what is essential difference between to ecology.pdfnatural surroundings. difference from other...
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What is essential difference between snake behind glass versus a wild animal?
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� intact cells� physiological properties� genetics� some extent behavior
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�Caged animal is out of context
�Removed from natural surroundings
�Stripped of its natural history
�No loner interfaces with environment
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Discipline deals with myriad ways in which organisms
� interact with� influence� influenced byNatural surroundings
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Difference from other disciplinesFocus is directed upward and outward
from individual to environment
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Other disciplines in biology are reductionist
�Cell biology� Physiology� Anatomy� Genetics
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Ecology is holistic
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� “Painful elaboration of the obvious”Cavitt’s Mom
� “The total relations of the animal to its organic and inorganic environment”Haeckel 1869
If true then what isn’t ecology
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Ecology
Physiology
BehaviorGenetics
Evolution
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Ecology overlaps all these disciplines thus need stricter definition
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� “Scientific natural history” Charles Elton, Animal Ecology
�“Study of structure and function of nature” Eugene Odum
� “Scientific study of the distribution and abundance of organisms” H.G. Andrewartha
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� The science of how organisms interact with their living and non-living environment
� “The scientific interactions that determine the distribution and abundance of organisms” Charles Krebs
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Definition is � Specific� Workable
• Where are organisms found• How many are there• Why
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�Perhaps the oldest of the natural sciences
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�Ecology is a basic science�Applied sciences use the knowledge
generated by basic sciences to solve problems
�Ecology is hierarchical
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�Individuals • (organisms)
�Populations• (groups of individuals of 1 species)
�Communities • (species that interact)
�Ecosystems • (community + abiotic environment)
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�Evolutionary - investigating adaptive aspects of ecological phenomena
�Distribution & Abundance - investigating the contemporary processes that determine• where an organism occurs (distribution)• how many organisms (abundance)• how numbers of organisms change (dynamics)
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�Example: Bluegill sunfish (Lepomis macrochirus)
�Large individuals• found in open water• feed mostly on zooplankton
�Small individuals• found in vegetation• feed mostly on benthic insects
Image from Conrell Univ. Dept. of Natural Resources
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�Hypothesis: bluegill choose habitat with most profitable prey
�Hypothesis: small bluegill are safer from predation in the vegetation
�Hypothesis: large bluegill aggressively exclude small bluegill from open water
�Evolutionary hypotheses --advantageous for individuals to choose certain locations
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�Hypothesis: bass predation on young bluegill
�Hypothesis: human predation on older bluegill
�Hypothesis: limited nest sites �Ecological hypotheses --
contemporary population level effects
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�How do bluegill affect populations of pumpkinseed sunfish?
�How do bluegill affect populations of Daphnia?
�What would be the effect of a 2oC increase in water temperature on bluegill populations?
�Ecological questions -- contemporary processes affecting distribution and abundance
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Ecologists utilize scientific method
2 approaches to answering questions and testing hypotheses in ecology
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Observation and Experimentation
1. Observation
Not all ecological phenomena are amenable to experimentation
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� Natural events can not be manipulated� Scale of question can be important� Ethics of experimentation can be
important
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� Observation or comparative studies often rely on correlation
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Major Goal
Want to understand causality
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AB
Leads us to infer that A “causes” B
AB
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A
B
Problem with correlation
Some other unknown event could be “causing” both to occur
X
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So what’s the problem with correlation?
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# Baptist Ministers
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� The problem is -
What can infer from correlation?
“Correlation does not prove causation”
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The number of Baptist Ministers is not causing an increase in the number of Taverns in Wisconsin.
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Roots of Ecology lie in natural history so technically it is as old as humans
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1. Primitive peoples relied ona. Huntingb. Fishingc. Food gathering
so needed detailed information on when/where to find food
2. Agriculture – increased need for knowledge of relationships of plants and animals to their environments
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3. Plagues attracted early writers attentiona. Exodusb. Aristotle – explained plagues of mice and
locusts.
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4. Principle of Ecological HarmonyGreeks established
term “Balance of Nature” – idea that nature is designed to benefit and preserve each species
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Each species had a special place in nature and extinction does not occur because it would upset the balance of nature.
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Today you can still find subscribers to this principle.
Why?
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1. Graunt 1662Described human population change
quantitatively (father of demography)• Birth rate• Death rate• Sex ratio• Age structure
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2. Leeuwenhoek 1687Studied reproductive rate of grain beetles,
carrion flies and human lice.
Counted # eggs laid by flies and extrapolated (746,496 in 3 months)
1st attempt to calculate theoretical rates of increase for an animal
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3. Buffon – Natural History 1756First to recognize all organisms are subject
to same processes
4. Malthus – Essay on Populations 1798Caluculated organisms can increase
geometricallyReproductive rate held in check by food
supply
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During this time
What prevents populations from reaching a point where they deplete their food supply?
What checks operate against a tendency toward geometric increase?
3 Centuries later ecologists are still asking these same questions
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1. Doubleday – “True Law of Population”1841
Questioned Malthus
Whenever species threatened with extinction, nature made a corresponding effort to increase fertility
Used humans in less developed countries
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Verhulst 1838 – Derived equation to describe rapid increase and stabilization of populations
Called logistic growth curve
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During most of time thinking hadn’t changed from idea of Plato
2 ideas during this time undermined “Balance of Nature”
1. Many spp becoming extinct2. Resources are limiting and competition is
important
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Consequences became clear �Malthus�Lyell�Darwin
“Providential Ecology” and “Balance of Nature” replaced with Natural Selection
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“Balance of Nature” continues to persist in modern ecology
� natural systems are stable� systems in equilibrium unless disturbed
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Ecology as Science
Roots � Natural History�Human Demography�Biometry�Applied problems in Agriculture,
Medicine
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Struggle to understand how nature works
Alfred Lotka – Metropolitan Life Insurance Co
Laid groundwork for mathematical ecology
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Until 1960’s ecology not considered important science
What happened?
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� Increase in human population�Destruction of natural environment
Caused public outcry and realization of problems