
Certificate in Education
10th Placer Licensure Exams for Teachers (1998)
20 years of experience as a teacher (College, High School, Elementary)
Expert in Internet, Computer (Software, Hardware, and Repair)
Researcher and BloggerArt Restorer/Conservator
Publication:Prime Journal of Microbiology Researchhttp://primejournal.org/PJMR/abstract/2012/may/Paner.htm
http://primejournal.org/PJMR/cont/2012/may.htm
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http://primejournal.org/PJMR/abstract/2012/may/Paner.htmhttp://primejournal.org/PJMR/cont/2012/may.htmhttp://allaboutweightmanagement.blogspot.com/http://thepregnancyplanner.blogspot.com/http://cmpaner.blogspot.com/http://internet-moneymakingsecrets.blogspot.com/http://internet-moneymakingsecrets.blogspot.com/http://internet-moneymakingsecrets.blogspot.com/http://internet-moneymakingsecrets.blogspot.com/http://cmpaner.blogspot.com/http://thepregnancyplanner.blogspot.com/http://allaboutweightmanagement.blogspot.com/http://primejournal.org/PJMR/cont/2012/may.htmhttp://primejournal.org/PJMR/abstract/2012/may/Paner.htm