the macroinvertebrate stream fauna of martha's vineyard, with records from southeastern...

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The Macroinvertebrate Stream Fauna of Martha'sVineyard, with Records from Southeastern MainlandMassachusettsAuthor(s): Greg D. WhitmoreSource: Transactions of the American Entomological Society,134(3):431-457. 2008.Published By: The American Entomological SocietyDOI: http://dx.doi.org/10.3157/0002-8320-134.3.431URL: http://www.bioone.org/doi/full/10.3157/0002-8320-134.3.431

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431GREG D. WHITMORETRANSACTIONS OF THE AMERICAN ENTOMOLOGICAL SOCIETY VOLUME 134, NUMBER 3+4: 431-458, 2008

The Macroinvertebrate Stream Fauna ofMartha’s Vineyard, with Records from Southeastern

Mainland Massachusetts

GREG D. WHITMORE

The Trustees of Reservations, Vineyard Haven MA 02568

ABSTRACT

Stream sampling on the island of Martha’s Vineyard and in southeasternmainland Massachusetts produced 154 species, when added to other records, 157species have been documented, including 47 new state records. Unaltered streamson Martha’s Vineyard are cold water streams with temperatures similar to streamsat higher elevations in New England. Extended seasonal larval presence of sev-eral species was noted on Martha’s Vineyard.

INTRODUCTION

To date there has been no comprehensive published treatment of themacroinvertebrate stream fauna of the island of Martha’s Vineyard. Smith (2005)treated the non-insect macroinvertebrates of Massachusetts, including the islands.The Massachusetts Natural Heritage and Endangered Species Program created adraft checklist for the aquatic macroinvertebrates of Massachusetts (Chandler &Loose 2001), but it does not include records from Martha’s Vineyard. In 2000stream samples were taken over a two day period on Martha’s Vineyard by theMassachusetts Department of Environmental Protection (Fiorentino 2002) thatproduced 56 macroinvertebrate records, 14 of which were identified to specieslevel. The Ode News online database for the Odonata of southern New England(Nikula et al. 2006) lists 25 recent Odonata records for Dukes County (Martha’sVineyard) and 8 historical records (pre-1950). These records include pond andstream species.

Martha’s Vineyard occupies a transitional zone between northern and south-ern flora and fauna. The moderate climate has allowed some groups that cur-rently have a northern distribution to remain since the end of the most recentglaciation, while groups with a more southern distribution may have colonizedthe area more recently (Jones & Kimball 1943).

Martha’s Vineyard is located 3.5 miles from Cape Cod and is approximately100 sq.mi in size. Martha’s Vineyard, along with Nantucket marks the maximumadvance of the Laurentide ice sheet in New England (Oldale 2001). The streamfauna has been isolated from that of southeastern Massachusetts for 6,000 years,or since sea-level rise formed the Vineyard and Nantucket Sounds. The island isa combination of moraine and outwash deposits (Oldale 2001). There are severalwell developed small streams located on the morainal deposits of Martha’s Vine-yard. Their presence can be attributed to the large clay deposits underlying themorainal soils. The streams are groundwater fed with mainly sand bottoms andare less than four miles in length. Those streams which are unaltered by damsfunction as cold water streams with temperatures similar to streams found at higher

432 MASSACHUSETTS MACROINVERTEBRATES

elevations in New England. Streams which have been altered by the constructionof dams to create ponds were found to be distinctly warmer. The nearest perma-nent freshwater streams in southeastern mainland Massachusetts are on Cape Codapproximately seven miles northeast, though there may be ephemeral streams ata shorter distance. The streams found in southeastern mainland Massachusettshave similar characteristics to those on Martha’s Vineyard but tend to be larger insize and warmer. Water temperature measurements throughout the season showedthat the unaltered streams sampled on Martha’s Vineyard are distinctly colderthan the unaltered streams sampled in southeastern mainland Massachusetts. Ininitiating this study it was expected that the fauna on Martha’s Vineyard would besimilar to that of southeastern mainland Massachusetts because of the similaritiesin stream characteristics, geology and land use histories. It was also expectedthat the fauna of Martha’s Vineyard would be substantially smaller than the totalfauna found throughout the rest of mainland Massachusetts, due to the limitednumber of habitat types on Martha’s Vineyard.

PROCEDURE

Seven streams were sampled from Martha’s Vineyard. Three streams occur-ring on The Trustees of Reservations properties in southeastern mainland Massa-chusetts were also sampled to provide a preliminary faunal comparison with nearbystreams having similar characteristics. Streams were selected to represent a vari-ety of habitats. Unaltered cold water streams consisted of Roaring Brook, Un-named Priester’s Pond Feeder, Paint Mill Brook, and Witch Brook. Altered warmwater streams consisted of Mill Brook and Tiasquam River. Unaltered warm wa-ter streams consisted of Red Brook and Mashpee River. Ephemeral cold waterstreams consisted of Red Brook Feeder and Menemsha Hills Creek. Streamswere sampled at 2-3 week intervals in 2007 from March 13th through October15th. An additional sample was taken in Roaring Brook on January 16th, 2008.Paint Mill Brook was sampled only on October 15th, 2007, March 17th, 2008, andApril 1st, 2008. Witch Brook was sampled only on March, 17th, 2008. Streamswere sampled using aquatic kick nets and D-frame dip nets. A kick net was usedto sample high gradient sites and consisted of disturbing a 2m≤section of streambottom upstream of the net by kicking at the stream bottom to flip rocks anddislodge debris. A dip net was used to sample low gradient sites and consisted ofa series of jabs, focusing on debris piles, aquatic macrophytes, silt deposits andundercut stream banks. Both methods were based on EPA protocol (Barbour etal. 1999). Material collected was transferred to a white enamel pan in order tocondense the sample by removing larger pieces of debris. Samples were thentransferred to a 500ml jar containing 70% ethanol. Samples were sorted in awhite enamel pan containing water. All specimens were removed and the samplewas re-checked twice for quality purposes. All specimens were stored in glassvials containing 70% ethanol along with site location and collection date.

Water sampling was done on a monthly basis. Temperature, pH, conductiv-ity and total dissolved solids data were taken with a YSI EcosenseÆ pH10 meter.Stream discharge was recorded on a monthly basis and was obtained using aver-age stream depth across a set point and flow rate. Stream bottom compositiondata was recorded once in May and was based on the Wentworth scale for particlesize (Wentworth 1922). Canopy cover data was recorded once in May and is an

433GREG D. WHITMORE

estimation of the percentage of the stream that is shaded by overhanging trees/bushes.

Specimens of Insecta and Crustacea were identified using current species-level keys. Specialists were consulted for difficult groups (Don Chandler; alltaxa, John Burger; Simuliidae, both University of New Hampshire). Three spe-cies of Chironomidae that were collected by the Massachusetts Department ofEnvironmental Protection (MassDEP) during the Islands (Martha’s Vineyard) 2000Biological Assessment (Fiorentino 2002) were not collected during this projectand those records are included in the species records section of this paper. Mas-sachusetts species occurrence records were obtained from the MassachusettsNatural Heritage and Endangered Species Program (unpublished data; all spe-cies), and The Plecoptera Society of North America (Stark & Baumann 2006). Acomplete set of specimens is stored at the University of New Hampshire, DurhamNew Hampshire.

RESULTS

There were 154 species recorded during this project, with 137 species re-corded from Martha’s Vineyard. Three additional Chironomidae records fromthe 2000 Massachusetts DEP sampling are included, bringing the number of spe-cies on Martha’s Vineyard to140 and the total fauna of Martha’s Vineyard andsoutheastern Massachusetts to 157. All other previously recorded insects andcrustaceans found by the Massachusetts Department of Environmental Protec-tion on Martha’s Vineyard were also collected during this study, so the Massa-chusetts Department of Environmental Protection records for these species arenot included in the results. There were 70 species recorded from southeasternmainland Massachusetts. Of the 157 species, 47 are new state records. Thirty-five of these new records represent specimens collected solely on Martha’s Vine-yard. Six of these new records represent specimens collected solely in southeast-ern mainland Massachusetts. Six of these new records were recorded from bothMartha’s Vineyard and southeastern mainland Massachusetts. The large numberof new state records from a relatively small sampling area can be attributed to alack of intensive sampling throughout the rest of Massachusetts. Many of thenew records are species that are widely spread throughout New England, andmore extensive sampling throughout Massachusetts would likely produce similarrecords. The large difference in number of species collected on Martha’s Vine-yard compared to southeastern mainland Massachusetts can be attributed to thedifference in the number of sampling locations (7:3), as well as the presence ofcold water streams on Martha’s Vineyard.

This study produced a significant range expansion for Tetragoneuria spinosa.Collected from Mill Brook, T. spinosa is typically a southern coastal species,with the nearest records coming from New Jersey (Needham et al. 2000). Anaxlongipes was also found in Mill Brook, and is listed as a “Species of SpecialConcern” by the Massachusetts Natural Heritage and Endangered Species Pro-gram. A. longipes is typically found in coastal plain ponds and grassy ponds(Nikula et al. 2003). A number of species showed an extended seasonal presencein streams on Martha’s Vineyard. Several species, including Eurylophellafuneralis, Eurylophella verisimilis, Leptophlebia nebulosa, and Leuctra ferrugineawere present in streams several months longer on Martha’s Vineyard than in south-


eastern Massachusetts. Baetis tricaudatus and Amphinemura wui are found atlower elevations early in the season, and at higher elevations throughout the sea-son. On Martha’s Vineyard they were found in an unaltered stream (RoaringBrook) in large numbers throughout the season.

Martha’s Vineyard streams

Roaring Brook, Dukes Co., Chilmark, Gosnold’s Way, N41°22.685,W070°44.650. Elevation: 1m. Bottom: cobble 25%, gravel 25%, sand 50%.Canopy cover: 90%. Aquatic macrophyte cover: 30%. Water chemistry (IV-30-2007): temperature 10.8°C, electrical conductivity 90µS, pH 6.5, total dissolvedsolids 45ppm. Discharge (V-31-2007): .03m≥/s. Species recorded: 60.

Menemsha Hills Creek, Dukes Co., Chilmark, North Road, N41°21.932,W070°44.550. Elevation: 64m. Bottom: silt 100%. Canopy cover: 100%. Aquaticmacrophyte cover: none. Water chemistry (IV-30-2007): temperature 10.6°C, elec-trical conductivity 70µS, pH 5.6, total dissolved solids 35ppm. Discharge: toolow to record. Species recorded: 13.

Tiasquam River, Dukes Co., Chilmark/West Tisbury, State Road, N41°22.650,W070°40.615. Elevation: 5m. Bottom: gravel 5%, sand 75%, silt 5%. Canopycover: 80%. Aquatic macrophyte cover 20%. Water chemistry (IV-30-2007): tem-perature 12.7°C, electrical conductivity 74µS, pH 6.4, total dissolved solids 37ppm.Discharge (V-31-2007): .03m≥/s. Species recorded: 63.

Mill Brook, Dukes Co., West Tisbury, North Road, N41°24.141, W070°40.693.Elevation: 20m. Bottom:. Canopy cover: 50%. Aquatic macrophyte cover: 10%.Water chemistry (IV-30-2007): temperature 13.2°C, electrical conductivity 79µS,pH 6.3, total dissolved solids 39ppm. Discharge (V-31-2007): .06m≥/s. Speciesrecorded: 56.

Unnamed Priester’s Pond Feeder, Dukes Co., North Road, N41°24.248,W070°40.830. Elevation: 18m. Bottom: Canopy cover: 90%. Aquatic macrophytecover: 10%. Water chemistry (IV-30-2007): temperature 11.2°C, electrical con-ductivity 83µS, pH 6.1, total dissolved solids 41ppm. Discharge (V-31-2007):.01m≥/s. Species recorded: 60.

Witch Brook, Dukes Co., Seven Gates Road, N41°24.500, W070°41.500. El-evation: 72m. Bottom:. Canopy cover: 100%. Aquatic macrophyte cover: none.Water chemistry: not taken. Discharge: too low to record. Species recorded: 9.

Paint Mill Brook, Dukes Co., North Road, N41°23.326, W070°43.052. Eleva-tion: 36m. Bottom:. Canopy cover: 75%. Aquatic macrophyte cover: 25%. Waterchemistry: not taken. Discharge: not taken. Species recorded: 24.

435GREG D. WHITMORE

Martha’s Vineyard and Cape Cod streams.

Cape Cod streams

Mashpee River, Barnstable Co., Meetinghouse Road, N41°37.764,W070°29.029. Elevation: 46ft. Bottom: cobble 10%, gravel 40%, sand 40%.Canopy cover: 50%. Aquatic macrophyte cover: 10%. Water chemistry (V-8-2007):temperature 14.4°C, electrical conductivity 97µS, pH 6.9, total dissolved solids48ppm. Discharge (V-23-2007): .17m≥/s. Species recorded: 53.

Red Brook, Plymouth Co., Red Brook Road, N41°46.070, W070°38.150.Elevation: 5ft. Bottom: gravel 30%, sand 40%, silt 30%. Canopy cover: 25%.Aquatic macrophyte cover: 40%. Water chemistry (V-8-2007): temperature 11.4°C,electrical conductivity 88µS, pH 6.0, total dissolved solids 44ppm. Discharge (V-23-2007): .07m≥/s. Species recorded: 20.

Unnamed Red Brook Feeder, Plymouth Co., Red Brook Road, N41˚46.075,W070˚38.153. Elevation: 17ft. Bottom: sand 20%, silt 80%. Canopy cover: 100%.Aquatic macrophyte cover: 75%. Water chemistry: not taken. Discharge: too lowto record. Species recorded: 16.


Species Records and Notes

* New state records.** Species collected by MassDEPMV Martha’s VineyardSMA Southeastern mainland Massachusetts

CrustaceaAmphipoda

Crangonyctidae

Crangonyx pseudogracilis (Bousfield, 1958): [Martha’s Vineyard]: Tiasquam River, IX-13-2007 (1, net).

Notes: A single specimen was collected on MV in September.

Gammaridae

Gammarus fasciatus (Say, 1818): [Martha’s Vineyard]: Roaring Brook, VIII-9-2007 (4,net). Tiasquam River, III-27-2007 (1, kick; 6, net), IV-10-2007 (4 kick; 5, net), IV-30-2007 (1, kick; 6, net), V-16-2007 (2, net), V-31-2007 (1, kick; 2, net), VI-15-2007 (2, kick), VII-9-2007 (4, net), VII-25-2007 (2, net), VIII-22-2007 (2, kick), X-15-2007 (1, kick). Mill Brook, III-27-2007 (4, kick; 5, net), IV-10-2007 (1, kick; 2,net), IV-30-2007 (5, kick; 2, net), V-16-2007 (1, kick; 1, net), V-31-2007 (1, kick; 1,net), VI-15-2007 (8, kick; 8, net), VII-9-2007 (1, kick), VII-25-2007 (9, net), X-15-2007 (1, kick). Unnamed Priester’s Pond Feeder, IV-10-2007 (1, kick; 6,net), IV-30-2007 (3, kick; 4, net), V-16-2007 (19, kick; 14, net), V-31-2007 (9, net), VI-15-2007(5, kick; 9, net), VII-9-2007 (18, kick; 8, net), VII-25-2007 (6, kick; 5, net), VIII-22-2007 (7, kick; 3, net), X-15-2007 (2, net). [Southeastern mainland rivers]:Mashpee River, III-28-2007 (2, kick; 6, net), IV-20-2007 (3, kick; 7, net), V-8-2007(1, kick; 3, net), V-23-2007 (5, net), VI-15-2007 (10, net), VIII-13-2007 (4, kick; 7,net), IX-28-2007 (2, kick; 4,net). Red Brook, III-28-2007 (28, kick; 15, net), IV-20-2007 (10, kick), V-8-2007 (16, kick; 28, net), V-23-2007 (8, kick; 23, net), VI-15-2007 (23, net), VII-16-2007 (13, kick; 11, net), VIII-13-2007 (11, kick; 14, net), IX-28-2007 (20, kick; 5, net). Red Brook Feeder, IV-20-2007 (2, net), V-23-2007 (2,net), VI-15-2007 (1, net), VII-16-2007 (7, net), VIII-13-2007 (6, net).

Notes: Extremely common, collected from all larger streams throughout season.

Talitridae

Hyalella azteca (Saussure, 1858): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (1,kick), IV-10-2007 (4, net), IV-30-2007 (2, net), V-16-2007 (3, net), VI-15-2007 (1,kick), VI-27-2007 (2, net), VII-25-2007 (1, net), VIII-9-2007 (1, kick). TiasquamRiver, VI-15-2007 (1, net), VII-9-2007 (1, net). Mill Brook, III-27-2007 (1, net),IV-10-2007 (1, kick; 3, net), V-16-2007 (6, kick; 2, net), V-31-2007 (2, kick; 2, net),VI-15-2007 (8, net), VII-9-2007 (5, kick), VII-25-2007 (2, kick; 10, net), VIII-22-2007 (2, kick, 2, net), IX-13-2007 (8, kick; 1, net), X-15-2007 (2, kick). UnnamedPriester’s Pond Feeder, IV-10-2007 (1, kick; 2, net), IV-30-2007 (2, net), V-16-2007(3, kick), V-31-2007 (4, kick), VI-15-2007 (1, kick), VII-9-2007 (3, net), VIII-22-2007 (1, net), X-15-2007 (1, net). Paint Mill Brook, IV-1-2008 (2, net). [Southeast-ern mainland rivers]: Mashpee River, VI-15-2007 (2, kick).

Notes: Collected from larger streams throughout season.

437GREG D. WHITMORE

IsopodaAsellidae

Caecidotea communis (Say, 1818): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (3,kick; 6, net), III-27-2007 (3, net), IV-10-2007 (5, kick; 4, net), IV-30-2007 (2, net),V-16-2007 (1, kick; 2, net), VI-18-2007 (1, net), VI-27-2007 (1, net), VII-9-2007 (1,kick; 3, net), VII-25-2007 (4, net), VIII-9-2007 (2, net), VIII-22-2007 (1, kick), IX-13-2007 (2, kick; 1, net), X-15-2007 (3, net), X-25-2007 (2, kick). Menemsha HillsCreek, III-19-2007 (19, net), IV-10-2007 (7, net), IV-30-2007 (9, net), V-16-2007(4, net), V-31-2007 (7, net), VI-18-2007 (7, net), VII-9-2007 (7, net), VII-25-2007(9, net). Tiasquam River, III-27-2007 (1, kick; 5, net), IV-10-2007 (4, kick), IV-30-2007 (4, kick; 5, net), V-16-2007 (5, kick; 4, net), V-31-2007 (2, kick; 1, net), VI-18-2007 (2, kick; 3, net), VII-9-2007 (2, kick; 5, net), VII-25-2007 (1, kick; 5, net),VIII-22-2007 (2, kick), IX-13-2007 (1, net). Mill Brook, III-27-2007 (1, kick; 7,net), IV-10-2007 (1, kick; 3, net), IV-30-2007 (1, kick; 2, net), V-16-2007 (4, kick;2, net), V-31-2007 (2, kick; 2, net), VI-18-2007 (5, kick; 8, net), VII-19-2007 (2,kick), VII-25-2007 (5, kick; 5, net), VIII-22-2007 (1, kick; 1, net), X-15-2007 (1,kick; 4, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (3, net), IV-30-2007 (6,kick; 3, net), V-16-2007 (7, kick; 7, net), V-31-2007 (4, kick; 2, net), VI-18-2007 (3,kick; 4, net), VII-9-2007 (2, kick; 3, net), VII-25-2007 (2, kick; 5, net), VIII-22-2007 (2, kick; 3, net), IX-13-2007 (3, kick; 2, net), X-15-2007 (1, kick; 1, net).Witch Brook, III-17-2008 (8, kick; 5, net). Paint Mill Brook, X-15-2007 (5, net),III-17-2008 (4, kick; 7, net), IV-1-2008 (1, kick; 6, net). [Southeastern mainlandrivers]: Mashpee River, III-28-2007 (2, net), IV-20-2007 (2, kick; 5, net), V-8-2007(6, net), V-23-2007 (4, net), VI-15-2007 (1, kick; 10, net), VIII-13-2007 (1, kick; 1,net), IX-28-2007 (1, net). Red Brook, III-28-2007 (4, kick; 4, net), IV-20-2007 (2,kick; 2, net), V-8-2007 (2, kick; 4, net), V-23-2007 (4, kick, 2, net), VI-15-2007 (2,net), VII-16-2007 (2, kick). Red Brook Feeder, III-28-2007 (7, net), IV-20-2007 (8,net), V-23-2007 (19, net), VI-15-2007 (3, net), VII-16-2007 (9, net), VIII-13-2007(3, net), IX-28-2007 (3, net).

