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Triplodia, replacement name for Triodia A. M. Clark, 1970(Ophiuroidea: Amphiuridae), non Hübner, 1820 (Lepidoptera:Hepialidae)Author(s): Richard L. Turner and Joel K. HallanSource: Proceedings of the Biological Society of Washington, 124(1):7-8. 2011.Published By: Biological Society of WashingtonDOI: http://dx.doi.org/10.2988/09-32.1URL: http://www.bioone.org/doi/full/10.2988/09-32.1

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Triplodia, replacement name for Triodia A. M. Clark, 1970 (Ophiuroidea:Amphiuridae), non Hubner, 1820 (Lepidoptera: Hepialidae)

Richard L. Turner* and Joel K. Hallan

(RLT) Department of Biological Sciences, Florida Institute of Technology, 150 West University

Boulevard, Melbourne, Florida 32901-6975, U.S.A., e-mail: rturner@fit.edu;

(JKH) 11282 Taylor Draper Lane #335, Austin, Texas 78759, U.S.A.,

e-mail: jhallan@austin.rr.com

Abstract.—The extant ophiuroid genus Triodia A. M. Clark, 1970 is a

junior homonym of the lepidopteran genus Triodia Hubner, 1820. The

replacement name Triplodia is herein proposed to accommodate Triodia

abdita A. M. Clark, 1970, assigned to this monotypic genus.

Keywords: brittlestar, eastern tropical Pacific, ghost moth, replacement name

The genus Triodia was established by

Clark (1970) to include a new species

from material included by Lutken &

Mortensen (1899) in Amphiura dalea Ly-

man, 1879, presently Amphioplus (Unio-

plus) daleus (Lyman, 1879) (Clark 1970).

The single specimen from the Eastern

Tropical Pacific Ocean was among four

assigned to A. dalea by Lutken &

Mortensen (1899), who illustrated several

segments of an arm to show differences in

the morphology of tentacle scales from

those of the other three specimens. In

addition to the triangular shape of the

tentacle scales, Clark (1970) derived the

name Triodia from the presence of three

pairs of oral papillae on each jaw and the

trio of plates formed by the oral shield

and a pair of accessory adoral shields.

The name Triodia has not appeared in the

echinoderm literature since its original

description, and it remains monotypic.

One hundred and fifty years earlier,

Hubner (1820) erected the genus Triodia

for a group of ghost moths. Triodia

Hubner, 1820 presently includes seven

species that range from Europe to central

Russia to Jordan (Nielsen et al. 2000).

Triodia A. M. Clark, 1970 is a junior

homonym of Triodia Hubner, 1820. In

accordance with ICZN Art. 60, a replace-

ment name is herein proposed for Triodia

A. M. Clark, 1970.

Class Ophiuroidea Gray, 1840

Order Ophiurida Muller & Troschel, 1840

Family Amphiuridae Ljungman, 1867

Triplodia nom. nov. pro Triodia A. M.

Clark, 1970 (preoccupied)

Type species.—Triodia abdita A. M.

Clark, 1970 by monotypy. The holotype

(MCZ 1488) in the Museum of Compar-

ative Zoology, Harvard University, mea-

sures 14 mm in disk diameter (Clark

1970). No other material is known to exist.

Etymology.—Triplodia is of feminine

gender, derived from the original name to

retain Clark’s (1970) reference to the three

sets of tripled features: three-sided (trian-

gular) tentacle scales, three pairs of oral

papillae, triad of oral shield and two

accessory adoral shields.

Distribution.—Lutken & Mortensen

(1899) assigned to Amphiura dalea four

specimens collected by the Albatross

from the Gulf of Chiriqui in 1891. Clark

(1970) based her new species on one of

three specimens taken from green ooze at

Albatross Station 3361 (6u109N, 83u069W;* Corresponding author.

PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON124(1):7–8. 2011.

1471 fm; 206 km south of Punta Burica,

Costa Rica/Panama). Lutken & Morten-

sen (1899) reported one additional spec-

imen of A. dalea from Albatross Station

3414 (10u149N, 96u289W; 2232 fm; 590 km

south of Puerto Angel, Oaxaca State,

Mexico). Clark (1970) clearly indicated(pp. 43, 68, 69) that the three remaining

A. dalea were not her Triodia abdita. She

referred to ‘‘other ‘Albatross’ specimens’’

(presumably of A. dalea) collected in 1904

from Stations 4493 (Monterey Bay, Cal-

ifornia, U.S.A.) and 4670 (off Lima, Peru)

and stated, ‘‘Their identification is prob-

ably correct.’’ It seems, therefore, thatTriplodia abdita (A. M. Clark, 1970) is

known only from the holotype.

Acknowledgments

We thank David L. Pawson (Smithso-

nian Institution) and Kayla M. Duro(Florida Institute of Technology) for

assistance with a literature search.

Literature Cited

Clark, A. M. 1970. Notes on the family Amphiur-

idae (Ophiuroidea).—Bulletin of the British

Museum (Natural History), Zoology 19(1):

1–81.

Hubner, J. 1820. Verzeichniss bekannter Schmet-

tlinge. Part 13. Jacob Hubner, Augsburg, pp.

193–208.

Lutken, C. F., & T. Mortensen. 1899. Reports on an

exploration off the west coasts of Mexico,

Central and South America, and off the

Galapagos Islands, in charge of Alexander

Agassiz, by the U. S. Fish Commission

Steamer ‘‘Albatross,’’ during 1891, Lieut.

Commander Z. L. Tanner, U. S. N., com-

manding. XXV. The Ophiuridae.—Memoirs

of the Museum of Comparative Zoology at

Harvard College 23(2):97–208.

Nielsen, E. S., G. S. Robinson, & D. L. Wagner.

2000. Ghost-moths of the world: a global

inventory and bibliography of the Exoporia

(Mnesarchaeoidea and Hepialoidea) (Lepi-

doptera).—Journal of Natural History 34(6):

823–878.

Associate Editor: Rick Hochberg.

8 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON