ISSUE 16 . MICHAELMAS 2017

fusionTHE NEWSLETTER OF THE SIR WILLIAM DUNN SCHOOL OF PATHOLOGY

2 / FUSION . MICHAELMAS 2017

ContentsEditorial. Matthew Freeman ………2

NEWS

Fusion is 15 years old! …………3

Susan Lea elected to the Academy ofMedical Sciences …………………3

Corrigendum ……………………3

Jeremy Farrar delivers the 2016Norman Heatley lecture …………4

And the Winner is … Antibiotics!…4

Interview with Simon Shelly ………5

A year in the life of the GraduateStudents Association. The GSACommittee ………………………7

BOOK REVIEW

Penicillin and the Legacy of NormanHeatley by David Cranston and EricSidebottom. Annemieke Kok ……8

Innovation in Medical Teaching:Harnessing Technology for theTeaching of Anatomy. Chris Horton

…………………………………9

SPOTLIGHT

Transcription and DNA Damage: Friendor Foe? Monika Gullerova ………11

RESEARCH FEATURE

Quality control of membrane proteins.Pedro Carvalho …………………12

Editorial2016 was a special year for the Dunn School. It was the 75th anniversary of thefirst trial of penicillin in humans, and we celebrated accordingly. But 2017 is alsospecial, as it marks the 90th anniversary of the opening of the Dunn School. TheOxford Journal Illustrated, dated March 16th 1927, headlined Oxford's new home ofresearch, modernised by a generous gift of £100,000. It described how “so convincedwere the trustees that the surest and safest road to knowledge lay in research” thatthey had gone to the Court of Chancery to amend the terms of Sir William Dunn’s willto support a research department, instead of making a gift that was more directlymedical. Their aim was a “continual and sustained supply of men and women whoseminds lend themselves to the stress and strain of research work – resourceful, keen,and adaptable minds”. I like to think that, 90 years later, the trustees would feelthat their gift had fulfilled both of those goals. Top men and women scientistsresourcefully researching major biomedical challenges and, with antibiotics, thebonus of having developed a treatment that has saved hundreds of millions of lives:one of the greatest examples ever of the value of philanthropy.

We celebrate the past but must put our effortinto the future by focusing on what’s next.Recent recruitments highlight our continuingability to attract the very best. Pedro Carvalhois the new EPA Professor of Cell Biology. Hejoined us this year from one of Spain's topresearch institutes in Barcelona, where is hasbeen a rising star in European cell biology. Hisfocus is on quality control of proteins in cells,and the molecular mechanisms that regulatehow the identity of cellular organelles isestablished and maintained, a programme ofresearch he outlines in detail in this edition ofFusion. This is the kind of fundamental biologythat underpins many disease mechanisms andit epitomises how pathology in the 21st centurydepends on molecular and cell biology. Wehave also just recruited Monika Gullerova as anAssociate Professor in cancer biology. Monikahas been at the Dunn School as anMRC-funded Career Development Fellow forthe last four years and was appointed from astrong field of applicants from around theworld. Her research, likewise described later inthis edition, focuses on a new mechanism ofrepairing damaged DNA, which she discovered.Again, a fundamental piece of science of clearand direct relevance to disease. We are very

grateful to the trustees of the EPA ResearchFund and the Lee Placito Memorial Fund fortheir generosity in endowing these two posts.

As non-British Europeans, both Pedro andMonika also illustrate a significant concern thatwe must tackle over the next few years. Ofcourse, I speak of Brexit. While acknowledgingthat my column in Fusion is perhaps not theplace for a politically-charged polemic, I amunable to hide my own personal frustrationand disappointment about the outcome of theEU referendum, and my concern about itspotential effect on British science. But our jobnow is to react in a way that ensures that theDunn School can continue to be a trulyworld-leading research and teachingdepartment. We have, of course, been afully-international department for 90 years andthat will not change. Long before the EU wasdreamt of, the Dunn School was recruiting thebest from around the world and supporting thebest science. Indeed, the first three heads ofthe Dunn School were not British (Dreyer,Florey and Harris), nor was Chain, nor many ofthe other scientists whose photographs adornthe walls of the department – a lovely traditionthat we maintain to this day. If you take a lookat the homepage of the Dunn School’s websiteyou will see a ringing endorsement of ourdetermination to remain welcoming to all, fromwherever they come, and whatever the currentpolitical situation throws at us. The excellentrecruitments mentioned above exemplify myconfidence that we will succeed: by the time ofour centenary, I predict we will be asinternational as we are now, with equallyoutstanding science.

Front cover image:We are grateful to Simon Shelley for allowingus to feature a piece of his artwork on the frontcover. Please see the interview with Simon forfurther details.

Let me finish by saying, as I do each year,how much we value staying in touch withour former members and friends. Wegreatly appreciate hearing your news andmeeting you when you visit, and areespecially grateful for the support that theextended Dunn School family provides inso many ways. Please do stay in touchand keep an eye on the news pages of thewebsite, where we post stories about awide range of Dunn School events andachievements. In the meantime, we verymuch hope you will enjoy reading aboutthe many aspects of life within thedepartment, celebrated in the pages ofthis year’s edition of Fusion.

FUSION . MICHAELMAS 2017 / 3

The inaugural issue of Fusion was published inMichaelmas term 2002, an initiative firstintroduced by Herman Waldmann and edited bySusan Harrison, the Dunn School’s DevelopmentOfficer at the time. The first edition includeddetails of George Brownlee’scontribution to thetreatment ofhaemophilia B, aninterview withnewly-appointed groupleader, Keith Gull, anddetails of the completionof the EPA Building a yearprior to its official openingby Nobel Prize-winner Dr TimHunt. Since its modest beginnings, Fusion hasbeen published annually and distributed tomany hundreds of friends and alumni across 27countries, from Sri Lanka to Argentina and fromLuxembourg to Taiwan, but has also evolvedwith the times: full colour printing wasintroduced in 2009, a dedicated front cover wasadded in 2011 and the magazine is now madeavailable for download as a pdf file from thedepartmental website.

Over the years, Fusion has documented thearrival of many group leaders who continue to

work at the Dunn School including ChrisNorbury, Quentin Sattentau, Susan Lea andOreste Acuto as well as more recent recruits, ofwhich Pedro Carvahlo’s appointment is

showcased in this issue. It has alsocelebrated the work of

many key peoplewho, upon

retirement, haveleft a significant

scientific legacy tothe Dunn School

including GordonMacPherson, Siamon

Gordon, Neil Barclay,George Brownlee, Simon

Hunt and previous Head of Department,Herman Waldmann. Furthermore, it has servedas a forum to explore and celebrate the richhistory of the department and the colourfulcharacters that have worked here over theyears. Most importantly, however, the editorsfrequently receive letters from readers sharingtheir reminiscences of the years they spent inthe Department which invariably occupies aparticularly fond place in the hearts andmemories of many who worked here. We lookforward to receiving many more such lettersover the next 15 years!

NewsFusion is 15 years old!

On 28th June2017, Susan Lea,Professor ofMicrobiology at the

Dunn School, was formally admitted as a Fellowof the Academy of Medical Sciences inrecognition of her exceptional contributions to thefield of infection and immunity. The Academy isan independent body in the UK with a mission toadvance biomedical and health research and tofacilitate its translation into the clinic. This year,46 new Fellows were elected, of which 15 werewomen, the highest proportion so far elected bythe Academy.

Susan’s laboratory is renowned for its work onbiological structures with specific medicalsignificance in two important areas: thecomplement system and bacterial secretionsystems. Her recent research has focussed on theactivation and regulation of the alternative

pathway of complement by both membranebound and soluble proteins. Studying the hijackof complement components by microbialvirulence factors for both complement evasionand cell entry may influence the design ofantigens for new vaccines based on theseproteins, work that is now being developed withNovartis. Furthermore, her group has made majoradvances in defining the architecture of bacterialsecretion systems, including the pathogenic typeIII secretion system of Shigella flexneri and themajor integral membrane component of the TATsystem. Her election to the Academy of MedicalSciences formally recognises the quality of hercontribution to these important areas of medicalscience.

Susan Lea Elected to the Academy of Medical Sciences

FOCUS ON PATHOGENS

Influenza and Host Transcription: ALove-Hate Relationship. DavidBauer …………………………14

Host Pathogen Interactions: A CircularArgument. Jessica Martyn andChristoph Tang …………………16

Leishmania – The Shape of a Killer.Jack Sunter and Keith Gull ………18

Gene Therapy against HIV: Fightingthe Virus in Disguise. BernadetaDadonaite………………………19

TECHNOLOGY FEATURE

The New Ice Age: Cryo-ElectronMicroscopy Comes to South ParksRoad. Errin Johnson ……………22

Winning Entries of the ImagingCompetition ……………………24

HISTORY CORNER

The Dunn School archive at theBodleian library. CharlotteMcKillop-Mash…………………25

The First Systemic Use of Penicillin: A Personal Account by CharlesFletcher…………………………26

The curious tale of cows, quarantinedislands and Saddam’s WMD. EricSidebottom ……………………28

fusionT H E N E W S L E T T E R O F T H E S I R W I L L I A M D U N N S C H O O L O F P A T H O L O G Y

Welcome

to the f

irst edit

ion of fus

ion! This

is both

an exciti

ng and a

challeng

ing time

for

the Sir W

illiam Dun

n School

of Patho

logy. The

past few

years ha

ve seen

much bu

ilding

work bot

h on the

new EPA

building

and on t

he Medic

al Scienc

es Teachi

ng Block

, situated

behind t

he Dunn

School an

d access

ed by a w

alkway fr

om the S

chool.

This physical expansion of the department is

necessary to enable the Dunn School to maintain

its international leadership in research on

mechanisms and treatments for disease in the face

of increasing competition for research funding.

In this country the most co

mmon causes of

morbidity and mortality are related to cancer

and to cardiovascular and immunological

diseases. The historic development of antibiotics

in this department has, over the past 50 years,

reduced the consequences of bacterial infection,

although microbial resistance to the current

generation of antibiotics will m

ean that unless

new drugs are found bacterial diseases will

again constitute a major problem. Outside the

western world parasitic diseases and AIDS

constitute an immense problem.

The challenge for the Department is to maintain

strong scientific

excellence while positioning

itself to improve health in those major disease

areas. In the areas of immunology and chemical

pathology we have some of the strongest

scientific groupings in the UK, soon to be

strengthened even further by a recently

endowed Chair in molecular biology. We now

wish to expand our strong foundations in other

areas of pathology, in particu

lar in the areas of

molecular microbiology and in cell and cancer

biology. To that end, the Department is

fortunate to have been able to create a Chair

and a lectureship in microbiology. A further

lectureship is currently being fille

d. We are

delighted to welcome Professor Keith Gull, a

world authority on protozoal diseases, to

provide leadership in molecular microbiology. In

addition, new research fellowships have allowed

diversification of our effort in

bacterial diseases.

In the areas of cell and cancer biology recent

appointments to a Chair and lectureship in cell

biology will provide that added impetus. Our

next goal is to raise funds for a Chair in cancer

cell biology to continue the Department’s long-

standing expertise in that area. The area of

cardiovascular disease is evolving around the

crucial role of macrophages in determining lipid

accumulation in blood vessels, and clearly

remains a priority for the Dunn School.

The successful completion of the new E P

Abraham building and the overwhelming

endorsement of the Department’s research with

the award of a 5* rating in the recent RAE

exercise provide both opportunities and

challenges. Chief among the opportunities is our

ability to attr

act the best sc

ientists and their

groups to the Dunn School. Among the

challenges in the current financial clim

ate is the

need to find new resources to enable D. Phil

students and young, independent post-doctoral

research fellows – the scientists

of the future –

to study here, while also maintaining the services

and fabric of the Department as a whole. The

year ahead will see the start of a fundraising

campaign to generate new income to support our

goals in research and infrastructure, and I hope

you will lend your support to this new effort.

Herman W

aldmann

EditorialV O L U M E 1 . I S S U E 1 . M I C H A E L M A S 2 0 0 2

Contents

Editorial

Challenges ahead... 1

From clone to market

place

How haemophilia B

patients benefit...2

Introducing

Keith Gull

3

The EPA building at

the Dunn School4

When cell biology

met immunology6

Gunther Blobel gives

Heatley Lecture

7

Appointments and

distinctions

7

Prizes and awards 7

Forthcoming events 7

Fighting AIDS with

books

8

Herman Waldmann, FRS,

Head of Department

CCoConContConteContenContentContentsContents

Editorial: Herman Waldmann . . .1

Introducing Jordan Raff . . . . .2

Life and Trypanosomes –Catarina Gadelha . . . . . . . . . . .4

The Dunn School: Out ofAfrica – Keith Gull . . . . . . . . . . . .6

Tales of the Unexpected: TheTwists and Turns of a Life inScience – George Brownlee 8

The Foibles of Flow Cytometry –An Interview with Nigel Rust .11

Reminiscences – AlvinVolkman . . . . . . . . . . . . . . . . . . . . .13

News, Honours, Prizes . . . . .15

Champions: A History of theDunn School Cricket Club . . .16

Cultural Transmission: HowInformation about PenicillinSpread throughout Europeduring World War II – GilbertShama . . . . . . . . . . . . . . . . . . . . . .18

The Oxford MolecularPathology Institute (OMPI) –William James . . . . . . . . . . . . . .20

ISSUE 8 . MICHAELMAS 2009

fusionTHE NEWSLETTER OF THE SIR WILLIAM DUNN SCHOOL OF PATHOLOGY

Our hope is that the new space created will notonly enable us to fill our vacant posts with scientists at the highest level, but will also allowus to harbour more joint appointments with theClinical School. In the past 15 years the DunnSchool has undertaken a very high level ofrebuilding and refurbishment activity, all aimedat maintaining the Department as a world-leader in biomedical research. Our colleagueshave been remarkably patient and accepting ofthese intrusions, and I am pleased to say that

the morale and spirit of the School is as highas ever.

It is very satisfying for us that so many alumnihave taken pleasure from receiving past issues ofFusion, and we hope that we can continue to keepyou updated about our aspirations, developmentsand achievements as we meet the building andrecruitment challenges of the next three years.