Notes: Collected from all streams from March-October. Extremely common andabundant.

InsectaColeoptera

Note: Coleoptera adults and larvae were collected and recorded.

Chrysomelidae

Neohaemonia melsheimeri (Lacordaire, 1845): [Martha’s Vineyard]: Mill Brook, V-31-2007 (3 adults, net).

Notes: Collected on MV in May.

Dytiscidae

Acilius semisulcatus (Aube 1838): [Martha’s Vineyard]: Mill Brook, IV-10-2007 (1adult, net).

Notes: A single adult was collected on MV in April from a backwater area.

*Neoporus undulatus (Say, 1823): [Martha’s Vineyard]: Mill brook, IV-10-2007 (1 adult,net).

Notes: A single adult was collected on MV in April from a backwater area.


Elmidae

Macronychus glabratus (Say, 1825): [Martha’s Vineyard]: Roaring Brook, VII-25-2007(1 larva, kick). Tiasquam River, V-16-2007 (3 adult, kick; 1 adult, net), V-31-2007(1 adult, kick; 1 adult, net; 2 larvae, net), VI-19-2007 (2 larvae, net), VII-9-2007 (2larvae, net), IX-13-2007 (1 larva, net), X-15-2007 (1 larva, net). Mill Brook, IV-10-2007 (3 adult, net), IV-30-2007 (1 adult, kick), V-16-2007 (3 adult, kick; 10 adult,net; 3 larvae, net), V-31-2007 (4 adult, net; 2 larvae, kick), VI-18-2007 (3 adult, net;2 larvae, net), VII-19-2007 (2 larvae, kick), VII-25-2007 (1 larva, kick; 1 larva,net), VIII-22-2007 (3 larvae, net), IX-13-2007 (1 larva, kick). Unnamed Priester’sPond Feeder, VII-25-2007 (1 larva, kick). [Southeastern mainland rivers]: MashpeeRiver, III-28-2007 (1 adult, kick), V-8-2007 (2 adult, net), V-23-2007 (3 adult, net),VI-18-2007 (1 larva, kick), VIII-13-2007 (1 adult, kick).

Notes: Larvae were collected on MV from May-September and in SMA in June. Adultswere collected on MV from April-June and in SMA from March-August. Commonon MV and in Mashpee River.

Optioservus ovalis (LeConte 1863): [Martha’s Vineyard]: Roaring Brook, IV-10-2007 (1larva, kick), V-31-2007 (6 adult, kick; 10 larvae, kick), VI-18-2007 (2 adult, kick;10 larvae, kick; 1 larva, net), VII-9-2007 (1 larva, kick), VII-25-2007 (5 larvae,kick), VIII-9-2007 (17 adult, kick), IX-13-2007 (6 adult, kick; 1 larva, kick).Tiasquam River, III-27-2007 (4 larvae, kick), IV-10-2007 (5 larvae, kick), IV-30-2007 (3 adult, kick; 4 larvae, kick), V-16-2007 (3 adult, kick; 7 larvae kick), V-31-2007 (3 adult, kick; 1 larva, kick; 1 larva, net), VI-18-2007 (3 larvae, kick), VII-25-2007 (8 adult, kick; 5 larvae, kick), VIII-22-2007 (2 adult, kick; 2 larvae, kick), IX-13-2007 (1 larva, kick), X-15-2007 (2 adult, kick; 9 larvae, kick; 2 larvae, net). MillBrook, III-27-2007 (1 larva, kick), V-16-2007 (1 adult, net), V-31-2007 (2 adult,kick), VII-9-2007 (1 larva, kick), IX-13-2007 (1 larva, kick). Unnamed Priester’sPond Feeder, IX-13-2007 (2 larvae, kick). [Southeastern mainland rivers]: MashpeeRiver, III-28-2007 (2 adult, kick; 7 larvae, kick), IV-20-2007 (1 adult, kick; 5 larvaekick), V-8-2007 (2 larvae, kick), V-23-2007 (3 adult, kick; 4 larvae, kick), VI-15-2007 (2 adult, kick; 16 larvae, kick), VIII-13-2007 (1 adult, kick, 4 larvae, kick),IX-28-2007 (4 larvae, kick).

Notes: Larvae were collected on MV from March-October and in SMA from March-September. Adults were collected on MV from May-October and in SMA fromMarch-August. Common on MV and in Mashpee River.

Oulimnius latiusculus (LeConte, 1866): [Martha’s Vineyard]: Tiasquam River, IV-30-2007 (1 adult, kick), V-16-2007 (1 adult, kick), V-31-2007 (1 adult, kick).[Southeastern mainland rivers]: Mashpee River, III-38-2007 (1 adult, kick), IV-20-2007 (1 adult, kick), VI-15-2007 (2 adult, kick). Red Brook, III-28-2007 (1 adult,kick).

Notes: Adults collected on MV from April-May and in SMA from March-June. Larvaedid not turn up in sampling.

Promoresia tardella (Fall, 1925): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (2adult, kick; 4 larvae, kick), IV-10-2007 (4 adult, kick; 3 larvae, kick), V-16-2007 (5adult, kick; 2 larvae, net), V-31-2007 (11 adult, kick; 27 larvae, kick; 5 adult, net;12 larvae, net), VI-18-2007 (9 adult, kick; 13 larvae, kick; 4 adult, net; 19 larvae,net), VI-27-2007 (1 adult, net; 1 larva, net), VII-25-2007 (2 adult, kick; 8 larvae,kick; 12 adult, net; 28 larvae, net), VIII-9-2007 (2 larvae, kick), IX-13-2007 (2adult, kick; 1 larva, kick; 2 larvae, net). Tiasquam River, III-27-2007 (1 adult, kick;1 larva, kick), IV-10-2007 (1 adult, kick), IV-30-2007 (1 adult, kick; 1 adult, net),

439GREG D. WHITMORE

V-16-2007 (7 adult, kick; 3 larvae, kick; 1 larva, net), V-31-2007 (3 adult, kick; 3larvae, kick), VI-18-2007 (1 larva, kick), VII-9-2007 (1 larva, kick), VII-25-2007 (1adult, kick; 2 larvae, kick; 2 adult, net; 1 larva, net), VIII-22-2007 (1 larva, kick; 1adult, net), X-15-2007 (1 larva, kick; 1 larva, net). Unnamed Priester’s Pond Feeder,VIII-22-2007 (1 larva, kick).

Notes: Adults were collected on MV from March through September. Larvae werecollected on MV from March through October. Common on MV.

*Stenelmis crenata (Say, 1824): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (1adult, kick; 9 larvae, kick; 1 larva, net), IV-10-2007 (1 adult, kick; 3 larvae, kick; 1larva, net), IV-30-2007 (5 adult, kick; 6 larvae, kick; 1 larva, net), V-16-2007 (2adult, kick; 8 larvae, kick; 3 larvae, net), V-31-2007 (7 adult, kick; 9 larvae, kick; 3larvae, net), VI-18-2007 (7 adult, kick; 7 larvae, kick; 3 larvae, net), VII-9-2007 (3adult, kick; 15 larvae, kick; 8 larvae, net), VII-25-2007 (4 adult, kick; 12 larvae,kick; 3 larvae, net), VIII-22-2007 (16 larvae, kick; 3 larvae, net), IX-13-2007 (1adult, kick; 11 larvae, kick; 1 larva, net), X-15-2007 (5 larvae, kick; 2 larvae, net).Mill Brook, III-27-2007 (5 adult, kick; 7 larvae, kick; 1 larva, net), IV-10-2007 (1adult, kick; 8 larvae, kick; 1 adult, net), IV-30-2007 (31 larvae, kick; 1 adult, net, 5larvae, net), V-16-2007 (16 adult, kick; 16 larvae, kick; 2 adult, net; 16 larvae, net),V-31-2007 (1 adult, kick; 11 larvae, kick; 1 larva, net), VI-18-2007 (13 larvae, kick;4 larvae, net), VII-9-2007 (7 larvae, kick), VII-25-2007 (11 larvae, kick; 1 larva,net), VIII-22-2007 (10 larvae, kick), IX-13-2007 (5 larvae, kick), X-15-2007 (5larvae, kick; 1 larva, net). Unnamed Priester’s Pond Feeder, V-16-2007 (1 larva,kick), VI-18-2007 (1 larva, kick), VII-25-2007 (1 larva, kick; 1 larva, net).[Southeastern mainland rivers]: Red Brook, IV-20-2007 (1 larva, kick).

Notes: Adults were collected on MV from March through October. Larvae werecollected on MV from March through September. A single larva was collected inSMA in April. Common on MV.

Haliplidae

Haliplus immaculicollis (Harris, 1828): [Martha’s Vineyard]: Mill Brook, VII-25-2007 (1adult, net), [Southeastern mainland rivers]: Red Brook, V-23-2007 (1 adult, net).

Notes: A single adult was collected on MV in July. A single adult was collected in SMAin May. There is no regional key to larvae.

*Peltodytes dunavani (Young, 1961): [Southeastern mainland rivers]: V-23-2007 (1adult, net).

Notes: A single adult was collected in SMA in May. There is no regional key to larvae.

Peltodytes duodecimpunctatus (Say, 1825): [Martha’s Vineyard]: Tiasquam River, IV-30-2007 (1 adult, net).

Notes: A single adult was collected on MV in April. There is no regional key to larvae.

Peltodytes muticus (LeConte, 1853): [Martha’s Vineyard]: Unnamed Priester’s PondFeeder, IV-10-2007 (2 adult, kick; 1 adult, net), V-16-2007 (1 adult, kick; 1 adult,net), V-31-2007 (2 adult, net), VI-18-2007 (1 adult, net). [Southeastern mainlandrivers]: Mashpee River, IV-20-2007 (1 adult, net), V-8-2007 (1 adult, net).

Notes: Adults were collected on MV from April through June and in SMA from April-May. There is no regional key to larvae.

*Peltodytes shermani (Roberts, 1913): [Martha’s Vineyard]: Mill Brook, VII-25-2007 (1adult, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (1 adult, net), IV-30-2007(1 adult, kick).


Notes: Adults collected on MV in April and July. There is no regional key to larvae.

Psephenidae

Ectopria nervosa (Melsheimer, 1845): [Martha’s Vineyard]: Roaring Brook, III-27-2007(1 larva, kick), VI-18-2007 (1 larva, kick).

Notes: Larvae were collected on MV in March and June.

DipteraChironomidae

**Polypedilum aviceps (Townes, 1945): [Martha’s Vineyard]: Tiasquam River, VI-21-2000 (3, kick). Mill Brook, VI-21-2000 (9, kick).

Notes: Collected on MV in June.

**Polypedilum flavum (Johannsen, 1905): [Martha’s Vineyard]: Mill Brook, VI-21-2000(3, kick).


**Cricotopus vierriensis (Goetghebuer, 1935): {Martha’s Vineyard]: Tiasquam River,VI-21-2000 (1, kick).

Notes: A single specimen was collected on MV in June.

Ptychopteridae

*Bittacomorpha clavipes (Fabricius, 1781): [Martha’s Vineyard]: Menemsha Hills Creek,IV-30-2007 (3, net), V-16-2007 (1, net), VI-15-2007 (1, net), VII-25-2007 (1, net).Tiasquam River, IV-1-2007 (3, net), IV-16-2007 (1, net). Unnamed Priester’s PondFeeder, V-31-2007 (2, kick), VII-9-2007 (1, kick). Witch Brook, III-17-2008 (1,kick).

Notes: Collected on MV from March-July.

Simuliidae

Prosimulium sp.: [Martha’s Vineyard]: Roaring Brook, III-13-2007 (4, net).Notes: Collected on MV in March. Too small to be identified.

*Simulium murmanum (Enderlein, 1935): [Martha’s Vineyard]: Roaring Brook, VII-25-2007 (4, kick).

Notes: Collected on MV in July.

*Simulium tuberosum (Lundstrom, 1911): [Southeastern mainland rivers]: MashpeeRiver, VI-15-2007 (6, kick).

Notes: Collected in SMA in June.

Simulium venustum complex: [Martha’s Vineyard]: Roaring Brook, VI-15-2007 (15,net). Tiasquam River, IV-10-2007 (2, kick), IV-30-2007 (10, net), V-16-2007 (4,kick), V-31-2007 (4, kick), VI-15-2007 (4, kick; 13, net), VII-9-2007 (10, net), VII-25-2007 (6, net). Mill Brook, VI-15-2007 (2, kick; 9, net). Unnamed Priester’s PondFeeder, IV-30-2007 (1, net). [Southeastern mainland rivers]: Red Brook, IX-28-2007 (10, kick).

Notes: Collected on MV from April-July and in SMA in September. Can only beseparated to species via preparation of the chromosomes.

441GREG D. WHITMORE

*Stegopterna mutata/diplomutata complex: [Martha’s Vineyard]: Mill Brook, III-27-2007 (15, net). Paint Mill Brook, III-17-2008 (7, net).

Notes: Collected on MV in March. Can only be separated to species via preparation ofthe chromosomes.

Ephemeroptera

Baetidae

*Acentrella parvula (McDunnough, 1932): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (5, kick), VI-18-2007 (4, kick), VII-9-2007 (36, kick), VII-27-2007 (9, kick),VIII-22-2007 (5, kick), IX-13-2007 (2, kick). Mill Brook, V-31-2007 (1, kick; 1,net), VI-18-2007 (19, kick). Unnamed Priester’s Pond Feeder, V-31-2007 (1, net).

Notes: Collected on MV from May-September.

Acerpenna pygmaea (Hagen, 1861): [Martha’s Vineyard]: Tiasquam River, III-27-2007(57, kick), IV-10-2007 (42, kick; 1, net), IV-30-2007 (225, kick; 63, net), V-16-2007(18, kick), V-31-2007 (18, kick), VI-18-2007 (100, kick; 1, net), VII-9-2007 (39,kick; 6, net), VII-25-2007 (15, kick; 6, net), VIII-22-2007 (24, kick), IX-13-2007(22, kick), X-15-2007 (3, kick; 2, net). Mill Brook, III-27-2007 (1, kick), IV-10-2007 (15, kick; 8, net), IV-30-2007 (6, kick; 5, net), VI-18-2007 (42, kick; 16, net),VII-25-2007 (2, kick), X-15-2007 (1, kick). Unnamed Priester’s Pond Feeder, VII-9-2007 (12, kick), VIII-22-2007 (1, net).

Notes: Collected on MV from March-October. High abundance in the Tiasquam River.

Baetis flavistriga (McDunnough, 1921): [Southeastern mainland rivers]: Mashpee River,V-23-2007 (40, kick; 3, net), VI-15-2007 (96, kick; 2, net).

Notes: Collected in SMA from May-June.

Baetis tricaudatus (Dodds, 1923): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (14,kick; 10, net), III-16-2007 (34, kick), III-27-2007 (57, kick), IV-10-2007 (43, kick),IV-30-2007 (13, kick; 3, net), V-16-2007 (2, kick; 2, net), V-31-2007 (1, kick), VI-18-2007 (155, kick; 6, net), VI-27-2007 (1, net), VII-9-2007 (60, kick; 3, net), VII-25-2007 (4, net), VIII-9-2007 (152, kick), VIII-22-2007 (129, kick), IX-13-2007(253, kick), X-15-2007 (62, kick; 1, net).

Notes: Collected on MV from March-October. High abundance in Roaring Brook.

*Callibaetis fluctuans (Walsh, 1862): [Martha’s Vineyard]: Mill Brook, III-27-2007 (1,net), IV-10-2007 (1, net), VII-25-2007 (4, net), VIII-22-2007 (5, net).

Notes: Collected on MV from March-April and from July-August.

*Centroptilum triangulifer (McDunnough, 1931): [Martha’s Vineyard]: UnnamedPriester’s Pond Feeder, IV-30-2007 (1, kick). Notes: A single specimen wascollected on MV in April.

*Heterocloeon amplum (Traver, 1932): [Martha’s Vineyard]: Tiasquam River, IX-13-2007 (1, kick), Mill Brook, V-16-2007 (22, kick).

Notes: Collected on MV in May and September.

*Heterocloeon anoka (Daggy, 1945): [Martha’s Vineyard]: Roaring Brook, V-31-2007(199, kick; 69, net), VI-18-2007 (85, kick; 36, net), VII-9-2007 (8, kick). TiasquamRiver, V-31-2007 (33, kick), X-15-2007 (3, kick).

Notes: Collected on MV from May-July and in October. High abundance in RoaringBrook.


*Plauditus cestus (Provonsha & McCafferty, 1982): [Martha’s Vineyard]: TiasquamRiver, V-31-2007 (1, net).

Notes: A single specimen was collected on MV in May.

*Plauditus cingulatus (McDunnough, 1931): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (6, kick; 7, net). Unnamed Priester’s Pond Feeder, VII-19-2007 (1, kick).

Notes: Collected on MV in May and July.

*Plauditus dubius (Walsh, 1862): [Martha’s Vineyard]: Roaring Brook, III-16-2007 (2,kick). Tiasquam River, V-16-2007 (35, kick), VIII-22-2007 (5, kick), IX-13-2007(8, kick). [Southeastern mainland rivers]: Mashpee River, (9, kick), IX-28-2007 (1,kick).

Notes: Collected on MV from March-September, and in SMA from March-May and fromAugust-September.

*Plauditus punctiventris (McDunnough, 1923): [Martha’s Vineyard]: Roaring Brook,VII-19-2007 (1, net).

Notes: A single specimen was collected on MV in July.

*Pseudocloeon propinquum (Walsh, 1863): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (19, net), IX-13-2007 (5, net). Mill Brook, V-31-2007 (1, net). UnnamedPriester’s Pond Feeder, V-31-2007 (16, kick), VI-18-2007 (2, kick; 26, net), VII-19-2007 (25, net), VII-25-2007 (1, net), VIII-22-2007 (1, net), IX-13-2007 (1, kick, 3,net).

Notes: Collected on MV from May-September.

Caenidae

*Caenis latipennis (Banks, 1907): [Martha’s Vineyard]: Mill Brook, III-27-2007 (1, kick;1, net), IV-30-2007 (1, net), V-16-2007 (3, kick), V-31-2007 (3, kick; 8, net), VI-18-2007 (12, kick; 9, net), VII-25-2007 (7, net), VIII-22-2007 (1, net).

Notes: Collected on MV from March-August.