HHeHerHermHermaHermanHerman Herman WaHerman WHerman WalHerman WaldHerman WaldmHerman WaldmaHerman WaldmanHerman WaldmannHerman Waldmann

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On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has jIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joiIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joinIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joineIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joinedIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined aIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined asIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as tIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as theIt has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as then

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thene

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew M

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Mi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Mil

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Mils

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milst

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milste

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstei

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein C

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Ch

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Cha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chai

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair o

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of C

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Ca

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Can

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Canc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cance

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer M

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Mo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Mol

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Mole

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molec

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecu

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecul

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecula

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular B

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Bi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Bio

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biol

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biolo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biolog

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, an

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and w

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and wo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and wor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work h

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work ha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has b

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has be

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has bee

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been i

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been in

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been ini

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been init

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initia

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiat

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiate

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to rep

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to repl

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to repla

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replac

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace thef

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace thefo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace thefor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theform

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theforme

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer L

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Le

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Les

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Lesl

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Lesli

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie M

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Ma

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Mar

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Mart

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Marti

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin b

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin bu

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin bui

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin buil

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin build

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin buildi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin buildin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building w

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building wi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building wit

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with n

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with ne

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new l

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new la

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new lab

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new labo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new labor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new labora

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laborat

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laborato

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laborator

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratori

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratorie

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to b

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be k

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be kn

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be kno

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be know

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the O

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Ox

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxf

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxfo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxfor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford M

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Mo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Mol

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Mole

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Molec

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Molecu

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Molecul

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Molecula

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford Molecular

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularP

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPa

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPat

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPath

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPatho

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathol

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPatholo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPatholog

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology I

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology In

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Ins

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Inst

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Insti

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Instit

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institu

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institut

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (O

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OM

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMP

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI)

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI).

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI).

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). W

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We h

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We ha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We hav

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have al

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have als

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also rec

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also rece

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recen

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recent

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recentl

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently w

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently we

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently wel

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welco

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcom

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcome

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed P

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Pr

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Pro

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof B

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Ba

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bas

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass H

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Ha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Has

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hass

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassa

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan w

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan wh

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who h

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who ha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has rel

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has reloc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has reloca

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocat

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocate

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocated

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedh

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis l

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis la

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis lab

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis labo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis labor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis labora

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laborat

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laborato

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laborator

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratori

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratorie

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the D

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Du

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dun

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn S

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn Sc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn Sch

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn Scho

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn Schoo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, s

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so h

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so he

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so hel

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so help

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helpi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helpin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping e

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping es

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping est

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping esta

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping estab

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establ

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establis

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a c

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr i

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr it

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr iti

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itic

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itica

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical m

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical ma

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mas

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass i

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in c

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in ca

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in can

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in canc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cance

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer res

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer rese

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer resea

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer resear

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer research

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchw

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwit

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwith

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the D

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the De

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Dep

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Depa

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Depar

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Depart

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Departm

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Departme

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Departmen

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department.

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department.

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. O

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the s

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sa

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad s

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad si

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad sid

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, thr

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, thre

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three o

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of o

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of ou

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our m

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our mo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our mos

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most s

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most se

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most sen

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most seni

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senio

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior c

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior co

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior col

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior coll

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colle

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior collea

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleag

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagu

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleague

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (S

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Si

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Sia

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siam

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon G

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Go

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gord

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,G

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,Ge

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,Geo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,Geor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,Georg

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George B

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Br

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Bro

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brow

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brown

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownl

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownle

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee an

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and G

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Go

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gor

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gord

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordo

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon M

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Ma

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Mac

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macp

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macph

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macphe

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpher

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macphers

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherso

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson)

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson)

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) h

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) ha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) hav

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have ret

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have reti

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retir

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retire

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired,

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, l

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, le

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, lea

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leav

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leavi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leavin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a m

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a ma

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a mas

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a mass

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massiv

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive h

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive ho

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hol

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole i

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the res

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the rese

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the resea

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the resear

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researc

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the research

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researcha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchan

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand te

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand tea

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teac

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teach

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teachi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teachin

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching l

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching li

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching lif

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life o

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the d

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the de

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the dep

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the depa

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the depar

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the depart

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the departm

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the departme

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the departmen

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department.

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department.

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. T

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. Th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The c

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The ch

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The cha

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chai

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chair

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs v

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs va

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vac

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vaca

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacat

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacate

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated b

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by thes

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these r

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these re

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these ret

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these reti

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retir

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retire

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirem

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retireme

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retiremen

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirement

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements c

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements ca

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements can

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cann

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements canno

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot b

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be f

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be fi

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be fil

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be fill

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be fille

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filled

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filledu

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filledun

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filledunt

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filledunti

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil O

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OM

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMP

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI i

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is c

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is co

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is com

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is comp

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is compl

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is comple

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is complet

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is complete

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed a

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at t

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at th

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the e

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the en

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end o

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of 2

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of 20

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of 201

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of 2010

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of 2010.

It has been a dramatic year in the life of the Dunn School. On the good side, the Departmentscored first nat ionally in the recent research assessment exercise, Jordan Raff has joined as thenew Milstein Chair of Cancer Molecular Biology, and work has been initiated to replace theformer Leslie Martin building with new laboratories to be known as the Oxford MolecularPathology Institute (OMPI). We have also recently welcomed Prof Bass Hassan who has relocatedhis laboratories to the Dunn School, so helping establish a cr itical mass in cancer researchwithin the Department. On the sad side, three of our most senior colleagues (Siamon Gordon,George Brownlee and Gordon Macpherson) have retired, leaving a massive hole in the researchand teaching life of the department. The chairs vacated by these retirements cannot be filleduntil OMPI is completed at the end of 2010.

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ISSUE 11 . MICHAELMAS 2012

fusionTHE NEWSLETTER OF THE SIR WILLIAM DUNN SCHOOL OF PATHOLOGY

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Corrigendum

In the last edition of Fusion, we reportedhow Elizabeth Robertson had beenawarded the Royal Medal for her manycontributions to developmental biologyand was one of a number of distinguishedmembers of the Dunn School over theyears who had likewise been honoured fortheir contributions to science. One namethat was inadvertently omitted from thelist of those who had received the RoyalMedal was that of James Gowans, whosepioneering work on lymphocyterecirculation was recognised in 1976.

The Editors apologise for this oversight.

4 / FUSION . MICHAELMAS 2017

And the Winner is … Antibiotics!

On 15th June 2016, BBC2 broadcastBritain’s Greatest Invention, a live eventfrom an aircraft hangar near Swindon, whichserves as a storage facility for some of theScience Museum’s largest exhibits.

A number of well-known celebritieschampioned a wide range of inventionsfrom the jet engine to concrete and fromtelevision to the humble fridge. Butjournalist and former news reader, Angela

Rippon was on a personal mission:owing her very survival as a childto the use of antibiotics, she madean impassioned case in favour ofpenicillin. In support of herarguments, she visited the DunnSchool to interview thedepartment’s archivist, EricSidebottom, and reported on her

findings from the iconic bridge overlookingthe Bridge Café. Her estimate that 200million lives have been saved by antibioticssince the advent of penicillin appeared tostrike a chord with the general public whovoted overwhelmingly that antibioticsshould be considered Britain’s greatestinvention!

Jeremy Farrar Delivers the 2016 Norman Heatley Lecture

Against the impressive backdrop of thenewly-refurbished Weston Library, the 2016Norman Heatley lecture was delivered byProfessor Jeremy Farrar OBE, FRS, Director ofthe Wellcome Trust, one of the world’slargest and most influential medicalresearch charities. The occasion provided anopportunity to celebrate the 75th anniversaryof the first systemic administration ofpenicillin in man which heralded theantibiotic era. The event drew an audienceof several hundred which included manycurrent members of the Dunn School as wellas numerous people with personal links tothe penicillin story. These included membersof the Heatley family who continue tosupport the annual lecture founded in 1992to commemorate the vital contribution madeby Norman Heatley to the purification ofpenicillin that made its subsequent clinicaluse possible. The lecture was also attendedby Mark Fletcher, son of Charles Fletcher,

the physician responsible fortreating the first patient withpenicillin and who subsequentlywent on to become the nation’sfirst TV doctor, responsible forbringing medical education to manyof the British public.

Jeremy Farrar proved a very fittingchoice of speaker for such asignificant occasion since he has

previously served as Professor of TropicalMedicine and Global Health at Oxford andDirector of the Oxford University ClinicalResearch Unit in Vietnam. With along-standing interest in the control ofinfectious disease, Professor Farrar spoke ata time when outbreaks of MERS werethreatening Asia, a pandemic of influenzavirus was top of the government’s riskregister and the WHO had recently declaredthe antibiotic era to be over and drugresistance to be one of the greatest threatsto the future of humanhealth. He outlined howdrug resistance is evident notonly among commonbacterial pathogens such asStaphylococcus but alsoamong the malaria parasitePlasmodium which hasrecently rendered theartemisinin derivatives largely

obsolete in some parts of the world. Usingthe recent outbreak of Ebola in Sierra Leonein 2014 as a sobering example, ProfessorFarrar made the case that interconnectivityof travel across the globe poses anunprecedented threat to human health:“What happens in Freetown today”, hecommented, “affects us here in Oxfordtomorrow.” Drawing an important parallelwith the development of penicillin, ProfessorFarrar commented that clinical aspects ofdrug development commonly take 10-20years before approval is granted: in contrastpenicillin made the journey from mouse toman in less than a year. In a call to arms totoday’s medical researchers, Professor Farrarargued that new paradigms are requiredthat work in parallel on different aspects ofdrug discovery in order to make more rapidadvances to meet future threats frominfectious microorganisms. The lecture was followed by a celebratorydinner at Exeter College.

FUSION . MICHAELMAS 2017 / 5

What is your connection with the Dunn School?I was a DPhil student on the top floor in the Leslie Martin Building, or‘new building’ as it was more commonly called, of which GeorgeBrownlee was the head. My supervisor was Francisco Ernesto (Tito)Baralle. I started as a research technician in Tito’s lab in 1982 andwas subsequently a DPhil student from 1983-86, based at MagdalenCollege.

How has your career developed since leaving the Dunn School?When I left the Dunn School in October 1986, my first postdocposition was at Children's Hospital, part of Harvard Medical School, inBoston, Massachusetts. I then did additional postdoc stints at theHarvard Medical School institutions of the Center for Blood Researchand Massachusetts General Hospital (MGH). In 1991 I was appointedInstructor in Medicine and in 1995 Assistant Professor of Medicine atHarvard Medical School. While at MGH, I held sabbaticalappointments at the Institut d’Hématologie, Hôpital Saint-Louis, inParis and the Institut de Recherches sur le Cancer de Lille in France.In 2008 I moved to La Crosse, Wisconsin to take up the position ofDirector of the Kabara Cancer Research Institute. During this time, Iwas appointed faculty at Viterbo University and the University ofWisconsin, La Crosse. In 2014 I was appointed Adjunct ClinicalProfessor of Medicine at the University of Wisconsin School ofMedicine and Public Health, but in 2015 I moved back to Boston tofound the biotech firm Leukemia Therapeutics, LLC. In addition tobeing President and CEO of this company, I continue to be involved inacademic collaborations with researchers in Britain, France, Italy,Germany, Japan, the USA, Australia, and Nigeria. I am also activelyinvolved in the Cancer Genome Atlas (TCGA) program of the USNational Cancer Institute.

What are your recollections of the time you spent in thedepartment?It was probably the best time I have had as a scientist! Great people,extremely supportive, motivated, engaged, inspiring, creative,hard-working: you knew what you, and those around you, were doingwas world-class and cutting edge. I have so many recollections of the

years I spent at the Dunn School that it is difficult to single out anyone in particular, however, the Dunn School was the place that I firstdrank coffee. My breaks in the coffee room felt a little short with justa can of orange juice which could be downed in a couple of seconds.The answer was, of course, the free instant coffee which was hot andtook longer to drink. I hated the taste and added five large spoonsof sugar for masking purposes. But then after a few weeks with toomuch coffee and far too much sugar and my hands starting totremble, I was forced to go cold-turkey on the sugar, which I havemaintained until this day.

I well recall taking two weeks to synthesize and purify a singleoligonucleotide (the normal time frame in those days!), pickinghundreds of colonies in triplicate for a cloning screen while watchingthe test match on a small black and white TV, and melting glassmicropipettes over a Bunsen burner and pulling them to make a finehollow strand capable of loading an acrylamide gel. Tito’s officeprovided the best light for holding up x-ray film at an angle to thewindow and at arm’s length to try to imagine a signal!And then there was Kris Jenner, Marshall Scholar, lab mate and friendconfessing over a farewell pint in the King’s Arms that for the first sixmonths he knew me he just smiled and nodded when we spoke, sincehe couldn’t understand a word I said through my Yorkshire accent!

When did you first develop an interest in art?My mother is an artist: a sculptor and potter and an art teacher. Shewent to Goldsmiths, University of London and Batley School of Artand was heavily influenced by the arts-and-crafts movement, thepotters, Bernard Leach and Shōji Hamada, and the sculptors BarbaraHepworth and Henry Moore. My father is an amateur artist.Consequently, art supplies were always around the house.Furthermore, my mother took me out with her students on ‘plein-air’drawing expeditions and made sure I was exposed to art from a veryearly age, from watching art shows on the TV (eg Kenneth Clark’sCivilization), to visiting art museums both in my home town(Huddersfield) and on trips to London. On these expeditions, Irealized I liked the shared visceral passion of Medieval and Modernart. Both were making strident statements about existence that Ithought was often lacking in the centuries between. Being anadolescent during the Punk era also helped! I loved the later worksof Turner and considered him, rather than Cezanne and the FrenchImpressionists, as the father of modern art.

How would you describe the art form you have developed?The term ‘outsider art’ was coined when I was a boy and really spoketo me as someone with a complex ethnic background. My father wasborn and raised a Muslim in India: his father an Indian Muslim, hismother a German Jew. His mother’s father won the Iron Cross in theFirst World War: he wore his medal in public next to the Star of David

Following the publication of an article about Art at the Dunn School in last year’s edition of Fusion, theEditors received an email from alumnus Simon Shelley who, since leaving Oxford, has pursued parallelcareers in art and science. Intrigued, Fusion dispatched a reporter to find out more…

Interview with Simon Shelley

but ‘disappeared’ during WWII. Our family narrative was, therefore,of a Muslim in Hindu India with the traumas of Partition and of a Jewin Hitler’s Germany.

My childhood was the product of a mixed marriage in working classYorkshire in the 1960’s which led me to witness violent school-yardracism and my father being carried off a cricket field, the onlymember of the factory team targeted by body-line fast bowling. Theseexperiences made me intrinsically distrustful of the mass and themob: my brother and I were always the outsiders. And then therewas Punk and the Pistols and New Wave and Joy Division and FrancisBacon and Mark Rothko and Bosch and the Flemish Primitives andsuddenly I felt at home!

So a good short-hand for my work is ‘outsider art’. As far astechnique is concerned, it could be described as sculptural or 3Ddecoupage, mixed media using found materials such as newspapers,magazines, catalogues, scientific journals, advertising, clothing andtoys, acrylic artist paint, household paint, automotive spray paint, soiland wood stain.

Where do you find inspiration for your artwork?I find inspiration in the chaotic and ephemeral nature of life aroundus every day: the daily struggle, banal as it often seems, but whichembodies the nobility of striving to do better, to know more, to bewiser.

Art done well is the other side of the same coin as science done well.Both require imagination, hard work and the production, exhibitionand publication of something new, something novel. Both art andscience build on the past to press forward, groping in the darkness tonew knowledge and insight. An artist, like a scientist, strivesto be relevant and impactful. There is no beginning, no end,only the process, the act of doing. A finished piece of art islike a scientific publication, a milestone in a journey thatnever ends, a transient fleeting moment of accomplishmentwith which the artist and the scientist become bored,dissatisfied, frustrated at the very instant of completion but,nevertheless, lays the groundwork for the next step;inspiration for the next new and exciting chapter of the story;the next chapter that will again become boring, frustratingand imperfect but inexorably drives the scientist and artistforward.

What are the challenges of maintaining two verydifferent careers in parallel?When I was a very young boy I was asked like all children‘what do you want to be when you grow up’. Instead of theexpected answer of fireman or policeman, my family wasdismayed by my answer of a monk! I think I must have seensomething on TV about a monk’s life being filled with artistic(illuminated manuscripts) and scientific (Mendel) pursuits andthis looked great to me. Later at 16, after “O” levels anddeciding on “A” levels, I had to make a choice: art subjects orscience subjects. I reasoned that I could do art in my sparetime but not science in my spare time. So I chose science andwent to Imperial in London as an undergraduate. I also felt I

did not want to have a career at the mercy of subjective, artisticjudgements of what is good and what is bad and have to be ashowman to be successful. Success in science seemed to me to bebased on objective peer judgements instead. I understand now thatthis was probably a little naïve!

So I chose as a profession to be a scientist with the intention ofbeing an artist in my spare time, free of the subjective whims of artcritics. But what spare time? Science can consume 25 hours a dayand it did so for me for decades. In the end I realized that if I wasreally going to be an artist I had to deliberately and consistently carveout space, both physically in the form of a studio and psychologicallyby being more disciplined, efficient and focused as a scientist. So Irented space in a warehouse as a studio and made myself go to it inthe evenings and at the weekends. A bit like making myself drinkcoffee for the first time at the Dunn School! Like coffee at the DunnSchool, over time going to the studio became an easy, familiar habit,not a strange and bitter wrench. When I do art, it frees my mind todwell on science ideas; a kind of meditation. When I do science, theimages in journals, in westerns, northerns, southerns, in vivo imaging,immunohistochemistry etc drive artistic ideas. For me, there iscross-fertilization between art and science which are different sides ofthe same coin.

Simon Shelley exhibits his artwork under the pseudonym MichaelMetcalfe. His works of art can be viewed athttp://michaelmetcalfe.studio.

6 / FUSION . MICHAELMAS 2017

FUSION . MICHAELMAS 2017 / 7

We are lucky to have a large and diverse student community in theDunn School, which currently comprises over 65 graduate studentsfrom all over the world. The Dunn School Graduate StudentAssociation (GSA) makes an important contribution to student life, byactively promoting a social and supportive environment for studentsin the department.

The GSA is run entirely by a committee of student volunteers, whomeet regularly to share ideas and organise a wide range of social,scientific and career-related events. Since being founded by PattySachamitr in 2012, the GSA has flourished, and currently has a recordnumber of 14 committee members (Figure 1). We are very gratefulfor the enthusiastic backing and generous financial support wereceive from the department, which enables us to organise a widerange of events. Here, GSA members summarise some of the excitingevents and initiatives we have led in the past year, giving you aninsight into ‘a year in the life’ of the GSA.

Welcome events for new students Mariya LobanovskaGSA welcome events are designed to introduce new students to theDunn School and help them make a smooth transition into theirDPhil. Two years ago, the GSA created a ‘buddy’ system, where eachnew student is paired with a more experienced student from adifferent lab. Buddies can be a great source of advice, helping toprepare new students for their first year. The GSA organises socialevenings (Figure 2) and a tea party in the first week of term for thefreshers and their buddies, which provides an excellent platform tomeet students from all year groups and find out about the GSA. Thisyear, following the tea party, the GSA led a tour around thedepartment for the new students. They explored the departmentalfacilities, learned about the history of the Dunn School and found outmore about our research themes. We plan to make this tour a yearlyevent, giving new cohorts of graduates an early flavour of the vibrantand inclusive research community at the Dunn School.

Medical Sciences Careers Day Jessica HardyIn September 2016, we hosted our largest and most ambitious eventyet – the Medical Sciences Careers Day. Over 150 students andpost-docs attended, and were able to gain an invaluable insight into

a diverse range of post-DPhil career pathways. Speakers includedpharmaceutical scientists, a medical writer, a management consultantand a BBC producer, amongst others. A particular highlight was thelively discussion panel based around transitions in the academiccareer ladder (Figure 3). In addition, networking sessions over lunchand tea gave participants the chance to approach the speakersdirectly and chat with them in more detail. We receivedoverwhelmingly positive feedback from participants, and owing to thissuccess, we plan to run similar large-scale events once a year. Thenext Careers Day will take place in 2018, while this year’s event,taking place on Thursday 28th September, will feature an excitingrange of talks and discussions around the theme of ‘The Future ofScience’.

Speed Science Jessica ValliIn April, the GSA hosted a Speed Science Evening sponsored by theMedical Sciences Division. This event aimed to promote Dunn Schoolinter-lab relationships by allowing students and postdocs to find outabout their colleagues’ research. A broad range of research areaswere represented, and each participant had one-on-one conversationswith every other participant (Figure 4). Each conversation lasted 5

minutes, within which time both members had to summarise theirresearch with the help of graphical abstracts. This event providedinvaluable practice in clear and concise communication, and receivedexcellent feedback from all of those involved. At the end of theevening, each participant nominated their favourite conversation.

Figure 1. Some current members of the GSA committee.

A Year in the Life of the Dunn SchoolGraduate Student AssociationThe GSA Committee

Figure 2. A social evening with board games and pizza.

Figure 3. Discussion panel at the career’s day.

The runner up prize went to Bernadeta Dadonaite, and first prize waswon by Derek Xu, whose Drosophila illustrations left a lastingimpression.

Promoting contact with Dunn School alumni Felicia TanIt comes as no surprise that graduates of the Dunn School pursueambitious and exciting careers after completing their DPhils, oftenachieving success not only in academia, but also in industry,publishing, government, and the corporate world. Last year, the GSAinvited several Dunn School alumni to speak at our Careers Day,allowing students to draw from their experiences and gain inspirationand insight into a variety of possible career paths. This year, we hopeto continue connecting with alumni not only for the benefit of currentstudents, but also to encourage alumni to maintain links with theDunn School and each other. If readers of Fusion are interested inhelping current students by sharing experiences and advice, or wouldlike to reconnect with old friends or make new contacts, the DunnSchool Alumni Facebook and LinkedIn groups are looking for newmembers. Alternatively, readers are welcome to contact the GSA atgraduate.studies@path.ox.ac.uk.

Figure 4. Speed science event.

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If you have been at the Dunn School for any length of time, you arelikely to know about Norman Heatley and his role in developingpenicillin as a therapeutic agent, under the leadership of HowardFlorey, then Chair of Pathology and Head of Department. However,this story is not well known outside of Oxford’s medical and scientificcommunity. This book, by two authors who knew Norman Heatleyduring his life in Oxford, includes a foreword by Professor MatthewFreeman, current Head of Department, and is the first biography on hislife and work.

Eric Sidebottom completed his DPhil at the Dunn School, but afterretirement has become a keen medical historian. So the book startswith an interesting summary of the history of penicillin, starting fromearly evidence (as far back as ancient Egypt) on the application ofmoulds to treat wounds and infections, to the discovery and earlyattempts at isolating penicillin by Alexander Fleming and hiscolleagues, and ending with the invitation of Norman Heatley onto theteam of Florey and Ernst Chain, who were interested in developingpenicillin as a therapeutic antibacterial agent, and who famously werehonoured, along with Fleming, with a Nobel Prize for their work.Norman Heatley did not share this Nobel Prize, but after reading thisbook many will agree he should have.

Despite the title, the book covers the whole of Norman Heatley’s life -building a picture of how he became such a valued scientist andacademic, and of his kind and charitable personality - from his earlyyears in Suffolk and at boarding school in Kent, his pre-WWIIundergraduate and postgraduate studies in Natural Sciences at

Cambridge, to his main scientific career and later life in Oxford. Theauthors have spoken to Norman’s family and friends, and also hadaccess to Norman’s own observations, frequently cited in the book,which he recorded in his many laboratory notebooks and personaldiaries (we are told these have been preserved for posterity in thearchives at the Wellcome Trust). Through these sources, and theauthors’ sympathetic writing style, we get a real feeling for Norman’sinquisitiveness, ingenuity, practical skills, and warm personality, almostas if we were able to meet the man himself.

This biography also illustrates poignantly how science needs more thanjust great thinkers and visionary scientific leaders with extensivenetworks. In order to make, and exploit, scientific discoveries, it alsoneeds the makers, the developers of equipment and methods, who,with determination and tenacity, endeavour to make the ideas work inpractise. This book positions Norman Heatley as a kind of patron saintof that particular strain of scientist.

I highly recommend this beautifully produced little book. It can beread in its entirety in one or two evenings, by lay persons as well asscientists. And I would predict that a fair few readers will enjoy goingback for a second reading, to savour the exquisite little details,detours, and asides that bring the main story to life.

Penicillin and the Legacy of Norman Heatley by David Cranston andEric Sidebottom in available from Amazon ISBN: 978-1-909075-46-7.

Book review: Penicillin and the Legacy of Norman Heatley by David Cranston and Eric Sidebottom

Annemieke Kok

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Currently at Oxford, all medical students have access to an online libraryof anatomy modules, designed and written by the medical school, whichaims to support lecture-based and tutorial teaching. Modules are closelyaligned to the Oxford pre-clinical course syllabus and take the form ofdetailed notes, diagrams, dissection photographs, videos, interactivequizzes and references to experimental and clinical matters of interest.

In recent years, the Anatomy Section of the Medical School has run anAnatomy Fellowship programme, offering students the chance to becomeinvolved in the design and development of such teaching material. Themajor aims of the Summer Anatomy Fellowship, supervised by Mr TomCosker, Director of Human Anatomy at Oxford, are to investigate howand why current pre-clinical medical students use the medical school’sWebLearn Computer Assisted Learning (CAL) material for anatomy andidentify common recommendations to further improve provision. Withthis information, we aimed to review and revise a CAL anatomyeModule, whilst investigating other ways in which anatomy could betaught at the medical school. We therefore designed a comprehensiveonline survey, to be completed by current pre-clinical medical students.Questions were selected to assess how and why students use CALmaterial, which modules they found most and least helpful and how theythink modules could be further improved. Students were able to givehonest and anonymised feedback of the current provision in the freeresponse sections at the end of the survey.

The results of the online survey, taken by 76 first and second yearpre-clinical medical students, led us to produce an entirely revisedGastrointestinal Tract module on WebLearn. However, a number ofcommon student recommendations could not be easily implementedusing the current WebLearn (Magnolia) software. 63% of students saidthat it was not easy to quickly find information that they were lookingfor within the CAL material, and around 50% indicated that the currentself-check questions could be made more relevant and useful to mimicexamination-style, multiple choice questions in order to identify gaps inknowledge. A frequent comment in the “free answer” section of thesurvey was that it would be useful for anatomy modules to be availablein a mobile app format, allowing students to study in an interactive waywithout the need for internet access and/or a computer. Othersmentioned that some information was quite unorganised, recommendingthe use of clearer subsections and a more natural progression throughthe eModules. As such, a pilot module covering the Shoulder Jointanatomy was written and designed using eLearning course designerArticulate Storyline, incorporating as many suggestions made by students

as possible. The aim was to provide students with the opportunity tostudy discrete, syllabus-aligned anatomy modules in an interactivemanner, on both desktop and mobile platforms.

The intention of the Anatomical Walkthrough app, now known asOperation: Anatomy, is to build up the student’s knowledge of ananatomical structure from the inside-out, progressively increasing thecomplexity of anatomy until the entire structure is understood. In thecase of the shoulder joint, the app begins with the basic bonyarticulations, which then progresses to stabilising structures, blood supplyand innervation, musculature acting across the joint, movements possibleand finally clinically relevant information. Simple, digitally edited publicdomain Gray’s Anatomy diagrams were used throughout the module,highlighting important features discussed in the first-year course syllabus.In addition, joint movement animations were included to help studentsvisualise the actions and positions of various muscles.

Each subsection contains a set of mid-topic check-up questions, writtenin a variety of short-answer, multiple choice, drag-and-drop and “clickingthe named structure” style questions. If not answered correctly, studentscan readily navigate back to previous sections to find the relevantinformation. To ensure ease of navigation, a drop-down menu with asearch bar function is available for students to type in key words, whilstrelevant lecture slides and links to other webpages can be easilyaccessed by clicking the “Resources” tab.

Chris Horton, a former FHS student in Paul Fairchild’s lab, and Tom Kirk, an Engineering Masters student,discuss how they are aiming to keep medical students interested and engaged in anatomy by developing amobile and desktop application designed for the Oxford Medical Course.

Innovation in Medical Teaching: Harnessing Technology for the Teaching ofAnatomy

Chris Horton

Figure 1. Digitally edited Gray's Anatomy diagrams were included throughoutthe pilot app

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At the end of the module, understanding is checked by an end-of-topicquiz, which mimics the styles of questions encountered in Oxford’s end ofyear examinations. One of the key features of the app is the option torun the module as a “Walkthrough”. This means students cannotprogress to the more complicated parts of the anatomy beforesatisfactorily completing previous sections and mid-topic check-ups. Ifrun in this mode, the entire joint can be taught in the space of around 40minutes. If students are looking to quickly access specific information,the module can also be run in “Revision Resource” mode, allowingstudents to freely access all of the module’s resources without the needto complete each subsection.

The pilot module of this app was accepted for an oral presentation at TheAnatomical Society Winter Conference 2015 in Cambridge. Following thetalk, a number of senior anatomy lecturers from around the UK, includingthe University of Cambridge, University of Warwick and Royal College ofSurgeons of Edinburgh provided very positive feedback.

The app has been trialled with a number of medical students whocommented on its ease of use, comprehensiveness and direct relevance totheir course. Comments included: “It's by far the best anatomy resourceon the web or in our books” and “I think the questions at the end ofevery section are really useful in reinforcing what you have learnt becausethere have been too many instances when we read through largesections of text only to not remember anything. Having questions at theend really helps to highlight what we may have missed out / notremembered”.

Due to the positive responses received from students, the pilot modulewas submitted to the 2016 OxTALENT Competition run by OxfordUniversity IT Services. We were thrilled to be awarded the prize of best“Student IT Innovation”. The judges considered The AnatomicalWalkthrough to be an excellent example of an initiative to produce a

valuable learning and revision resource that can benefit future cohorts.They particularly liked the simplicity in the structure of the learningmaterials and the flexibility of the navigation, which offers a choicebetween a directed walkthrough or free access to individual resources forrevision purposes.

Following on from the pilot module and OxTALENT competition, we haveworked to develop the app in our spare time and have undergone a bit ofa rebrand. We have recently decided to code the app from the ground-upourselves, without the use of costly third-party software, as this offers usmore flexibility to add features students would like in an anatomylearning resource. New additions include an anatomy glossary, a dynamicquestion bank that regularly updates itself based on the user’s answersand a central webserver on which the app can be hosted and accessed.

We hope that other students would be interested in getting involved withwriting, designing and inputting new anatomy modules onto the appover the next few years, using the pilot module code as a template. Thehope is to gather a team of pre-clinical medical students during the longvacation to begin work on subsequent modules. In this way, acomprehensive, interactive anatomy resource can be made, whilstallowing students themselves to learn and revise anatomy by making andreviewing the modules. Further modules are currently being designed,which include further musculoskeletal modules, in addition tocardiovascular, respiratory and renal anatomy. We hope these will beavailable for use by the new cohort of pre-clinical medical studentsbeginning their studies in October 2017.

We have thoroughly enjoyed working on this project so far and wouldactively encourage others with similar interests in undergraduate

education and technology to give ita go. We’ve learned a great dealfrom the material writing and appdesign process and hope thatultimately students will benefit fromour modules. Perhaps in the future,if this project proves to be of use tostudents, modules for other topicscovered in the medicine course,including physiology, pharmacologyand pathology, may be developed.Should you have any suggestionsor comments, or would like to getinvolved in the project, we wouldbe delighted to hear from you!

Figure 2. University resources, including lecture notes and WebLearn pages canbe readily accessed through the app's "Resources" tab

Figure 3. - The application was presented at The Anatomical Society Winter Meeting (2015)at Magdalene College, Cambridge

Figure 4. App code in Ruby programming language.

Figure 5. The app has undergone a recentrebrand, has been re-written in Ruby codefor added flexibility and is now hosted on adedicated web server

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SPOTLIGHTMonika Gullerova was recently appointed to an Associate Professorship within the Dunn School. Here wecast a spotlight on the ground-breaking research for which her laboratory is already known and which sheintends to pursue within the Department.

Transcription and DNA Damage: Friend or Foe?

Monika Gullerova

Every living organism consists of minimal functional units or cells. Cellshave “bodies” called cytoplasm and “brains” called nuclei. The wholegenetic information, essential for the life of every cell, is stored andencoded in a single type of molecule, DNA, which is located in thenucleus. DNA is a very long, double stranded helix-shaped moleculewhich contains coding regions, genes, which are interrupted bynon-coding parts. Genes are transcribed into RNA molecules, which arein turn translated into proteins. Proteins are functional units whichperform various tasks essential for cell maintenance, chemicalprocessing, energy management and other functions in the nucleus andalso outside in the cytoplasm. Therefore, DNA and its transcriptiondetermines the fate of a cell. Unfortunately, DNA is continuouslyexposed to a multitude of environmental factors which can cause DNAdamage. If this is not immediately and efficiently fixed, a damaged cellcan die or become cancerous. The study of DNA repair mechanisms is,therefore, of fundamental importance. Not only do we learn about howthis process works, but, in doing so, we also develop knowledge thatcan help the design of future therapeutic strategies.

So far, most DNA repair research has focused on the functions of thevarious proteins involved. How do they recognize the damaged sites?How do they work to fix them? The primary research interest of my labis the novel role which transcription plays in DNA damage repair. Wehave discovered that as soon as damage occurs in some sections ofDNA, nascent transcription is initiated, resulting in production ofrelatively short non-coding RNA molecules. These molecules aretranscribed from both strands of DNA and due to theircomplementarity, they hybridize and form double stranded RNA(dsRNA) molecules. Normally, such dsRNA is very dangerous for a cell,because it can stimulate the interferon response. Indeed, the interferonpathway was originally developed as a protection mechanism againstmany viruses, as lots of them have their genomes encoded in a form ofdsRNA. Once dsRNA is sensed in a cell, interferons activate apoptosisultimately leading to cell death. So how can DNA damage lead to denovo transcription and formation of dsRNA without triggering theinterferon response and subsequent apoptosis? We actually found thatdamage-induced dsRNA is quickly processed into small effector RNAmolecules called DDRNA (damage derived RNA) by an enzyme calledDicer, which works like scissors. Dicer is mostly localized in thecytoplasm of cells, but upon DNA damage it is modified. Thismodification is called phosphorylation and it occurs on one particularamino acid within Dicer’s protein structure. This modification triggersDicer’s re-localization from the cytoplasm into the nuclei, where Dicerfinds its dsRNA substrates, close to DNA damaged sites, in order toprocess them into small RNA molecules (Figure 1).

We are now very interested in why these small RNAs are important forefficient DNA repair, how they function and which proteins they bind to.We employ state-of-the-art experimental and bioinformatics techniquesin order address these questions. RNA-dependent DNA damage repair(DDR) is a novel concept, which is currently not well understood.

Being based at the Dunn School, we are privileged to use its facilities todissect the molecular mechanisms of DDR, leading to efficient DNArepair. In particular, we are interested in the interplay between,transcription, RNA and the RNA interference pathway upon DNAdamage. These factors possess canonical primary functions in geneexpression and gene silencing. However, within physiological conceptsthey act as the key players in DDR, exhibiting their non-canonicalfunctions in processing of endogenous dsRNA, antisense transcriptionand mediation of DDR.

We therefore study RNA-dependent DDR on four connected levels: 1. Regulation: What regulates transcription at double strand breaks(DSBs)? 2. RNA: What are the features and function of newly synthesizedtranscripts around DSBs? 3. Processing: What factors are important in damage derived RNAprocessing? 4. Function: What is the role of RNA in the DDR pathway?

Overall, we aim to provide mechanistic insights into novelRNA-dependent DDR and its role in effective DNA damage repair. Ourresearch may unravel new insights into DDR and the maintenance ofgenome stability which are fundamental to a better understanding ofthe molecular mechanisms that cause cells to become cancerous.

Figure 1. Dicer shuttling between cytoplasm and nucleus upon DNA damage.

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RESEARCH FEATURE

Quality Control of Membrane Proteins

Pedro Carvalho

In cells, proteins are everywhere and participate in virtually all cellularactivities. Synthesized as linear, spaghetti-like molecules, proteinsbecome functional only upon acquiring defined three-dimensionalstructures. To ensure that this folding process is successfully executed,cells evolved exquisite protein quality control processes. Like in anymodern production line, quality control mechanisms operating duringprotein biogenesis identify aberrant molecules and promote theirdegradation. Defects in these protein quality control systems arefrequently linked to pathologies, such as cystic fibrosis and proteinaggregation diseases, highlighting their impact on normal cellphysiology.

A major interest of our laboratory has been the characterization ofprotein quality control mechanisms and their contribution to overallcellular homeostasis. In particular, we have focused on proteinsembedded in the lipid environment of cellular membranes, and whosebiogenesis takes place in a dedicated cellular compartment, theendoplasmic reticulum (ER). After folding in the ER, membrane proteinscan eventually traffic to their final cellular destination to function in awide range of processes, from lipid synthesis to signaling orcommunication in and between cells. However, not so rarely, moleculesfail to fold correctly and become substrates of a dedicated qualitycontrol process called ER-associated protein degradation or ERAD1. Thismultistep process encompasses recognition of the faulty molecules,their removal from the membrane and tagging for degradation in thecytoplasm (Figure 1).

Although ERAD was first discovered in human cells, most of itscomponents were identified in S. cerevisiae, the common baker’s yeast.This unicellular organism shares many features with human cells and itsfacile manipulation has led to a wealth of information on a variety ofbasic cellular mechanisms. Over a decade ago, during my postdoctoraltraining using yeast, we came to the realization that the various stepsof ERAD, from substrate recognition to degradation, were coupled inwell-defined protein assemblies (or protein complexes).2, 3

Importantly, we found that cells contain distinct flavors of thesecomplexes, each with specificity for different types of misfolded

proteins. Now, a major question that we are addressing in the lab ishow this specificity for substrates is achieved. How do these complexesdetect and recognize their substrates while all other proteins in thecrowded environment of the ER membrane are spared fromdegradation.

Aside from a canonical role in protein quality control, it was longknown that ERAD could also target a specific folded protein. This wasHMG-CoA reductase (HMGR in mammals and Hmg2 in yeast) requiredfor the biosynthesis of sterols, lipid molecules with essential cellularfunctions and linked to common diseases. Curiously, the degradation ofHMG-CoA reductase by ERAD is dependent on the levels of specificsterol metabolites in cells: if low levels of these metabolites werepresent, HMG-CoA reductase was stable; however, if the levels rose toa certain threshold the protein was quickly degraded. Thismetabolite-induced degradation of HMG-CoA reductase establishes afeedback mechanism required in maintaining sterol homeostasis.Moreover, HMG-CoA reductase degradation by ERAD involves a specificsubstrate adaptor which is not necessary for the recognition ofmisfolded membrane proteins. These findings suggested that ERADcould potentially have a more general role in the turnover ofproperly-folded ER proteins and consequently regulate ER functionsbeyond protein quality control.

To investigate how general was the role of ERAD in the degradation offolded proteins we took a global approach using mass spectrometry inour favorite model system, baker’s yeast. By comparing wholeproteomes of ERAD mutant and control cells we identified severalendogenous substrates of the different ERAD complexes. For example,we found that a second sterol biosynthetic enzyme, squalene epoxidase(SQLE in mammals and Erg1 in yeast), was also degraded by ERAD.4

Interestingly, the degradation of squalene epoxidase was also controlledby a sterol metabolite, however distinct from the one involved inHMG-CoA reductase regulation. Moreover, the degradation of squaleneepoxidase and HMG-CoA reductase was mediated by distinct ERADcomplexes. Subsequently, we and others showed that ERAD-mediateddegradation of these two sterol enzymes was conserved from yeast tomammals which is quite remarkable considering that a billion years ofevolution separates these organisms (Figure 2). Thus, ERAD has acentral and ancestral role in sterol homeostasis.1

Regulated degradation of HMG-CoA reductase and squalene epoxidasein response to specific sterol metabolites implies that cells have themeans to sense and measure the concentration of those metabolites inmembranes. However, how this is achieved is not known and is anarea of great interest in our lab. Insights on the molecular mechanismsof these sensing processes may allow us to interfere with sterolhomeostasis and eventually provide therapeutic strategies forhypercholesterolemia.

Figure 1. Biogenesis and quality control of membrane proteins within the ER.Newly synthesized proteins are inserted in the membrane of the ER in an unfolded state [1].Upon successful folding, proteins remain in the ER or traffic to their final destination [2].Misfolded proteins are recognized by ER-associated degradation (ERAD) that promotes theirextraction from the membrane into the cytoplasm for degradation [3].

FUSION . MICHAELMAS 2017 / 13

Other folded proteins not related to sterol metabolism were alsoidentified as ERAD substrates. While the characterization of thosesubstrates is still ongoing, the contribution of ERAD in defining thefunctional ER proteome is becoming increasingly clear.

The proteomic analysis revealed an interesting set of substrates that didnot follow any of the known ERAD complexes, suggesting theinvolvement of some previously-unidentified components. This wasindeed the case and through a number of experiments we identifiedthese elusive ERAD components. Unexpectedly, these were actingexclusively on the inner nuclear membrane (INM), a specialized regionof the ER facing the interior of the nucleus and involved in nuclearorganization, such as chromosome positioning or transcriptionalcontrol.5 To carry out these specialized functions, the INM has a uniqueset of membrane proteins distinct from the rest of the ER, even if thetwo membranes are continuous. Thus, it turns out that a major functionof this eccentric ERAD branch is to safeguard the unique composition ofthe INM by degrading ER proteins fortuitously mislocalized to thisspecialized membrane region (Figure 3). These findings support the

notion that cells carry out distinct forms of quality control based notonly on the folding state of proteins but also their localization insidecells. A challenge for the future will be to understand how spatial cuesare encoded in proteins and deciphered by quality control processes,such as ERAD. Another major priority in the lab is to identify thecomponents of INM quality control in mammalian cells and understandits contribution to organelle homeostasis.

In early days, the concept of protein quality control was perceived todeal exclusively with the folding state of proteins. The picture nowemerging places protein quality control processes, such as ERAD, in abroader context. These same pathways, by controlling the activities offolded proteins both in space and time, appear as major contributors tooverall architecture and organization of organelles and cells. Bydefining the principles guiding these protein quality control processeswe expect to learn about basic mechanisms of cell physiology. Webelieve these insights may eventually shed light on a variety of commondiseases linked to defects in protein homeostasis.

References1. Ruggiano A et al, (2014) J Cell Biol 204:869–792. Carvalho P et al, (2006) Cell 126:361–3733. Carvalho P et al, (2010) Cell 143:579–5914. Foresti O et al, (2013) Elife 1–17 doi:10.7554/eLife.009535. Foresti O et al, (2014) Science 751. doi:10.1126/science.1255638

Figure 2. A central role for ERAD in sterol homeostasis.Schematic representation of the feedback inhibition systems required for sterol homeostasis inyeast (left) and mammals (right). ERAD complexes are in bold and the enzymes targeted byERAD regulated degradation are enclosed in grey boxes.

Figure 3. Protein quality control by ERAD at the inner nuclear membrane (INM). The INM (in green) is continuous with the rest of the ER membrane (grey) but contains a uniqueprotein composition. An INM-specific ERAD complex (see inset) contributes to the unique INMproteome by promoting the degradation of mislocalized ER membrane proteins.

Making a gift to the Dunn SchoolThe Dunn School owes its existence to a philanthropic gift, from the Trustees of SirWilliam Dunn, and over the years has been the beneficiary of many acts ofphilanthropy, not least from those who have worked here. Any gift made to the DunnSchool helps to further research here, whether it is made to support a specific initiativesuch as the ones described in this newsletter, or at the discretion of the Head ofDepartment.

If you would like to make a gift to the Department this year, please use the gift formenclosed with this edition of Fusion. Please make sure that you have completed a giftaid form so that we can reclaim tax on your gift, and note that if you are a higher ratetax-payer, you can also set your gift against your tax liability for the year. All gifts madeto the Dunn School from the USA are also fully tax-deductible, when made through theUniversity’s ‘giving vehicle’ there, the Americans for Oxford, Inc organization.Ph

otograph courtesy of Paul Fairchild

14 / FUSION . MICHAELMAS 2017

Figure 1. Summary of known and hypothesised interactions between the influenza virus and host transcription across the transcription cycle. In this article, we discuss the role ofcap-snatching by the viral polymerase (FluPol) at host RNA Polymerase II (Pol II) transcription start sites and the role of the host nuclease Xrn2. In the body of genes, interactions have beenproposed to be mediated by splicing factors (SF) mutually required for host and viral mRNA splicing. Lastly, at the poly(A) site, the viral non-structural protein (NS1) can target host 3’-endmRNA processing by direct interaction with the CPSF30 subunit of the cleavage and polyadenylation (CPA) complex.

FOCUS ON PATHOGENSInfectious diseases remain a significant contributor to the global burden of disease. In this edition ofFusion, we showcase on-going research by different groups at the Dunn School into three distinct classesof pathogen: viruses, bacteria and eukaryotic parasites.

Influenza and Host Transcription: A Love-Hate Relationship

David Bauer

While it might not be the ‘hottest’ virus of the year or feature in thelatest round of Hollywood disaster movies, influenza remains one of thetop challenges to global health. Despite substantial public healthinfrastructure, the annual number of deaths attributable to seasonalinfluenza in developed countries (37-174 deaths/million population inthe United States) is only a single order of magnitude behind thenumber of deaths during the 2014-16 Ebola outbreak in West Africa(480 deaths/million population). These statistics do not include periodicglobal pandemics of new influenza strains, which have historicallyproduced much higher mortality rates (~300 deaths/million populationin 1968, 290 deaths/million in 1957, and 10,000-30,000deaths/million population in 1918).1

The molecular architecture of the influenza virus explains why the virusremains difficult to combat. The virus is a (-) sense, single-strandedRNA virus, with a genome divided into 8 segments, each containing 1-2major genes. This segmentation allows “reassortment” of a givensegment between two different viruses that happen to co-infect thesame cell. Evolution, in effect, converged again upon a type of sexualreproduction to increase diversity in the overall population of influenzaviruses. On a single-virus level, the origin of genetic diversity lies in theerror-prone replication of the viral genome. Like most RNA viruses,influenza encodes its own viral RNA polymerase that is responsible forcarrying out both transcription and replication.2 The lack ofproofreading by the polymerase during RNA synthesis leads to thegeneration of multiple mutations throughout the course of an infection,and rapid changes to the structure of the antigens that are present onthe surface of the viral particle.

Overall, the influenza RNA polymerase is a major determinant of thevirulence and host range of influenza viruses, and is a key regulator ofthe adaptation of avian influenza viruses to mammalian hosts. TheFodor group at the Dunn School – where I am a postdoc – studies themolecular mechanisms of how the polymerase replicates the viral RNAgenome and how it transcribes viral genes into mRNA in the context ofvarious virus-host interactions in an infected cell. Host-Shutoff and TranscriptionViruses employ a variety of mechanisms to achieve “host shutoff” withthe aim of increasing the amount of resources available to the virusand preventing the host cell from mounting an effective response to theinvading pathogen (Figure 1). Since the generation of an effectiveantiviral state fundamentally requires a change in gene expression bythe host, viruses have evolved mechanisms that target host mRNA atnearly all points in gene expression – from transcription, processing andexport from the nucleus to mRNA translation and stability in thecytoplasm.3

The most obvious strategy for a virus that encodes its own RNApolymerase might be to prevent host gene expression in preventingtranscription from initiating in the first place. Both (+) sense RNAviruses such as polio and (-) sense viruses in the Rhabdoviridae familydegrade or inhibit transcription Factor II D (TFIID), which is responsiblefor recognising the TATA box of promoters and recruiting RNApolymerase II.4, 5 Similarly, (-) sense viruses in the Bunyaviridae familytarget TFIIH, the helicase responsible for unwinding DNA to allowtranscription bubble formation.6

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Neither mechanism, however, is readily available to influenza. Thevirus, which transcribes and replicates its RNA genome in thenucleus, relies on the host RNA polymerase to generate 5’-cappedRNAs that the viral polymerase then co-opts to prime its ownmRNA transcription. This process of “cap-snatching” is carried outin a coordinated manner between the three subunits of theinfluenza polymerase,7 which bind the 5’ cap of the nascent hosttranscript, cleaves the transcript 10-15 nucleotides downstream ofthe cap, and feeds this fragment into the polymerase active site toprime transcription,8 to yield transcripts that appear to be “normal”host transcripts at their 5’ ends. The price the virus must pay forthis advantage is that it cannot completely shut off hosttranscription. Indeed, the virus must balance two competing needsduring the early stages of infection: to keep the host RNApolymerase active and supplying 5’-caps, while suppressing hostgene expression.

A Tale of Two PolymerasesOver 10 years ago, our lab showed that the viral polymerasedirectly binds the host RNA polymerase.9 This interaction is theresult of a specific interaction with the long, unstructuredC-terminal domain (CTD) of the host polymerase, which consists of52 heptad repeats with the sequence(Tyr-Ser-Pro-Thr-Ser-Pro-Ser)52. The CTD normally acts both as aplatform for the assembly and recruitment of various componentsof the transcription complex, and as a master regulator of Pol IIactivity via specific phosphorylation marks on the amino acids ofthe CTD.10

We have recently shown that the viral polymerase binds to Pol IIspecifically when the fifth serine residue of the CTD isphosphorylated (Ser5P), and that this binding requires a fullyassembled viral polymerase: the 3 individual subunits alone areinsufficient.11 This specificity provides an explanation for how theviral polymerase manages to efficiently sort through the ~80,000Pol II molecules in each nucleus12 in order to find those withnascent capped mRNAs available for cap-snatching: the Ser5Pmark is uniquely associated with Pol II that is in the initial stagesof transcription at the start of genes,13 while unassembled viralpolymerase subunits alone do not compete for host Pol II binding.

Once the viral polymerase has successfully obtained capped RNAfrom the host Pol II, the host polymerase is effectively redundant asfar as the virus is concerned. Recently, we have employedmammalian native elongating transcript sequencing (mNET-seq)that provides single-nucleotide resolution information of thedynamics of Pol II.14 The results generate a “snapshot” of thelocation of host Pol II and its various phosphorylated isoformsacross the genome during influenza infection. From these data, wecan examine behaviour on individual genes, as well as create moregeneralised “metaprofiles” of Pol II across the entire genome.This recent work – a collaboration between the Fodor, Murphy and

Proudfoot groups at the Dunn School – has shown that viralinfection leads to a depletion of Pol II from the gene bodies,downstream of transcription start sites. We have found that thisdepletion is likely specific to active viral transcription, and not justthe presence of the viral polymerase alone or other viral proteins.This result suggests that the very act of cap-snatching and RNAcleavage leads to suppression of host transcription elongation and,therefore, full gene expression. We are currently exploring theexact mechanism by which displacement of Pol II occurs. Oneintriguing coincidence is that the normal process of transcriptiontermination at the polyadenylation site (pAs) of a gene begins withthe cleavage of the RNA transcript from the elongating Pol II inorder to be passed to the 3’-end mRNA processing machinery.15

The product of this cleavage reaction is an RNA polymerase thatcontains a short piece of uncapped RNA trailing behind that can berecognised by the cellular exonuclease Xrn2, which chews back thisRNA and can then “torpedo” Pol II off the chromatin to terminatetranscription.16 It is possible that viral cap snatching thereforeinduces premature termination – in this case just a few hundredbases downstream of the transcription start site – by the samemechanism.

This process of premature termination comes with an added bonusfor the virus: in addition to preventing the host from transcribingany activated antiviral genes, the displaced Pol II is now free tore-initiate transcription and feed viral polymerase transcription inthe process. By the same token, the dependence of the virus onthe host’s own transcription initiation processes is a ripe target fornew antiviral agents against influenza that act by targeting theviral polymerase and the host’s transcriptional machinery itself, anapproach that we are now exploring.

References1. Viboud C et al, (2016) J Infect Dis 213:738–745 2. Te Velthuis AJW & Fodor E (2016) Nat Rev Microbiol14:479–493 3. Herbert KM & Nag A (2016) Viruses 8:154 4. Sharma R et al, (2004) Virology 320:195–205 5. Yuan H et al, (2001) J Virol 75:4453–4458 6. Le May N et al, (2004) Cell 116:541–550 7. Hengrung N et al, (2015) Nature 527:114–117 8. Te Velthuis AJW et al, (2016) Nat Microbiol 1:16029 9. Engelhardt OG et al, (2005) J Virol 79:5812–5818 10.Zaborowska J et al, (2016) Nat Struct Mol Biol 23:771–777 11.Martínez-Alonso M et al, (2016) J Virol 90:6014–6021 12.Zhao ZW et al, (2014) Proc Natl Acad Sci USA 111:681–686 13.Milligan L et al, (2016) Mol Syst Biol 12:874 14.Nojima T et al, (2015) Cell 161:526–54015.Proudfoot NJ (2016) Science 352:aad9926–aad9926 16.West S et al, (2004) Nature 432:522–525

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Host Pathogen Interactions: A Circular ArgumentJessica Martyn and Christoph Tang

A small step for bacteria - a giant leap backwards for mankindOur bodies are teeming with microorganisms. Many cannot be grownin the laboratory but the full catalogue of our microbial passengers hasbecome apparent by molecular methods to detect their presencethrough their tell-tale DNA sequences. Advances in sequencingtechnology have enabled us to look deep into ourselves and at themicrobes we carry, and appreciate the scale and diversity of ourbacterial passengers.

In most instances, our microbial flora is completely harmless to us. Theorganisms that make up our flora are called commensals, stemming fromthe word ‘to eat at the same table’. They are part of us and take a shareof our food. In return, they synthesise vitamins that we absorb asnutrients, and contribute to innate immune defence against the incursionof other microorganisms. There is compelling evidence that bacteria inthe intestinal tract stimulate our immune responses, locally at mucosalsurfaces, and also beyond to prime our systemic immune system.

However despite our mutually-beneficial coexistence with the vastmajority of microbes to which we are exposed, there is a small group ofbacteria, the pathogens, which have the capacity to inflict harm.Pathogens can establish themselves in niches usually devoid ofcommensal microbes, breaching anatomical barriers and evadingimmune defences. The distinction between pathogens and our floracan be blurred; there are several bacterial species that can be foundcolonising healthy people yet can cause disease in others.

In contrast, other pathogens are highly virulent, and have emergedfrom harmless bacteria in a single genetic step. A remarkable exampleis Shigella. To all intents and purposes, Shigella is identical toEscherichia coli which is a ubiquitous component of the flora of themammalian intestinal tract. However the four species of Shigella havearisen from harmless strains of E. coli between 500 and 20,000 yearsago by the uptake of one piece of circular DNA called a plasmid. Thisplasmid contains all the genetic information needed for the bacteriumto invade human cells, including cells of our immune system, elicitinflammation, and cause dysentery. The small step of acquiring aplasmid has led to a quantum leap in evolution and allowed thebacterium to exploit new niches in the human body. As a result,Shigella now causes over 100 million cases of dysentery per year, witharound 700,000 deaths among children living in impoverishedcircumstances, and eclipsing malaria as a cause of mortality.

Plasmids: going around in circlesPlasmids are circular, double-stranded DNA molecules that are distinctfrom a bacterium's chromosome. Often, genes carried by plasmidsprovide their host with fitness advantages, such by conferring virulenceor antimicrobial resistance (AMR). Resistance plasmids, R-factors, werefirst characterised in the 1950s as mediators of antimicrobial resistancein bacteria such as Salmonella. There then followed a period of intenseinvestigation into and modification of plasmids that allowed them to be

tamed as tools for the laboratory. Plasmids are exploited as vectors toclone, transfer, and manipulate genes. Then, because bacteria dividerapidly, plasmids are used as nano-factories to make DNA fragments inlarge quantities.

Plasmids have several interesting features. Their DNA must beduplicated within bacteria. For this, plasmids, which are inherentlyselfish genetic elements, hijack bacterial enzymes for their replication,similar to viruses in mammalian cells. DNA replication is carefullycontrolled to keep the number of copies of a plasmid inside abacterium relatively constant from generation to generation. Thenumber of plasmids in a bacterium can be in the hundreds, as for manyplasmids used in the laboratory. However for plasmids that exist in thewild, more is often less. Plasmids canimpose a significant burden on theirbacterial hosts, so many plasmids involvedin virulence and drug resistance are onlyfound as a single copy inside each bacterialcell. And even this can be a heavy load fora rapidly dividing bacterium. For example,Shigella readily loses its plasmid in thelaboratory, then grows at a far higher ratecompared with a strain which has kept theplasmid (Figure 1).

Maintaining single copy plasmids presentsa major challenge for bacteria to ensurethat each daughter cell receives one of thetwo copies generated following replication.Replication of plasmid DNA must be co-ordinated with cell division, andthen one plasmid must move into each daughter cell before division iscomplete. These aspects have been extensively studied in standardlaboratory plasmids: segregation can be achieved by plasmid-encodedsystems that either push (ParMR) or pull (ParAB) plasmids to the farends of dividing cells. However, complexities abound. For instance, theShigella virulence plasmid, pINV, surprisingly contains versions of bothsystems, a canonical ParAB sequence and an uncharacterised version ofParMR.

Coping with failureBut what happens if the systems to maintain a plasmid fail, and adaughter cell ends up without a plasmid? Plasmids even have a strategyin place to deal with this. As a selfish element, bacteria that areplasmid-free after division can be eliminated from a population through aprocess known as post-segregational killing (PSK) (Figure 2). PSKinvolves plasmid encoded toxin:antitoxin (TA) systems. In the normalstate of affairs when a bacterium has a plasmid, a toxin is produced andneutralised by a specific antidote, the antitoxin. There are six differenttypes of TA system depending on the nature of the antitoxin andmechanism of inactivation. Type II TA systems are abundant on plasmids,and are characterised by having both the toxin and antitoxin as proteins

Figure 1. Shigella flexnerigrown on Congo red agarshowing virulent bacteria (blackarrow), avirulent bacteria (whitearrow) which have lost part oftheir plasmid.

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which form a complex resulting in toxin neutralisation. If a plasmid islost, antitoxins are degraded in the cell as they are inherently unstable,allowing the more stable toxin to access its cellular target and arrestgrowth or kill the plasmid-less bacterium. In this way, bacteria lackingthe plasmid (which would have a growth advantage) are kept in check.