Ephemerellidae

Ephemerella invaria (Walker, 1853): [Southeastern mainland rivers]: Mashpee River,III-28-2007 (140, kick; 40, net), IV-20-2007 (1600, kick; 100, net), V-8-2007 (200,kick; 40, net), V-23-2007 (183, kick; 72, net), VI-15-2007 (15, kick; 3, net), IX-28-2007 (3, kick).

Notes: Collected in SMA from March-June and in September. Highest abundance of anyspecies collected.

Ephemerella subvaria (McDunnough, 1931): [Southeastern mainland rivers]: MashpeeRiver, III-28-2007 (1, kick; 1, net).

Notes: Collected in SMA in March.

Eurylophella funeralis (McDunnough, 1925): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (1, kick; 23, net), III-27-2007 (25, net), IV-10-2007 (1, kick), VI-27-2007(2, net), VII-9-2007 (1, kick), VII-25-2007 (74, net), VIII-22-2007 (8, kick), IX-13-2007 (6, kick; 2, net), X-15-2007 (1, net). Unnamed Priester’s Pond Feeder, VIII-22-2007 (58, net). [Southeastern mainland rivers]: Mashpee River, V-8-2007 (4,net).

Notes: Collected on MV from March-October and in SMA in May.

443GREG D. WHITMORE

Eurylophella minimella (McDunnough, 1931): [Southeastern mainland rivers]: MashpeeRiver, V-23-2007 (1, net).

Notes: A single specimen was collected in SMA in May.

Eurylophella verisimilis (McDunnough, 1930): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (4, kick; 48, net), IV-10-2007 (42, net), IV-30-2007 (30, net), V-16-2007(1, kick; 20, net), V-31-2007 (3, kick; 1, net), VIII-9-2007 (2, kick), X-15-2007 (1,kick). Tiasquam River, III-27-2007 (85, kick; 64, net), IV-10-2007 (99, kick), IV-30-2007 (9 kick; 125, net), V-16-2007 (9, kick; 80, net), V-31-2007 (1, kick; 20,net), VI-18-2007 (3, net). Mill Brook, III-27-2007 (13, kick; 3, net), IV-10-2007 (7,kick; 1, net), IV-30-2007 (4 kick; 3, net), V-16-2007 (13, kick; 2, net), V-31-2007(4, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (33, kick; 5, net), IV-30-2007(27, kick; 9, net), V-16-2007 (31, kick; 8, net), V-31-2007 (5, kick; 12, net), VI-18-2007 (10, kick; 9, net). Paint Mill Brook, IV-1-2008 (1, net). [Southeasternmainland rivers]: Mashpee River, III-28-2007 (8, net), IV-20-2007 (8, net), V-23-2007 (42, net), VI-15-2007 (3, net).

Notes: Collected on MV from March-August and in October, and in SMA from March-June. Common on MV.

Serratella deficiens (Morgan, 1911): [Southeastern mainland rivers]: Mashpee River, III-28-2007 (34, kick).

Notes: Collected in SMA in March.

Heptageniidae

Maccaffertium modestum (Banks, 1910): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (47, kick; 6, net), IV-10-2007 (11, kick), IV-30-2007 (20, kick; 1, net), V-16-2007 (28 kick; 2, net), V-31-2007 (12, kick; 3, net), VI-18-2007 (2, net), VII-9-2007(11, kick; 3, net), VII-25-2007 (10, kick; 1, net), VIII-22-2007 (17, kick; 4, net), IX-13-2007 (16, kick), X-15-2007 (8, kick; 2, net). Mill Brook, III-27-2007 (76, kick;17, net), IV-10-2007 (40, kick; 15, net), IV-30-2007 (46, kick; 6, net), V-16-2007(53, kick; 16, net), V-31-2007 (26, kick; 12, net), VI-18-2007 (11, kick; 3, net), VII-9-2007 (11, kick), VII-25-2007 (21, kick; 11, net), VIII-22-2007 (11, kick; 2, net),IX-13-2007 (1, kick; 9, net), X-15-2007 (1, kick; 4, net). Unnamed Priester’s PondFeeder, IV-30-2007 (1, kick; 1, net), IX-13-2007 (4, kick). [Southeastern mainlandrivers]: Mashpee River, III-28-2007 (13, kick; 8, net), IV-10-2007 (140 kick; 13,net), V-8-2007 (16, kick; 13, net), V-23-2007 (14, kick; 20, net), VI-15-2007 (24,kick; 13, net), VIII-13-2007 (25, kick; 4, net), IX-28-2007 (12, kick; 1, net).

Notes: Collected on MV from March-October and in SMA from March-September.Common on MV and in Mashpee River.

Leptophlebiidae

*Leptophlebia cupida (Say, 1823): [Martha’s Vineyard]: Roaring Brook, IV-30-2007 (1,net). Tiasquam River, IV-30-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (22, net), IV-30-2007 (4, kick: 23, net), V-16-2007 (6, net), IX-13-2007 (1,net). [Southeastern mainland rivers]: Mashpee River, V-8-2007 (3, net).

Notes: Collected on MV from April-May and in SMA in May.

*Leptophlebia intermedia (Traver, 1932): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (29, net), III-27-2007 (1, kick; 74, net), IV-10-2007 (27, net). UnnamedPriester’s Pond Feeder, IV-10-2007 (4, net), X-15-2007 (2, kick; 12, net). WitchBrook, III-17-2008 (1, kick). Paint Mill Brook, X-15-2007 (5, net), III-17-2008 (14,net). [Southeastern mainland rivers]: Mashpee River, IV-20-2007 (9, net).


Notes: Collected on MV from March-April and in October, and in SMA in April. Highabundance in Roaring Brook.

*Leptophlebia johnsoni (McDunnough, 1924): [Martha’s Vineyard]: Roaring Brook, III-16-2007 (1, kick), V-16-2007 (8, net), VI-18-2007 (2, net), VII-9-2007 (23, net).Unnamed Priester’s Pond Feeder, IV-30-2007 (2, kick; 32, net), V-16-2007 (1, kick;22, net), V-31-2007 (18, kick), VI-18-2007 (3, net), VIII-22-2007 (3, kick).


*Leptophlebia nebulosa (Walker, 1853): [Martha’s Vineyard]: Roaring Brook, IV-30-2007 (5, net), V-16-2007 (3, net). Tiasquam River, III-27-2007 (3, net), IV-10-2007(1, kick; 1, net), IX-13-2007 (1, kick). Mill Brook, III-27-2007 (4, net), IV-10-2007(2, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (3, kick). [Southeasternmainland rivers]: Mashpee River, III-28-2007 (10, net).

Notes: Collected on MV from March-May and in October, and in SMA in March.

Paraleptophlebia debilis (Walker, 1853): [Martha’s Vineyard]: Roaring Brook, VI-18-2007 (1, kick), VI-27-2007 (35, net), VII-9-2007 (79, kick; 17, net), VII-25-2007(3, kick; 4, net), VIII-9-2007 (58, kick; 29, net), VIII-22-2007 (130, kick), IX-13-2007 (75, kick; 27, net), X-15-2007 (8, kick; 29, net). Unnamed Priester’s PondFeeder, IV-10-2007 (2, kick).

Notes: Collected on MV from April-October. High abundance in Roaring Brook.

*Paraleptophlebia strigula (McDunnough, 1932): [Martha’s Vineyard]: Roaring Brook,IV-10-2007 (3, kick). Tiasquam River, III-27-2007 (1, kick), IV-10-2007 (2, kick).

Notes: Collected on MV from March-April.

*Paraleptophlebia volitans (McDunnough, 1924): [Martha’s Vineyard]: Roaring Brook,III-13-2007 (1, kick; 1, net), III-27-2007 (4, kick).

Notes: Collected on MV in March.

Hemiptera

Belostomatidae

Belostoma flumineum (Say, 1823): [Martha’s Vineyard]: Roaring Brook, III-3-2007 (1,net). Unnamed Priester’s Pond Feeder, IX-13-2007 (1, net), X-15-2007 (1, net).

Notes: Collected on MV in March and from September-October.

*Belostoma lutarium (Stal, 1855): [Martha’s Vineyard]: Mill Brook, VII-25-2007 (6,net), VIII-22/2007 (2, net), IX-13-2007 (1, net).

Notes: Collected on MV from July-August.

Hebridae

*Hebrus buenoi (Drake & Harris, 1943): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (1, kick).


445GREG D. WHITMORE

Mesoveliidae

Mesovelia mulsanti (White, 1879): [Martha’s Vineyard]: Tiasquam River, IX-13-2007 (1,net). Mill Brook, VII-25-2007 (1, net), IX-13-2007 (1, net).

Notes: Collected on MV in July and September.

Naucoridae

Pelocoris femoratus (Palisot de Boisvois, 1820): [Martha’s Vineyard]: Mill Brook, V-31-2007 (1, net), VII-25-2007 (3, net), VIII-22-2007 (1, net).

Notes: Collected on MV in May and from July-August.

Notonectidae

*Notonecta irrorata (Uhler, 1879): [Martha’s Vineyard]: Menemsha Hills Creek, IV-30-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (1, net), V-16-2007 (2,net), VII-9-2007 (1, net), X-15-2007, (1, net). Witch Brook, III-17-2008 (1, kick).

Notes: Collected on MV from March-July and in October.

Notonecta lunata (Hungerford, 1926): [Martha’s Vineyard]: Mill Brook, VIII-22-2007(4, net), IX-13-2007 (2, net).

Notes: Collected on MV from August-September.

*Notonecta undulata (Say, 1832): [Southeastern mainland rivers]: Red Brook, V-8-2007(1, net). Notes: A single specimen was collected in SMA in May.

Pleidae

Neoplea striola (Fieber, 1844): [Martha’s Vineyard]: Menemsha Hills Creek, V-31-2007(1, net). Unnamed Priester’s Pond Feeder, V-31-2007 (2, kick; 1, net), VI-18-2007(2, net).

Notes: Collected on MV from May-June.

Veliidae

*Microvelia americana (Uhler, 1884): [Martha’s Vineyard]: Menemsha Hills Creek, V-31-2007 (2, net). Unnamed Priester’s Pond Feeder, V-31-2007 (2, kick), VII-25-2007 (3, net), VIII-22-2007 (1, kick), IX-13-2007 (4, net). Paint Mill Brook, IV-1-2008 (1, net).

Notes: Collected on MV from April-September.

*Microvelia buenoi (Drake, 1920): [Martha’s Vineyard]: Unnamed Priester’s PondFeeder, VII-9-2007 (1, net), VII-25-2007 (1, net), VIII-22-2007 (1, net), IX-13-2007 (2, net).

Notes: Collected on MV from July-September.

Megaloptera

Corydalidae

Chauliodes pectinicornis (Linnaeus, 1763): [Martha’s Vineyard]: Tiasquam River, IV-10-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-30-2007 (1, net), VIII-22-2007(1, kick).

Notes: Collected on MV in April and August.


*Chauliodes rastricornis (Rambur, 1842): [Martha’s Vineyard]: Mill Brook, VII-25-2007(1, net). Unnamed Priester’s Pond Feeder, V-16-2007 (1, net), VII-9-2007 (1, net),VII-25-2007 (1, kick).

Notes: Collected on MV in May and July.

*Nigronia serricornis (Say, 1824): [Southeastern mainland rivers]: Mashpee River, III-28-2007 (3, kick), VIII-13-2007 (6, kick), IX-28-2007 (5, kick).

Notes: Collected in SMA in March and from August-September.

Sialidae

*Sialis iola (Ross, 1937): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (3, net).Unnamed Priester’s Pond Feeder, IV-10-2007 (8, kick), IV-30-2007 (3, kick), V-31-2007 (2, kick; 1, net), X-15-2007 (3, net). Witch Brook, III-17-2008 (2, kick). PaintMill Brook, X-15-2007 (1, net), IV-1-2008 (1, net). [Southeastern mainland rivers]:Red Brook Feeder, III-28-2007 (2, net), IV-20-2007 (11, net), V-8-2007 (4, net), V-23-2007 (8, net), VI-15-2007 (3, net), VIII-16-2007 (7, net), VIII-13-2007 (2, net),IX-28-2007 (5, net).

Notes: Collected on MV in March and from May-October, and in SMA from March-August.

*Sialis velata (Ross, 1937): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (10, net),X-15-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (1, net).

Notes: Collected on MV from March-April and in October.

Odonata

Aeshnidae

Aeshna umbrosa (Walker, 1908): [Martha’s Vineyard]: Menemsha Hills Creek, V-16-2007 (1, net). Unnamed Priester’s Pond Feeder, V-31-2007 (1, kick).


Anax junius (Drury, 1773): [Martha’s Vineyard]: Mill Brook, IX-13-2007 (1, net).Notes: A single specimen was collected on MV in September.

Anax longipes (Hagen, 1861): [Martha’s Vineyard]: Mill Brook, VIII-22-2007 (1, net).Notes: A single specimen was collected on MV in August.

Basiaeschna janata (Say, 1839): [Southeastern mainland rivers]: Mashpee River, V-23-2007 (1, net).

Notes: A single specimen was collected in SMA in May.

Boyeria vinosa (Say, 1839): [Martha’s Vineyard]: Roaring Brook, VIII-22-2007 (1, kick).Menemsha Hills Creek, IV-30-2007 (1, net). Tiasquam River, III-27-2007 (1, kick),V-16-2007 (1, kick; 1, net), V-31-2007 (2, net), VI-19-2007 (2, net), VII-9-2007 (5,net), VIII-22-2007 (1, net). Mill Brook, IV-10-2007 (1, net), V-31-2007 (1, net).Unnamed Priester’s Pond Feeder, IV-30-2007 (2, net), VI-18-2007 (2, net), VII-25-2007 (5, net), VIII-22-2007 (8, net), IX-13-2007 (1, net), X-15-2007 (3, net).[Southeastern mainland rivers]: Mashpee River, IV-20-2007 (1, net), V-23-2007 (4,net), VI-15-2007 (1, kick; 2, net), IX-28-2007 (2, kick). Red Brook Feeder, III-28-2007 (1, net).

Notes: Collected on MV from March-October, and in SMA from March-June and inSeptember. Most commonly collected dragonfly.

447GREG D. WHITMORE

Calopterygidae

Calopteryx maculata (Beauvois, 1805): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (1, net), IV-30-2007 (1, kick), V-31-2007 (1, kick; 3, net), VI-19-2007 (2, net),IX-13-2007 (1, net), X-15-2007 (2, net). Mill Brook, III-27-2007 (4, net), IV-10-2007 (1, net), V-31-2007 (3, net), X-15-2007 (1, kick). Unnamed Priester’s PondFeeder, V-31-2007 (2, kick), VII-9-2007 (1, net), IX-13-2007 (1, kick; 1, net), X-15-2007 (1 kick; 1, net). [Southeastern mainland rivers]: Mashpee River, IV-20-2007 (1, net), V-8-2007 (1, net), V-23-2007 (8, net), VI-15-2007 (3, net), VIII-13-2007 (4, net), IX-28-2007 (4, net).

Notes: Collected on MV from March-October and in SMA from April-September. Mostcommonly collected damselfly.

Coenagrionidae

Argia fumipennis violacea (Walker, 1853): [Martha’s Vineyard]: Tiasquam River, V-16-2007 (1, net), V-31-2007 (2, net), VI-19-2007 (2, net), VII-9-2007 (2, net), X-15-2007 (1, kick).

Notes: Collected on MV from May-July and in October.

Chromagrion conditum (Hagen, 1876): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (1, net).

Notes: A single specimen was collected on MV in March.

Enallagma divagans (Selys, 1876): [Martha’s Vineyard]: Mill Brook, IV-30-2007 (5,net).

Notes: Collected on MV in April.

Enallagma durum (Hagen, 1861): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (1,net), VI-19-2007 (1, net). Mill Brook, IV-10-2007 (1, net), IV-30-2007 (5, net), V-31-2007 (11, net), VI-18-2007 (3, kick; 8, net), VII-25-2007 (13, net), VIII-22-2007(8, net), IX-13-2007 (29, net), X-15-2007 (20, net). Unnamed Priester’s PondFeeder, X-15-2007 (1, net).

Notes: Collected on MV from April-October.

Enallagma geminatum (Kellicott, 1895): [Martha’s Vineyard]: Tiasquam River, IX-13-2007 (2, net).

Notes: Collected on MV in September.

Cordulegasteridae

*Tetragoneuria spinosa (Hagen, 1878): [Martha’s Vineyard]: Mill Brook, V-16-2007 (1,kick), VIII-22-2007 (1, net).

Notes: Collected on MV in May and August.

Libellulidae

Ladona julia (Uhler, 1857): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (1, net).Notes: A single specimen was collected on MV in May.


Plecoptera

Capniidae

Allocapnia recta (Claassen, 1924): [South-eastern rivers]: Red Brook Feeder, VIII-13-2007 (1, net).

Notes: A single specimen was collected in SMA in August.

Paracapnia angulata (Hanson, 1961): [Martha’s Vineyard]: Paint Mill Brook, III-17-2008 (1, kick).


*Paracapnia opis (Newman, 1839): [Martha’s Vineyard]: Roaring Brook, I-16-2008 (1,kick).

Notes: A single specimen was collected on MV in January.

Leuctridae

*Leuctra duplicata (Claassen, 1923): [Martha’s Vineyard]: Menemsha Hills Creek, VI-18-2007 (8, net).


Leuctra ferruginea (Walker, 1851): [Martha’s Vineyard]: Roaring Brook, III-13-2007(13, kick), III-27-2007 (34, kick), IV-30-2007 (7, kick; 2,net), V-16-2007 (47, kick),V-31-2007 (68, kick; 5, net), VI-18-2007 (27 kick; 18, net), VII-9-2007 (34, kick),VII-25-2007 (13, kick; 7, net), VIII-9-2007 (44, kick), VIII-22-2007 (60, kick), XI-13-2007 (8, kick). Menemsha Hills Creek, IV-30-2007 (103, net), V-16-2007 (43,net), V-31-2007 (89, net). Tiasquam River, III-27-2007 (1, kick). Paint Mill Brook,IV-1-2008 (2, net). [Southeastern mainland rivers]: Mashpee River, III-28-2007 (2,kick), IV-20-2007 (30, kick; 1, net), V-8-2007 (1, kick), VI-17-2007 (1, kick), IX-28-2007 (5, kick). Red Brook, IV-20-2007 (2, kick). Red Brook Feeder, VII-16-2007 (9, net), IX-28-2007 (3, net).

Notes: Collected on MV from March-September, and in SMA from March-September.Most commonly collected stonefly.

*Leuctra maria (Hanson, 1941): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (5,net), III-29-2007 (34, net), IV-10-2007 (20, net), IV-30-2007 (1, net). [Southeasternmainland rivers]: III-28-2007 (9, kick), IV-20-2007 (1, net), V-8-2007 (1, kick).Red Brook Feeder, III-28-2007 (2, net), IV-20-2007 (3, net).

Notes: Collected on MV from March-April and in SMA from March-May.

Leuctra tenuis (Pictet, 1841): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (58, net),IV-10-2007 (55, kick; 45, net). Menemsha Hills Creek, III-19-2007 (75, net), IV-10-2007 (90, net). Tiasquam River, IV-10-2007 (1, net). Paint Mill Brook, III-17-2008(25, kick), IV-1-2008 (16, kick). [Southeastern mainland rivers]: Mashpee River,III-28-2007 (2, net), V-23-2007 (1, kick). Red Brook Feeder, V-18-2007 (10, net).

Notes: Collected on MV in March-April and in SMA from March-May. Collected in highabundance on MV.