Interestingly pINV from Shigella has three functional TA systems.1 Thisseems like overkill as, for PSK, one TA system should be enough.However the TA systems operate under different circumstances. Onesystem, MvpAT, helps to keep pINV in bacteria when they are grown attemperatures found in the human body; another (GmvAT) serves thesame purpose but when the bacterium is in the external environment.Another role for the multiple TA systems on a single piece of DNA is thatthey can act to prevent loss of nearby sequences. MvpAT is near theorigin of replication while GmvAT is located adjacent to a gene thatcontrols the expression of virulence genes. So TA systems act locally aswell as globally.

TA systems are not just interesting because of how they contribute toplasmid biology. There are a plethora of TA systems in bacteria. E. colihas 30 on its chromosome, where they mediate the transition of thebacterium into a dormant state (with the bacterium being viable but notgrowing), making them temporarily resistant to killing by antibiotics(which generally act on dividing bacteria).2 Understanding how TAsystems function could be exploited for the development of newantimicrobials. For instance, the toxin CcdB which is part of another TAsystem on pINV targets the same protein in the bacterial cell at preciselythe same site as the fluoroquinolones, a major class of antibiotics. Sowhat we learn from the activities of these toxins could be translated intofinding novel antibiotics, which are desperately needed. GmvT encodedby the Shigella plasmid prevents protein translation in bacteria (anAchilles heel exploited by antibiotics such as streptomycin andaminoglycosides) by transferring a small chemical group to an as yetunknown molecule. We are currently studying how TA systems arecontrolled and how this could be used in biotechnology. Can we learnmore about TA systems to make plasmids that are never lost frombacteria which would be valuable for vaccine design and gene expressionstudies? Can we find ways to inhibit plasmid maintenance systems andmake them lose their virulence or resistance against antibiotics?

What goes around comes aroundThe rise in AMR in bacteria threatens to return certain areas of medicalpractice to a pre-antibiotic era in which there are no drugs to treatpeople with common infections. Resistance genes can cluster togetheron plasmids, which means that resistance against most classes ofantibiotics can be acquired in a single genetic event.

There is a correlation between the global dissemination of Shigella andthe acquisition of antimicrobial-resistance genes located on mobilegenetic elements such as plasmids, transposons, and integrons.Stably-inherited extended-spectrum β-lactamase (ESBL)-producingplasmids are increasingly being recovered from Shigella in developingcountries. Acquiring these plasmids, renders treatments with antibioticsineffective, and promotes spread of bacteria within and betweencountries.3 Plasmids are also prevalent in Neisseria gonorrhoeae, abacterium that infects the genitourinary tract, causing the sexuallytransmitted disease gonorrhoea. There has been a steady rise inantimicrobial resistance in this bacterium, which has now beenaccorded ‘Superbug’ status. Although the incidence of gonococcaldisease is highest in Africa, where it can act as a co-factor for thespread of HIV, remarkably little is known about strains of gonococcusfrom this continent. Through a collaboration with the Wellcome TrustUnit in Kilifi, Kenya, we have analysed a series of gonococcal isolatesfrom over 70 sex workers. What was striking is that over 98% ofstrains in this population carry plasmids. We found two main plasmids:one plasmid (pTetM) is virtually ubiquitous and confers resistanceagainst tetracycline, while the other plasmid (pBlaTEM) was found inaround half of the strains, and makes bacteria untreatable withpenicillin. While both plasmids have been found in gonococci before,this is the first time they have been seen at such high frequency. Forexample, only 4-5% of gonococcal strains in the UK and USA carryplasmids. Interestingly, pTetM can help the spread of pBlaTEM, sotreatment of high risk patients with tetracycline (for other genitalinfections) can promote resistance against penicillin, which waspreviously the mainstay of treatment.

PerspectivesPlasmids are common in bacterial populations, and have a profoundeffect on how bacteria interact with our cells and our attempts toeradicate them. While many plasmids have been studied and are usedin the laboratory, much is still to be learnt about plasmid biology inpathogenic bacteria. How do multiple plasmid maintenance systemsfunction on a single genetic element? How are plasmid genesregulated and integrated in the transcriptional networks of newbacterial hosts? Can understanding TA systems lead to novelantimicrobials to kill bacteria or to disarm them? And can we wake updormant bacteria in order to kill them with conventional antimicrobials?One thing is certain about the future, plasmids will be around.

References1. McVicker G et al.(2016) Nat Microbiol 2:162042. Harms F et al. (2016) Science 354:63183. The HC et al. (2016) Nat Rev Microbiol 14:235-250.

Figure 2. Post-Segregation Killing. Toxin-antitoxin (TA) systems are composed of an toxingene and an antitoxin gene. Presence of the plasmid ensures enough expression of theantitoxin to counteract the toxin. If the plasmid is lost, the antitoxin is degraded byproteases and its expression is not replaced by the plasmid. As there is no antidoteantitoxin present, the stable toxin kills the plasmid-less cell.

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It is an unfortunate truism that agents - as diverse as the spitfire fighterplane and the rattlesnake - that deliver injury and death often have abeautifully designed shape and form. This precise fitting of shape tofunction is seen in protozoal parasites causing devastating diseasessuch as malaria, trypanosomiasis and leishmaniasis. Such parasiteshave complicated life cycles in which they traverse significant transitionsbetween vector and host and host and vector. Evolutionary forces haveled to variations on a theme in the design of differentiated cells withshapes fitted to function – leading to beautifully “designed” infectiousagents.

Leishmania and leishmaniasisIn the Gull laboratory, we are interested in eukaryotic microbialparasites. One strand of our research is focussed on understanding howthe shape and form of the Leishmania parasite is fitted to itspathogenicity. We are not the first Dunn School researchers to studythe Leishmania parasite; whereas we now concentrate on the parasiteitself, in the 1970s David Bradley produced critical insights into thegenetics of mouse host susceptibility to, and immunology of,Leishmania infection.

There are numerous different species of Leishmania that cause diseasein humans, with the disease predominantly affecting people living inpoverty in the less-developed countries of South and Central America,Africa and Asia. The various forms of leishmaniasis range fromself-healing cutaneous lesions to severe and often fatal visceralleishmaniasis. The Leishmania parasite is spread by sandflies and, aswith other vector borne diseases, the distribution of the parasite iscompletely dependent upon the range of the vector, which in turn isdetermined by the environment. Climate change is likely to significantlyimpact sandfly distributions and there have been recent rises in thenumber of cases of leishmaniasis in southern Europe.

Life cycle transitions are accompanied by dramatic shape changesWhen an infected sandfly bites a human, infective Leishmania parasitestermed metacyclics (Figure 1) are injected into the bite site. Thesemetacyclic cells are taken up by macrophages where they differentiateinside a parasitophorous vacuole into the amastigote form. It is thisform that then proliferates, spreading the infection to furthermacrophages. A circulating infected macrophage is then taken up witha blood meal when another sandfly feeds and, in the midgut of thesandfly, the amastigote forms differentiate into motile promastigoteforms (Figure 1). These promastigotes traverse the sandfly digestivetract to a position close to the mouthparts where the parasitedifferentiates to the non-proliferating metacyclic forms, which are thencompetent to re-initiate the cycle.

A dramatic change in cell shape occurs when the Leishmania parasitetransitions from the sandfly vector and infects the macrophage. In thesandfly, Leishmania cells have an elongated cell body with a long,motile flagellum that provides the propulsive force enabling the cell tomove from the midgut to the mouthparts. After macrophage infectionthe Leishmania amastigote form has a short, non-motile flagellum thatbarely emerges from within the more spherical cell body. This flagellumis more likely to perform sensory functions pertinent to life inside themacrophage.

Understanding Leishmania shapeHow are these different parasite cell shapes achieved and modulated?The key to understanding the parasite cell shapes lies in defining howthe internal cytoskeletal architecture is built and how it influences theflagellum position and the flagellar pocket. The flagellar pocket ishugely influential in parasite biology: it is a plasma membraneinvagination at the base of the flagellum that has a vase-like shape,bulging out at the bottom and narrowing towards the top with theneck sheathing the flagellum for a short distance (Figure 2). The

Jack Sunter joined the Gull laboratory in October 2011 and has thoroughly enjoyed working at the benchand in the department itself. He will soon set up his own group up at Oxford Brookes University takingthis project with him and concentrating on understanding links between Leishmania cell shape, form andpathogenicity.

Leishmania – The Shape of a KillerJack Sunter and Keith Gull

Figure 1. Lifecycle of the Leishmania parasite, highlighting the different shapes and formsobserved in the vector and the host. The proliferative promastigote is found in the sandflymidgut; this form then migrates to the sandfly mouthparts and differentiates into the infectivemetacyclic. The metacylics are taken up by the host macrophages after being injected into thehost when the sandfly bites. In the macrophage, the metacylics differentiate into theintracellular amastigote form. (Leishmania cartoons courtesy of Richard Wheeler)

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flagellar pocket is a critical interface between the parasite and its host,as it is the only site for the uptake and secretion of largemacromolecules and is the likely location for critical surface receptors.

Insight into the cytoskeletal architecture of the flagellar pocket region inpromastigote forms has come from using electron tomography, anelectron microscopy technique that enables the generation ofhigh-resolution 3D models. Despite first impressions, the Leishmaniapromastigote cell is not radially symmetrical and a key break in thesymmetry of the cell is the attachment of the flagellum to one side ofthe neck region of the flagellar pocket. This attachment is mediated bya large cytoskeletal filament/membrane structure called the flagellumattachment zone. Significant changes to the flagellum attachment zonein the amastigote form seem to be associated with the remodelled cellshape and flagellar pocket.

Cell shape mutants and organisationWe have identified many of the protein components of the flagellum,

the flagellum attachment zone filaments and the membraneattachments. Our hypothesis was that deletion mutagenesis of genesencoding some of these proteins may produce shape changes inflagellar pocket organisation, allowing us to interrogate specific links topathogenicity. Leishmania are genetically tractable organisms and it isrelatively facile to express fluorescently-tagged copies of specificproteins of interest and also to generate gene knockout mutants. Aftercharacterising the cellular positions of flagellum attachment zoneproteins we produced a panel of knockout mutants.

As expected, the deletion of specific flagellum attachment zoneproteins resulted in the loss of attachment of the flagellum to the cellbody. But we noticed some unexpected consequences including adistinct change in the shape and organisation of the flagellar pocket(Figure 2). More surprisingly, we also observed a change in the overallshape of these parasites, with the Leishmania cell body becomingshorter and wider. Yet despite the changes observed, the cells were stillable to proliferate in culture, suggesting that these significant changesto flagellar pocket structure were easily tolerated. However, we wantedto establish whether these changes had consequences in the process ofinfection, in both the sandfly and mammalian macrophage.

In collaboration with groups in the UK and Czech Republic, we lookedat our mutants in vivo in established infection models. Early indicationsfrom these infections show that our mutants have much reducedinfectivity in both sandflies and animals in comparison to wild typeparasites and add-back controls in which those specific flagellumattachment zone proteins are present. There are a number of potentialexplanations for this loss of infectivity as the flagellar pocket region is anexus for many processes in the cell. However, changing the parasitecell shape in a subtle way appears to have had dramatic effects onpathogenicity in both host and vector.

Figure 2. Deletion of the FAZ5 gene causes a dramatic change in the shape and organisationof the flagellar pocket in the Leishmania amastigote with a large reduction in the length of theflagellar pocket neck.

Photograph courtesy of Tim Davies

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foreign entity and can remain dormant there until the environment issafe to reveal itself. The ART drugs are only able to kill an activelyreplicating virus but the dormant virus can remain integrated into thegenome for many years. Consequently, HIV patients have to use ARTdrugs every single day for the rest of their lives to prevent virus fromre-emerging, killing the immune cells and leading to AIDS.

The challenges for treating HIV infections still remain great, howeverthe incredible advancements in gene editing and therapy technologiesare paving a way for a brighter future. Gene editing is basically anon-demand ability to change and modify any gene within a givengenome. Because the human genome has thousands of genes, thetargeting of specific genes, without interfering with the functions of

Figure 1. Schematic showing the mode of action of the CRISPR/Cas9 system.

Earlier this year, Bernadeta Dadonaite, a DPhil student in Ervin Fodor’s group, won first prize in theBritish Society for Gene and Cell Therapy (BSGCT) science writing competition. Here we reproduce herwinning essay with kind permission of the Society

Gene Therapy against HIV: Fighting the Virus in DisguiseBernadeta Dadonaite

The world around you is teeming with pathogens. Lucky, your immunesystem is well trained to protect and guard you from any unwantedinvaders. Pathogens, such as viruses, are in a permanent arms racewith our immune system, constantly evolving to evade our defenses.Perhaps the most successful pathogens are the ones which havelearned to disguise their weapons. And what better disguise can therebe than becoming a physical part of the target - just the kind of tacticsused by HIV?

HIV, or human immunodeficiency virus, is a pathogen, which infectshumans and causes AIDS (acquired immunodeficiency syndrome).Around 38 million people worldwide are infected with HIV and, while

with appropriate treatment patients can live for many years, there is stillno cure or vaccine against the virus.1 The major treatment for peoplewith AIDS is an antiretroviral therapy (ART). ART is essentially a cocktailof various drugs, which stop the virus at different stages in its life cycle.However, preventing the virus from replicating is only a part of thechallenge, as HIV is very good at playing hide-and-seek. HIV primarilytargets our immune cells - the very same cells which are meant to seekand destroy the invaders. When HIV encounters an immune cell itenters and becomes part of the cell by integrating into the cell’sgenome. Integration into a host genome is an essential part of HIV’slife cycle and it is also a perfect Trojan horse strategy. When HIVbecomes part of the cell’s genome it is no longer recognized as a

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inserts its key and opens the door into the cell. Just like with any lock, aslight change in the CCR5 can prevent the key from fitting.Interestingly, it has been observed that a small deletion in the CCR5gene can prevent or attenuate HIV infection. 1% of Caucasiansnaturally carry the CCR5 deletion, indicating that it is not deleterious forhumans. This observation suggested that engineering patient cells tocarry the CCR5 deletion could be a potential treatment against the HIV.Indeed, experiments in a lab using CRISPR to introduce the CCR5deletion have successfully produced immune cells that are resistant toHIV infection.6 The first clinical trial exploring the same principles hasalso shown some promise. The patients whose blood cells whereisolated and modified to carry the CCR5 deletion showed a muchslower re-emergence of the virus in their blood in the absence of ART.7

Gene therapy has traditionally been used to treat genetic diseases,however, the advancements in gene editing technologies now enablethe use of gene therapy to treat infectious diseases as well. Manychallenges still remain for the use of CRISPR against HIV. HIV is

renowned for its ability to change and evade many treatmentstrategies. Potentially, HIV could outsmart CRISPR by changing itssequence, so that CRISPR can no longer recognize it. Even more so, theuse of CRISPR in humans has still many ethical and safety issues to beresolved. However, until effective vaccines against HIV are available,combination of ART and gene therapy approaches may be the only wayof developing a cure against HIV.

References1. World Health Organization. HIV/AIDS; 2016 Nov 30 [cited: 2017 Jan 16] http://www.who.int/mediacentre/factsheets/fs360/en/2. Kaminski R et al, (2016) Gene Therapy 23:690-6953. Liao HK et al, (2015) Nat Commun 6:6413 doi: 10.1038/ncomms74134. Kaminski R et al, (2016) Sci Rep 6:22555 doi: 10.1038/srep225555. Ebina H et al, (2013) Sci Rep 3:2510 doi: 10.1038/srep025106. Ye L et al, (2014) Proc Natl Acad Sci 111:9591-95967. Tebas P et al, (2014) New Engl J Med 370:901-10

Figure 2. Novel strategies to combat HIV-1 infection.

other genes, has always been a challenge. However, experiments usingCRISPR (clustered regularly interspaced short palindromic repeats)technology (Figure 1) suggest that there is a way of eliminating thedormant form of HIV.

The dormant HIV virus, also called a provirus, is essentially a gene indisguise. The cell’s surveillance system cannot distinguish the provirusfrom any other cellular gene. With the use of CRISPR, however, there isa possibility of cutting out this non-native gene from the genome(Figure 2). CRISPR can be specifically targeted to a single gene via aguide RNA molecule and the guide RNA can be made to recognise theHIV sequence in the genome. Once the CRISPR system has identifiedthe site of the provirus, it can cut it out, which leaves the cell free of theHIV. While in laboratory experiments this strategy has been verysuccessful,2-4 the challenge has remained in being able to deliver theCRISPR system into a living organism. Recently, however, a

combination of CRISPR and decades-long advancements in genedelivery methods showed the potential of this strategy in animals too.Adeno-associated virus vectors (AAVs) have for many years been usedas treatment-delivery systems. AAVs can be made to carry almost anygene and target specific cell types. The genes carried by AAVs integrateinto a genome and are expressed inside the cells like any normal gene.Last year the AAVs engineered to carry a provirus-targeting CRISPRhave been used to remove an integrated HIV genome from mice.5 Asmuch as 90% of blood cells were successfully depleted of provirus inthe mice, indicating the potential for the clinical use of this approach.

While gene therapy approaches to remove the provirus from a genomehave still a long way to go, gene therapy methods that prevent thevirus from entering a cell in the first place have already reached clinicaltrials. When HIV encounters an immune cell it has to bind a receptoron its surface. The receptor, called CCR5, acts like a lock into which HIV

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TECHNOLOGY FEATURE

The New Ice Age: Cryo-Electron MicroscopyComes to South Parks Road

Errin Johnson

The field of structural biology has undergone a spectacular transformation in recent years due to the rapid rise ofcryo-electron microscopy (cryo-EM) as a viable alternative to x-ray crystallography for protein characterisation. This wasfacilitated by a convergence of advancements in electron microscope design, signal detection, high performance computingand data acquisition/analysis software, which led to a significant improvement in the structural resolution achievable withcryo-EM. A wide range of proteins, as well as many other types of samples (eg: ribosomes, viruses, membranes andbiological polymers), can now be imaged using cryo-EM and modelled at sub-15 Å resolution, and in some cases near atomicresolution, using single particle analysis and sub-tomogram averaging techniques. With the recent establishment of astate-of-the-art cryo-EM Facility in the Science area on South Parks Road, led by the Dunn School’s Susan Lea, structuralbiologists across Oxford now have this powerful and cool (pun intended) technique at their fingertips.

What is cryo-EM?Proteins and other particulate samples, such as viruses, DNA, exosomesand liposomes, can readily be imaged with transmission electronmicroscopy (TEM) using the negative stain technique, which involvesdrying samples in a thin layer of heavy metal stain (such as uraniumsalts) on a TEM grid, so that the particles are visible in the TEM aswhite against the dark background of the stain. In this way, particlesize, behaviour (e.g. ability to aggregate) and purity (e.g. amount ofbackground protein) can quickly be assessed. However, negativestaining induces numerous artefacts, chief of which is the significantdistortion to particle structure which occurs as the sample is air dried inthe stain. Cryo-EM avoids these problems, as it allows particles to beimaged fully hydrated and in their near native state. This is achieved bythe process of vitrification, where the sample is applied to a TEM grid,blotted to remove excess solution and then frozen by immediatelyplunging it into liquid ethane, such that the particles are suspended atrandom orientations in a thin layer of amorphous ice within the foilholes of the grid. All subsequent handling and imaging (usingspecialised cryo-TEM holders) is done at cryogenic temperatures, whichinvolves numerous technical challenges. For instance, if the grid warmsabove -155°C, crystalline ice will be formed, which distorts structure, asdoes exposure to the electron beam itself. Cryo-EM samples are,therefore, imaged under low electron dose conditions, which, in theabsence of heavy metals to scatter the electrons, means that contrast incryo-EM images is low. To overcome the low contrast, 3D models aregenerated using many tens of thousands of particles. Every particleneeds to be correctly identified in the original image then theorientation with respect to the three dimensional object established.Combining signal from all the particles can then allow calculation of amodel of the object imaged. This is computationally challenging as, inaddition to the large size of the data sets, issues of heterogeneity(sample or conformational) also need to be resolved.

Cryo-EM in structural biologyCryo-EM was first developed in the early 80s, though it is only in thelast 5 years that it has been increasingly applied to structural biology,as multiple technical, analytical and computing limitations have beenovercome synergistically. A new cohort of microscopes, spearheaded by

The 300 kV Titan Krios cryo-TEM at the Old Observatory and part of the team who helpedwith the installation. From left to right: Alex Buzduga (FEI installation engineer), Susan Lea(Director of COSMIC), Errin Johnson (COSMIC & Dunn School EM Facility manager) and JoeCaesar (postdoc in the Lea group and computing wizard).

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the 300 kV FEI Titan Krios, with brighter, more stable electron sources,improved optics and higher thermal stability, allow data to be collectedautomatically over several days, with the help of new automatedacquisition software. Arguably the biggest game-changer for cryo-EMhas been the development of a completely new generation of cameras,called direct electron detectors, with higher sensitivity and much fasteracquisition rates than CCD cameras. Instead of taking one frame perexposure, direct electron detectors acquire hundreds of frames perexposure under very low dose conditions (e.g. 50 electrons/Å2 for a 50sexposure); these frames are then either summed into an integratedimage or recorded as a movie. Improved image processing algorithmsare then used to correct for the motion blurring that occurs when theparticles are exposed to the electron beam, resulting in an improvedresolution. At the same time, high performance computing has becomemore powerful and less expensive, which has made it more feasible tohandle the huge data output of these cameras (e.g. 10-20 TB for 1experiment) and subsequently perform the computationally-intensivedata analysis. Taken together, these advancements make it possible tomodel macromolecules from the MDa range down to ~200kDa in sizeat resolutions where it is possible to construct atomic models withcryo-EM. Although it is still challenging to resolve structures using thistechnique, it does open up possibilities for systems where the proteincannot be crystallised, as this is not required.

The Central Oxford Structural Microscopy and Imaging Centre(COSMIC)The critical need for a high resolution cryo-EM facility sited on SouthParks Road was recognised by Susan Lea and Matthew Freeman at theDunn School who, together with new Dunn School recruit TanmayBharat, formed a cross-departmental consortium with scientists fromChemistry (Carol Robinson) and Biochemistry (Francis Barr, MatthewHiggins, Simon Newstead) to establish one. This was successfullyachieved with funding from the Wolfson Foundation, the Wellcome Trustand the EPA Research Fund, and the backing of the Medical SciencesDivision. Commissioning of the Central Oxford Structural Microscopyand Imaging Center (COSMIC) began in the summer of 2016 withdelivery of the first two 200 kV cryo-EMs and has now been completedby the recent successful installation of the third TEM, a 300 kV FEI TitanKrios. COSMIC comprises a fleet of high end instrumentation andcomputing for every stage of the cryo-EM workflow:

• FEI Tecnai T12 TEM (in association with the Dunn School EMFacility) for screening samples using negative staining

• FEI Talos 200 kV cryo-TEM with STEM detector for screeningcryo-grids and for cryo-tomography and cryo-STEM tomography

• FEI Arctica 200 kV cryo-TEM with FEI Falcon 3 direct electrondetector for high throughput screening of cryo-grids and singleparticle analysis

• FEI Titan Krios 300 kV cryo-TEM with FEI Falcon 3 and GatanBioquantum direct electron detectors for single particle analysis,cryo-tomography and cryo-STEM tomography

• Four FEI Mark IV Vitrobots for preparing cryo-EM grids• High performance computing for data storage and processing

These microscopes will be flagship instruments of the new DorothyHodgkin Institute for Physical approaches to life-science research, dueto be completed in 2020. In the meantime, they are currently locatedin the Dunn School and at the Old Observatory. The good news is thatCOSMIC is now open to users and we are looking forward tofacilitating an exciting and diverse range of structural biology researchat Oxford with cryo-EM. We are also keen to push the boundaries ofwhat is possible with our new instrumentation and analysis tools bydeveloping new capabilities (e.g. in cryo-STEM and cryo-correlativemicroscopy) and breaking new resolution limits on a wide array ofbiomolecules.

Further readingIf you are interested in finding out more about COSMIC and the DunnSchool Bioimaging Facility, please see our website for more details andcontact information: www.dunnschoolbioimaging.co.ukThe following articles are a good starting point for learning more aboutcryo-EM:• Milne JL et al. (2013) Cryo-electron microscopy – a primer for thenon-microscopist. FEBS Journal 280:28-45• Bai XC et al. (2015) How cryo-EM is revolutionizing structuralbiology. Trends Biochem Sci, 40:49-57• Lučić V et al. (2013) Cryo-electron tomography: the challenge ofdoing structural biology in situ. J Cell Biol 202:407-419

Photograph courtesy of Thomas Dunwell

A B C

D E F

Dunn School Bioimaging Facility Image Awards 2016

A: Runner-up, Most Humorous - Screaming Leishmania by Flavia Moreira-Leite (Gull lab) on the FEI Tecnai 12TEM.

B: Winner, Light Microscopy - Day 14.5-of-pregnancy whole mount mouse mammary gland expressingmembrane tomato by Salah Elias (Robertson-Bikoff lab) on the Olympus FV1000 confocal.

C: Winner, Electron microscopy - Cryo-EM of a bacterial membrane protein complex and the correspondingclass averages created by particle alignment (inset) by Justin Deme (Lea lab) on the FEI Talos 200c TEM.

D: Winner, Most Humorous - Jack Cellington by Marvat Shurhi (Hassan lab) on the Olympus FV1000 confocal. E: Runner-up, Electron Microscopy - Three adjacent flagellar profiles with flagellar attachment zones on a

mutant Trypanosoma brucei bloodstream form cell by Jessica Valli (Gluenz lab) on the FEI Tecnai 12 TEM. F: Runner-up, Light Microscopy - Horizontal view of a Drosophila adult retina by Sonia Muliyil and Clemence

Levet (Freeman lab) on the Zeiss 880 Airyscan confocal.

24 / FUSION . MICHAELMAS 2017

FUSION . MICHAELMAS 2017 / 25

Carefully gathered by Eric Sidebottom over decades, the archive of theSir William Dunn School of Pathology has now been transferred to theBodleian and is being catalogued as part of the Wellcome Trust-fundedproject ‘75 Years of Penicillin in People’. The project has not onlyhistorical but scientific and social aims: from the full genetic sequencingof two isolates of the antibiotic fungi Penicillium and Cephalosporium,to sending Dunn School volunteers into local primary schools toencourage children to consider a future in science. And for the nextcouple of months, science writer and biographer Georgina Ferry will beconducting oral history interviews with current and former members ofthe department in an effort to capture the experience of being ascientist in the modern era, and to fill the historical gap between thewell-recorded penicillin and cephalosporin period and the present day.

On the archival side, the material gathered by Dr Sidebottom, whichderives from four main sources, will be made available to researchersvia four separate catalogues. The first describes a four-box personalarchive, a collection of degree certificates and family photographsbelonging to Peggy Pickles, who received her medical training in Oxfordin the 1930s and 40s, and went on to work as a clinical pathologist inthe Radcliffe Infirmary. She later married Alastair Robb-Smith who, in1937, at the fairly astonishing age of 29, became the head ofpathology at what is now the Nuffield Department of Medicine.

The next catalogue covers Gwyn Macfarlane’s working papers for his1985 biography Alexander Fleming, The Man and the Myth which, asthe title suggests, went a long way towards correcting the record onFleming’s contributions to the development of penicillin. What makesthis material particularly rich is the candid correspondence with peoplewho helped develop penicillin or knew people who did, includingNorman Heatley, Edward Abraham, and Dorothy Hodgkin.

Hodgkin was awarded the 1964 Nobel Prize in Chemistry for her workin identifying the molecular structure of organic compounds throughX-ray crystallography, and high among her achievements was her workon penicillin. In 1945 she was able to confirm E.P. Abraham and ErnstChain’s hypothesis that penicillin had a β-lactam ring, which madesynthesising the drug practically impossible. In a letter to GwynMacfarlane written on 26th December 1982, she neatly sums up hercontribution: “As the x-ray analysis of a chemically unknown structureof some complexity, the penicillin analysis was something of a landmark and plotted essential stages which have been followed since”[MS. 12387/3].

The third catalogue lists administrative papers, and these will join anexisting collection of the Dunn School’s papers in the Oxford UniversityArchives, which takes care of the administrative records of theUniversity.

There are intriguingstories to be foundhere, including earlyinvoices(Figure 1) andpersonnel records withevocative notes like‘left to get married’and ‘left for national service’ or ‘emigrated to Canada’. The papers alsoreveal a little-known fact about the Dunn School: it had in-houseinstrument makers. Stephen Bush, the Consulting Mechanic to theDunn School from 1927 to 1952, enjoyed a slightly odd arrangement inwhich he took a salary and had the use of two workrooms and anoffice in the School, but was required to fit those out himself and toemploy his own mechanics. His first responsibility was to fulfil theDunn School’s apparatus needs and to take care of its electrical andmechanical plant (as the Dunn School’s workshop still does) but he wasalso free to tout for business, taking orders for items like clamps andsyringes from institutions across the country, among them Liverpool’sSchool of Tropical Medicine and Oxford’s Radcliffe Infirmary.