Leuctra variabilis (Hanson, 1941): [Martha’s Vineyard]: Roaring Brook, IX-13-2007 (2,kick). Mill Brook, VII-25-2007 (1, net). Paint Mill Brook, X-15-2007 (23, net).[Southeastern mainland rivers]: Mashpee River, VIII-13-2007 (2, kick). Red BrookFeeder, VI-15-2007 (1, net).

449GREG D. WHITMORE

Notes: Collected on MV from August-September and in SMA from June-August.

Paraleuctra sara (Pictet, 1841): [Martha’s Vineyard]: Witch Brook, III-17-2008 (4, kick).Notes: Collected on MV in March.

Nemouridae

Amphinemura wui (Claassen, 1936): [Martha’s Vineyard]: Roaring brook, III-13-2007(1, kick), III-27-2007 (6, kick), IV-10-2007 (20, kick), IV-30-2007 (23, kick; 3, net),V-16-2007 (18, kick; 1, net), V-31-2007 (73, kick; 15, net), VI-18-2007 (41, kick;25, net), VII-9-2007 (10, kick), VII-25-2007 (25, kick; 23, net), VIII-9-2007 (8,kick), VIII-22-2007 (3, kick).


Ostrocerca truncata (Claassen, 1923): [Martha’s Vineyard]: Paint Mill Brook, III-17-2008 (1, kick).


*Podmosta macdunnoughi (Ricker, 1947): [Martha’s Vineyard]: Roaring brook, III-13-2007 (1, kick). Paint Mill Brook, IV-1-2008 (2, net).


Prostoia completa (Walker, 1852): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (1,net). Tiasquam River, III-27-2007 (6, kick; 2, net). Mill Brook, III-27-2007 (1,kick). [Southeastern mainland rivers]: Mashpee River, III-28-2007 (17, kick; 3,net).

Notes: Collected on MV and in SMA in March.

Soyedina vallicularia/washingtoni complex: [Martha’s Vineyard]: Roaring Brook, III-27-2007 (1, net). Paint Mill Brook, III-17-2008 (1, net), IV-1-2008 (2, net).[Southeastern mainland rivers]: Red Brook, III-28-2007 (1, net).

Notes: Collected on MV from March-April and in SMA in March. Larvae of thesespecies cannot be separated with current keys.

Perlodidae

Isoperla bilineata (Say, 1823): [Southeastern mainland rivers]: Mashpee River, VIII-13-2007 (4, kick; 1, net), IX-28-2007 (7, kick).

Notes: Collected in SMA from August-September.

*Isoperla frisoni (Illies, 1966): [Southeastern mainland rivers]: Mashpee River, IV-20-2007 (16, kick), V-8-2007 (1, net), V-23-2007 (23, kick; 3, net), VI-15-2007 (19,kick).

Notes: Collected in SMA from April-June.

Isoperla lata (Frison, 1942): [Southeastern mainland rivers]: Mashpee River, IV-20-2007(1, net), V-8-2007 (16, kick).

Notes: Collected in SMA from April-May.

*Isoperla marlynia (Needham & Claassen, 1925): [Martha’s Vineyard]: Roaring brook,V-31-2007 (1, kick), VII-25-2007 (2, net), VIII-22-2007 (10, kick), IX-13-2007 (27,kick), X-15-2007 (37, kick). Paint Mill Brook, X-15-2007 (6, net).

Notes: Collected on MV from May-October.


Isoperla similis (Hagen, 1861): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (4,kick), III-27-2007 (9, kick; 4, net), IV-10-2007 (3, kick; 2, net). Paint Mill Brook,III-17-2008 (5, kick), IV-1-2008 (2, kick).


Isoperla signata (Banks, 1902): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (10,net).


*Isoperla transmarina (Newman, 1838): [Southeastern mainland rivers]: MashpeeRiver, III-28-2007 (35, kick; 2, net), IV-20-2007 (15, kick; 2, net).

Notes: Collected in SMA from March-April.

Taeniopterygidae

*Taeniopteryx burksi (Ricker & Ross, 1968): [Martha’s Vineyard]: Roaring Brook, I-16-2008 (1, kick).

Notes: A single specimen was collected on MV in January.

Taeniopteryx maura (Pictet, 1841): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (3,kick; 3, net), III-27-2007 (1, net). [Southeastern mainland rivers]: Red Brook, III-28-2007 (1, kick).

Notes: Collected on MV and in SMA in March.

Taeniopteryx nivalis (Fitch, 1847): [Martha’s Vineyard]: Roaring Brook, X-15-2007 (2,kick).

Notes: Collected on MV in October.

Trichoptera

Brachycentridae

Brachycentrus americanus (Banks, 1899): [Martha’s Vineyard]: Mill Brook, IV-10-2007(1, net).

Notes: A single specimen was collected on MV in April.

Glossomatidae

Agapetus sp.: [Martha’s Vineyard]: Roaring Brook, IV-30-2007 (1, kick).Notes: A single specimen was collected on MV in April. Larvae cannot be identified to

species using current keys.

Glossosoma nigrior (Banks, 1911): [Southeastern mainland rivers]: Mashpee River, IV-20-2007 (1, kick), V-8-2007 (3, kick), V-23-2007 (2, kick), VI-15-2007 (3, kick; 4,net), VIII-13-2007 (1, kick), IX-28-2007 (2, kick).

Notes: Collected in SMA from April-September.

Goeridae

Goera calcarata (Banks, 1899): [Martha’s Vineyard]: Unnamed Priester’s Pond Feeder,VIII-22-2007 (7, kick; 2, net). [Southeastern mainland rivers]: Mashpee River, VI-15-2007 (1, kick; 1, net), IX-28-2007 (1, kick).

Notes: Collected on MV in August, and in SMA in June and September.

451GREG D. WHITMORE

Hydropsychidae

Ceratopsyche sparna (Ross, 1938): [Martha’s Vineyard]: Roaring Brook, IV-10-2007(11, kick), V-16-2007 (11, kick), VII-25-2007 (18, net), VIII-9-2007 (18, kick),VIII-22-2007 (26, kick), IX-13-2007 (19, kick), X-15-2007 (24, kick). TiasquamRiver, V-16-2007 (3, kick; 1, net), VII-9-2007 (8, kick), VIII-22-2007 (4, kick), X-15-2007 (2, kick; 7, net). Mill Brook, IV-10-2007 (2, net), VI-15-2007 (21, kick),VII-25-2007 (2, net), IX-13-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-30-2007 (1, net), VII-25-2007 (7, kick), VIII-22-2007 (1, kick). [Southeasternmainland rivers]: Mashpee River, III-28-2007 (25, kick; 1, net), IV-20-2007 (2,net), V-8-2007 (41, kick; 2, net), V-23-2007 (14, kick; 1, net), VI-15-2007 (18, kick;3, net), VIII-13-2007 (13, kick), IX-28-2007 (5, kick).

Notes: Collected on MV from April-October and in SMA from March-September.Common on MV and in Mashpee River.

Cheumatopsyche sp.: [Martha’s Vineyard]: Tiasquam River, III-27-2007 (18, kick; 11,net), IV-10-2007 (21, kick), IV-30-2007 (15, kick; 3, net), V-16-2007 (28, kick), V-31-2007 (16, kick; 2, net), VI-15-2007 (4, kick; 9, net), VII-9-2007 (5, kick), VII-25-2007 (5, kick; 1, net), VIII-22-2007 (5, kick; 5, net), IX-13-2007 (16, kick), X-15-2007 (21, kick; 4, net). Mill Brook, III-27-2007 (11, kick; 13, net), IV-10-2007(16, kick; 18, net), IV-30-2007 (7, kick; 8, net), V-16-2007 (9, kick; 14, net), V-31-2007 (1, kick; 1, net), VI-15-2007 (8, kick; 9, net), VII-9-2007 (15, kick), VII-25-2007 (31, kick; 15, net), VIII-22-2007 (39, kick; 5, net), IX-13-2007 (22, kick; 16,net), X-15-2007 (17, kick; 17, net). Unnamed Priester’s Pond Feeder, V-16-2007 (1,net), V-31-2007 (1, net), VII-25-2007 (5, kick), VIII-22-2007 (17, kick), IX-13-2007 (3, kick). [Southeastern mainland rivers]: Mashpee River, VI-17-2007 (2,kick).

Notes: Collected on MV from March-October and in SMA in June. Larvae cannot beidentified to species using current keys. Common on MV.

Diplectrona modesta (Banks, 1908): [Martha’s Vineyard]: Roaring Brook, III-13-2007,(9, kick; 13, net), III-27-2007 (13, kick; 7, net), IV-10-2007 (6, net), IV-30-2007 (5,kick; 1, net), V-31-2007 (19, kick; 4, net), VI-15-2007 (8, kick; 4, net), VIII-25-2007 (3, kick), IX-13-2007 (13, kick), X-15-2007 (5, kick). Unnamed Priester’sPond Feeder, IV-30-2007 (1, kick). Paint Mill Brook, X-15-2007 (16, net), III-17-2008, 3, kick), IV-1-2008 (10, kick).

Notes: Collected on MV from March-October.

Hydropsyche betteni (Ross, 1938): [Martha’s Vineyard]: Roaring Brook, VII-9-2007 (3,kick). Tiasquam River, III-27-2007 (3, kick; 2, net), IV-10-2007 (2, kick; 1, net), IV-30-2007 (8, kick; 1, net), V-16-2007 (20, kick; 1, net), V-31-2007 (19, kick), VI-15-2007 (18, kick), VII-9-2007 (10, kick; 6, net), VII-25-2007 (22, kick; 6, net), IX-13-2007 (2, kick), X-15-2007 (1, kick). Mill Brook, III-27-2007 (15, kick; 9, net), IV-10-2007 (10, kick; 5, net), IV-30-2007 (23, kick; 3, net), V-16-2007 (18, kick; 14,net), V-31-2007 (59, kick; 2, net), VI-15-2007 (3, kick), VII-9-2007 (9, kick), VII-25-2007 (17, kick), VIII-22-2007 (7, kick), IX-13-2007 (9, kick), X-15-2007 (5,kick; 3, net). Unnamed Priester’s Pond Feeder, VII-9-2007 (2, kick), VII-25-2007(12, kick), VIII-22-2007 (1, net), IX-13-2007 (3, kick). [Southeastern mainlandrivers]: Mashpee River, III-28-2007 (28, kick), IV-20-2007 (29, kick), V-8-2007(15, kick), V-23-2007 (4, kick), VI-15-2007 (10, kick; 5, net). VIII-13-2007 (1, net),IX-28-2007 (7, kick).