The fourth catalogue covers the historical archive, which includes 16mmcine film, audio recordings, and thousands of photographs of staff andstudents. The archive ranges from a mid-century score book of theDunn School cricket team (good record against Zoology; crushed byDyson Perrins) to a Department of Pathology ink stamp. Notoriously,the Dunn School was so strapped for cash in the 1930s thatthen-professor Howard Florey had to ban the use of headed stationaryas a money-saving measure, and all outgoing letters were stampedinstead. It is pleasing to imagine that the exquisitely-tooled stamp inthe collection is Florey’s famous penny-pincher…but it seems morelikely to have been the Dunn School library stamp.

Many of the cine films date back to Florey’s era and can be grimviewing for the squeamish, with more than one featuring open andgangrenous war wounds in the process of being treated, all in gloriouscolour. But (look away modern ethicists) there’s light relief too in theform of a little film parody of the MGM studio’s famous roaring-lionlogo, made with a yawning lab rat. The photographs, meanwhile, forma priceless record of the Dunn School’s Heath Robinson-style penicillinfactory, and can be paired productively with resources like a 1977recording of a Norman Heatley lecture about penicillin.

The Dunn School archives will be available in the Weston Library, to get a reader’s card please seehttps://www.bodleian.ox.ac.uk/using/getting-a-readers-card. Thecatalogues will be found online athttp://www.bodley.ox.ac.uk/dept/scwmss/wmss/online/online.htm

Figure 1. They don’t make invoices like they used to. [Oxford University Archives PT 60/2].

HISTORY CORNER

The Dunn School Archive at the Bodleian Library

Charlotte McKillop-Mash

26 / FUSION . MICHAELMAS 2017

How I came to conduct the pilotclinical trials on penicillin is aremarkable example of being in theright place at the right time. I havehad this kind of good fortune onseveral occasions. Sheer coincidencedrew me into the penicillin story inJanuary 1941. I knew little of thepenicillin project until, on amid-January day, I called at Witts’office with some paperwork andwas invited in to meet HowardFlorey, the project’s leader.

The small team of scientists that Florey led at Oxford’s Sir William DunnSchool of Pathology had made its wartime quest the isolation ofpenicillin, the strongly antibiotic substance secreted by the mouldPenicillium notatum, with hopes of it proving of massive therapeuticvalue. Wonderfully innovative and resourceful, his team had overcomemajor early difficulties in extracting penicillin (a highly unstablebeta-lactam) from the culture medium of the mould into which it wassecreted in miniscule quantities. Using scarce early supplies, the teamhad shown its amazing effectiveness in combating life-threateningstreptococcal infection in mice. Since then the challenge had been inacquiring sufficient supplies of the antibiotic for clinical tests. For this,the laboratory had needed to transform into a factory culturing themould in hundreds of bedpan-like vessels, biscuit tins originally, just toaccumulate the necessary few grams of antibiotic for pilot testing oncases of severe streptococcal and staphylococcal infection.

The conversation between Witts and Florey that I interrupted was aboutgetting these tests under way at the Radcliffe Infirmary. Witts had beenasked to take this on. Knowing that I was short of a research project hequickly decided that I should. I had recently come to the end of myhouse job on his firm and was back as a Nuffield research student in hisDepartment considering research options. “Here’s your man”, he toldFlorey. “Fletcher can do the testing.” Becoming involved was as simpleas that.

Witts cannot have envisaged the historic role that he was giving me.Not even Florey could have envisaged the full magnitude of what I wasto reveal.

The first essential clinical test was that of proving that the penicillinbeing produced at the Sir William Dunn School was safe to give topatients. Side effects were a serious and feared possibility and could

even have been fatal. Who to give it to was a major ethical dilemma. Itwas Florey and Witts who decided that the initial test should be on apatient with limited time to live who agreed to take the risk in theinterests of medical research. I chose a woman (Elva Akers) who wasdying of inoperable cancer, with only a month or so to live, and sheagreed to be injected with the antibiotic. There were no ethicalcommittees then and I needed no other permission.

Records show that on 17 January 1941 I gave her an injection of 100milligrams of penicillin intravenously, while Witts and Florey looked on.Unfortunately, the antibiotic was not pure enough and she had a sharprise in temperature and a rigor (a shivering attack) about an hour and ahalf later, showing that there were still pyrogens in the penicillin sampleused. A rather unhappy Florey returned to his team at the Sir WilliamDunn School of Pathology to get more rigorous purification of theantibiotic under way.

With purer supplies produced, I set about testing the best way ofadministering penicillin. Giving it by mouth was soon ruled out. It didnot survive passage through the stomach. Tests of intravenous,intramuscular and rectal routes showed intravenous injection to be thebest way.

By then we were ready to treat a severe case of bacterial infection andearly in February I found one of remarkable severity in the Hospital’sseptic ward. All major hospitals had septic wards then for victims ofsevere septicaemia (bacterial infection of the blood), many of them withgrimly discharging abscesses. Mainly sufferers had dangerousstaphylococcal infection causing great pain. Septic wards were places ofmisery.

Florey had advised that I treat the worst case that I could find and thiswas of severe staphylococcal and streptococcal infection in a 43-year-oldOxford policeman (Albert Alexander). The infection had begun in a smallscratch on his face due to contact with a rose bush and had led tonumerous facial abscesses. This was a time when such scratches couldhave fatal consequences. He had been on the septic ward for severalweeks and treated with massive doses of the best anti-bacterial drug,sulphapyridine, without any effect. When I saw him on the 12thFebruary 1941 his eye had recently been removed due to the spread ofinfection, which had now reached his right shoulder and lungs. I judgedthat he had only days to live.

I began giving him intravenous injections of penicillin without delay,beginning with a dose of 200 milligrams, the largest that I had given,based on Florey’s calculation of an appropriate first injection for a man

Although seldom recognised for his pioneering work, Charles Fletcher is central to the story of penicillin,having been responsible for its first systemic use in patients. Here we reproduce an excerpt from MaxBlythe’s book ‘Pioneering Physician: The Life of Charles Fletcher 1911-1995’, with kind permission of theauthor, in which Charles Fletcher describes in an interview, his own involvement in these historical events.We are indebted to his son, Mark Fletcher, for allowing us to reproduce the pictures of his father.

The First Systemic Use of Penicillin: A Personal Account by Charles Fletcher

Figure 1. Charles Fletcher (circa 1953)

of his stature. Thiswas followed by 100milligram injectionsthree hourly. Thenearest I ever came toseeing a miracle wasin witnessing thispatient’s dramaticimprovement. After24 hours he lookedconsiderably betterand was able to sit upand eat. By the fourth

day he looked on the way to full recovery. Lazarus being raised from thegrave could not have been much more impressive. But regrettably ourpenicillin supplies ran out on the fifth day and about a week later herelapsed and never recovered. Treatment had not completely eradicatedthe infection by the time it prematurely ended.

Supplies of penicillin were a critical limiting factor. The penicillinlaboriously extracted from about a thousand litres of mould culturemedium was only enough for a day’s injections. Although Florey’s teamat the Sir William Dunn School supplemented the scarce suppliesavailable by laboratory reclamation of every milligram of penicillin in thepatient’s urine each day, we ran out too soon. We all shared bitterdisappointment at the patient’s death, but elation also that penicillinhad proved vastly more effective than we had dared to hope.

I have vivid memories of cycling from the Radcliffe Infirmary to the SirWilliam Dunn School of Pathology each evening with bottles of thepatient’s urine for penicillin extraction for re-use, and Florey and hiscolleague Ernst Chain anxiously asking how the patient was progressing.Florey, a laconic Australian, contained his delight well as penicillinrevealed its colossal therapeutic potential. “That’s good” he wouldcomment, while Chain was excitedly jubilant, in his mid-European way.

Over the next four months (February-June 1941) I treated a furtherseven patients with penicillin. Most were children to whom I could givesmaller doses of the antibiotic and so maximise the limited supplies thatFlorey’s team laboriously produced.

The first child that I treated was a young teenage boy of 15 who wasdesperately ill with septicaemia resulting from an infected surgical hipwound (following pinning of the hip). Sulphonamides had not relievedhis condition and I decided to test whether penicillin would. Again, theresult was astounding. Within two days of starting the treatment (on 22February 1941) he was feeling much better and remained so until thehip pin that had caused the infection was removed a month later.

Equally remarkable results followed the treatment of a man with agigantic carbuncle on his back (from 3-10 May, 1941). We don’t seecarbuncles nowadays: clusters of boils usually with numerouspus-discharging heads, resulting from severe localised staphylococcalinfection of the skin. This one was about four inches wide and causinggreat pain. Typically, recovery was expected to be slow, over at least amonth of hospital care, with various dressings being applied, but Idecided to test the effectiveness of penicillin and the difference was

recovery in a week. Daily, one watched the carbuncle shrinking. The most appalling case that I treated with penicillin was of a boy offour who was dying of cavernous sinus thrombosis. I still remember thegrim, cat-like cries that he was wailing. Infection that had begun locallyon the face had spread to the cavernous sinus, a cavity behind the eyeringed by numerous thin-walled veins, and resulted in a blood clot there.Watching this small boy recover from a virtually fatal condition wasamong the most memorable moments of my career. This was in thesecond half of May (from 13-22 May 1941).

Tragically this boy died of a brain haemorrhage a week later, due to therupturing of a weakened blood vessel. But from the post-mortemexamination we had confirmation that the infection had been completelyeradicated.

By then the amazing potential of penicillin was indisputable. Iremember saying to Florey “Sir, it must be marvellous to have achieved amedical breakthrough like this” and his typical no-frills reply “Yes, itdoesn’t happen to everybody and I don’t suppose I’ll do anything like itagain, but it is nice to have happened once in a lifetime”.Unfortunately, the magnitude of penicillin’s potential was not grasped bythe Medical Research Council and leading drug firms in Britain, wholargely resisted Florey’s requests for help in speeding penicillin researchand technology towards large-scale production. He ran into enormousmyopia in trying to promote its development, with key figures inresearch-funding and heads of pharmaceutical companies unable toperceive the importance of penicillin to the war effort, particularly itscapacity to heal war wounds and get troops back into action moreswiftly than ever before. Sir Edward Mellanby, my father’s successor ashead of the Medical Research Council, has to be black-marked for hisfailure to recognise the importance of penicillin production. It was thegreat black spot on an otherwise illustrious career. The lack of Britishinterest in penicillin production led to Florey seeking Americancollaborators and the antibiotic’s first commercial production being inthe USA, not Britain, with consequent colossal loss of revenue to thiscountry.

In addition to the tests of penicillin that I have mentioned, I also treateda urinary tract infection in a baby and several cases (4) of acute eyeinfection at the Oxford eye hospital, all successfully. But by June 1941penicillin production in Florey’s department developed problems.Pressures of penicillin production had exhausted the team and suppliesand my involvement ended.

I was dismayed and delighted to be named on the landmark penicillinresearch paper published by Florey and his team in the Lancet a shortwhile afterwards. My part had been so minor that I questioned myinclusion, but Florey insisted on the publication being by the wholeteam: Abraham, Chain, Fletcher, Gardner, Heatley, Jennings, Florey. Andlater, through Florey’s generosity, my name was included in the list ofpenicillin pioneers in the memorial garden established at MagdalenCollege, Oxford.

Max Blythe’s book (ISBN: 978-1-909075-42-9) is available to buyonline at www.wordsbydesign.co.uk at the recommended retail priceof £15.

Figure 2. Charles Fletcher talking with Norman Heatley onthe front steps of the Dunn School building. Heatley iswidely credited with having solved the difficulties ofpurifying penicillin from culture medium and patients’ urine.

FUSION . MICHAELMAS 2017 / 27

28 / FUSION . MICHAELMAS 2017

Few people will remember Roy Vollum as I do.He arrived in Oxford as a Rhodes Scholar fromCanada in 1921, the same year as Florey arrivedfrom Australia, likewise a Rhodes Scholar. Vollumwas my examiner in the Pathology andBacteriology BM exam in 1961. In those days allcandidates had practical exams and vivas and myown viva concentrated almost exclusively onRickettsia and their diseases, not generallyconsidered a major contributor to humaninfectious disease, and, I thought, rather belowthe belt as a major topic for a viva grilling.Vollum was also the senior bacteriologist in theRadcliffe Infirmary in 1965 when I was aResident Pathologist starting my professionaltraining. If we were presented with a CSFsample at night for suspected meningitis, we hadto phone Vollum and he usually arrived in thelab, whatever the time, because he feared wenovices might contaminate the sample. He wasnot an easy man!

Nevertheless, though not medically qualified, he was ahighly-regarded bacteriologist and had an importantrole in planning the new building for the Sir WilliamDunn School of Pathology which was completed andopened in 1927. He became an important member ofthe teaching staff at both the Dunn School and theRadcliffe Infirmary from that time onwards.

In the late 1930’s he received an ear from a cow thathad died of Anthrax and the culture he isolated wassubmitted to The American Type Culture Collection inVirginia, America. It was catalogued as 14578‘Vollum’. In 1942 the British Government authorisedbiological weapons tests using this strain of Anthrax

on Gruinard Island, off the Scottish coast. Sheepwere infected and subsequently died, requiring theisland to be quarantined for 48 years! However, morerecently and rather more significantly, on 2nd May1986 a sample of strain 14578 was shipped from theATCC in Virginia to the Iraqi Higher Education Ministryand a CIA report on 30th September 2004 confirmsthat Anthrax strain 14578 was Saddam’s exclusivechoice for bioweapons research. Perhaps ratherfortunately, Roy Vollum himself could never haveknown this story since he died in 1970, at the age of70. Who could possibly imagine that the Dunn Schoolplayed a role, however minor, in Saddam’s ambitionsfor WMD?

THE SIR WILLIAM DUNNSCHOOL OF PATHOLOGY

is a department of theUniversity of Oxford

website:www.path.ox.ac.uk

CONTACTS:Matthew FreemanHead of Department

Sir William Dunn School ofPathology, South ParksRoad Oxford OX1 3REmatthew.freeman@

path.ox.ac.uk

EDITORSPaul Fairchild

Tel: (44) (0)1865 285751email: paul.fairchild@

path.ox.ac.uk

Eric SidebottomTel: (44) (0)1865 285751email: eric.sidebottom@

path.ox.ac.uk

Fusion is produced by the Medical Informatics Unit, NDCLS, University of Oxford. Telephone +44 (0)1865 222746. Ref: Fusion2017/700

The Curious Tale of Cows, QuarantinedIslands and Saddam’s WMD

Eric SidebottomPhotograph courtesy of Paul Fairchild