Notes: Collected on MV from March-October. Common on MV.


Hydroptilidae

Dibusa angata (Ross, 1939): [Martha’s Vineyard]: Roaring Brook, V-31-2007 (1, kick),VI-18-2007 (1, kick).


Lepidostomatidae

Lepidostoma sommermanae (Ross, 1946): [Martha’s Vineyard]: Tiasquam River, IV-30-2007 (2, kick), V-31-2007 (1, kick).

Notes: Collected on MV from April-May.

Lepidostoma sp.: [Martha’s Vineyard]: Roaring Brook, III-13-2007 (4, net), III-27-2007(8, net), IV-10-2007 (4, net), IV-30-2007 (2, net), V-16-2007 (1, net), V-31-2007 (6,kick; 5, net), VI-15-2007 (1, kick; 19, net), VII-9-2007 (1, kick; 5, net), VII-25-2007 (5, kick; 9, net), VIII-9-2007 (3, kick). Tiasquam River, III-27-2007 (3, kick;7, net), IV-10-2007 (2, kick), IV-30-2007 (9, kick), V-16-2007 (5, kick), V-31-2007(2, net), VI-15-2007 (5, kick; 6, net), VII-9-2007 (2, kick; 2, net), VII-25-2007 (10,kick), VIII-22-2007 (1, kick; 6, net), X-15-2007 (8, kick; 2, net). UnnamedPriester’s Pond Feeder, X-15-2007 (1, net). Paint Mill Brook, III-17-2008 (1, kick).[Southeastern mainland rivers]: Mashpee River, V-8-2007 (1, kick; 7, net), V-23-2007 (12, net), VI-15-2007 (2, kick; 11, net), VIII-13-2007 (2, kick). Red Brook,IV-20-2007 (1, net), V-8-2007 (1, net). Red Brook Feeder, V-8-2007 (9, net), VI-15-2007 (10, net), VII-16-2007 (3, net), VIII-13-2007 (3, net).

Notes: Collected on MV from March-October and in SMA from April-August. Commonon MV and in SMA. Larvae cannot be identified to species using current keys.

Leptoceridae

*Ceraclea alagma (Ross, 1938): [Martha’s Vineyard]: Mill Brook, V-16-2007 (2, net).Notes: Collected on MV in May.

*Ceraclea alces (Ross, 1941): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (2, kick),VI-15-2007 (1, kick).


Ceraclea tarsipunctata (Vorhies, 1909): [Martha’s Vineyard]: Mill Brook, V-31-2007 (2,net), VI-15-2007 (2, kick).


Mystacides longicornis (Linnaeus, 1758): [Martha’s Vineyard]: Roaring Brook, IV-10-2007 (1, net), IV-30-2007 (1, net), X-15-2007 (1, net). Tiasquam River, VII-9-2007(2, net). Mill Brook, V-16-2007 (1, net), VII-25-2007 (3, net). Unnamed Priester’sPond Feeder, V-16-2007 (1, kick), VIII-22-2007 (1, net), X-13-2007 (1, net).[Southeastern mainland rivers]: Mashpee River, VIII-13-2007 (1, net). Red BrookFeeder, V-23-2007 (1, net).

Notes: Collected on MV from April-October, and in SMA in May and August.

Mystacides sepulchralis (Walker, 1852): [Martha’s Vineyard]: Tiasquam River, X-15-2007 (1, kick). Unnamed Priester’s Pond Feeder, IV-10-2007 (1, kick), V-16-2007(1, net), IX-13-2007 (2, kick), X-15-2007 (1, net). [Southeastern mainland rivers]:Mashpee River, V-23-2007 (4, net).

Notes: Collected on MV from April-May and from September-October, and in SMA inMay.

453GREG D. WHITMORE

Nectopsyche exquisita (Walker, 1852): [Martha’s Vineyard]: Tiasquam River, III-27-2007(1, net).


Oecetis avara (Banks, 1895): [Martha’s Vineyard]: Tiasquam River, V-31-2007 (1, net).Mill Brook, V-31-2007 (2, kick; 2, net).


Oecetis cinerascens (Hagen, 1861): [Martha’s Vineyard]: Tiasquam River, IV-30-2007(1, kick).

Notes: A single specimen was collected on MV in April.

Oecetis osteni (Milne, 1934): [Martha’s Vineyard]: Tiasquam River, VII-9-2007 (4, net).Notes: Collected on MV in July.

Oecetis persimilis (Banks, 1907): [Martha’s Vineyard]: Mill Brook, V-31-2007 (2, kick).Notes: Collected on MV in May.

Triaenodes ignitus (Walker, 1852): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (3,net), V-16-2007 (3, net), VII-9-2007 (2, kick; 2, net). Mill Brook, III-27-2007 (1,net). [Southeastern mainland rivers]: Mashpee River, VIII-13-2007 (1, net), IX-28-2007 (1, net). Red Brook, V-23-2007 (1, kick).

Notes: Collected on MV from March-July, and in SMA in May and from August-September.

Limnephillidae

Anabolia consocia (Walker, 1852): [Martha’s Vineyard]: Unnamed Priester’s PondFeeder, IV-30-2007 (4, kick), V-16-2007 (2, net). [Southeastern mainland rivers]:Red Brook, III-28-2007 (1, net).

Notes: Collected on MV from April-May and in SMA in March.

Hydatophylax argus (Harris, 1869): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (5,net), III-27-2007 (4, net), VI-27-2007 (4, net), IX-13-2007 (3, net), X-15-2007 (1,kick; 4, net). Tiasquam River, X-15-2007 (1, net). Mill Brook, V-16-2007 (2, kick).Unnamed Priester’s Pond Feeder, IV-30-2007 (1, kick), V-31-2007 (2, net), VIII-22-2007 (1, kick; 2, net), IX-13-2007 (2, net). Paint Mill Brook, III-17-2008 (1, kick;4, net). [Southeastern mainland rivers]: Mashpee River, III-28-2007 (1, net), VIII-13-2007 (1, net), IX-28-2007 (12, net). Red Brook, IV-20-2007 (2, net).

Notes: Collected on MV from March-October, and in SMA from March-April and fromAugust-September.

Ironoquia punctatissima (Walker, 1852): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (3, net), IV-10-2007 (1, kick). Mill Brook, IV-10-2007 (2, net). UnnamedPriester’s Pond Feeder, IV-10-2007 (9, net), IV-30-2007 (1, net), V-16-2007 (2, net).[Southeastern mainland rivers]: Red Brook, V-8-2007 (1, net).

Notes: Collected on MV from March-May and in SMA in May.

Limnephilus sp.: [Southeastern mainland rivers]: Red Brook, III-28-2007 (1, kick).Notes: A single specimen was collected in SMA in March. Larvae cannot be identified to

species using current keys.


Onocosmoecus unicolor (Banks, 1897): [Martha’s Vineyard]: Roaring Brook, V-16-2007(1, net), VI-15-2007 (1, net), VIII-9-2007 (2, net). Tiasquam River, V-16-2007 (14,net), V-31-2007 (1, net). Mill Brook, IV-30-2007 (2, net), V-16-2007 (1, net), V-31-2007 (1, net), VIII-22-2007 (1, net). Unnamed Priester’s Pond Feeder, V-16-2007(5, kick; 1, net), V-31-2007 (3, kick), VI-15-2007 (5, net), VII-9-2007 (2, net), VII-25-2007 (2, net), IX-13-2007 (3, net). [Southeastern mainland rivers]: MashpeeRiver, V-23-2007 (1, net). Red Brook, IV-20-2007 (1, net). Red Brook Feeder, IV-20-2007 (14, net), V-8-2007 (25, net), V-23-2007 (41, net), VI-15-2007 (36, net),VII-16-2007 (4, net), VIII-13-2007 (11, net), IX-28-2007 (5, net).

Notes: Collected on MV from April-September and in SMA from April-September.

Platycentropus radiatus (Say, 1824): [Martha’s Vineyard]: Tiasquam River, IV-10-2007(2, net). [Southeastern mainland rivers]: Red Brook Feeder, III-28-2007 (17, net),V-23-2007 (20, net).

Notes: Collected on MV in April, and in SMA in March and May.

Pycnopsyche gentilis (MacLachlan, 1871): [Martha’s Vineyard]: Tiasquam River, V-16-2007 (1, net).


Pycnopsyche guttifera (Walker, 1852): [Martha’s Vineyard]: Roaring Brook, III-27-2007(1, kick; 9, net), IV-10-2007 (3, net), IV-30-2007 (5, net), V-16-2007 (8, net), VII-25-2007 (1, net). Menemsha Hills Creek, V-16-2007 (1, net). Tiasquam River, III-27-2007 (1, net), IV-10-2007 (1, net), IV-30-2007 (1, net). Mill Brook, IV-10-2007(1, kick; 3, net), IV-30-2007 (1, kick). Unnamed Priester’s Pond Feeder, V-16-2007(1, net), VIII-22-2007 (2, net). [Southeastern mainland rivers]: Mashpee River, IV-20-2007 (5, net), V-8-2007 (2, net), IX-13-2007 (1, net). Red Brook, IV-20-2007 (1,kick), V-8-2007 (1, kick). Red Brook Feeder, VI-18-2007 (1, net).

Notes: Collected on MV from March-August, and in SMA from April-June and inSeptember.

Pycnopsyche lepida (Hagen, 1861): [Martha’s Vineyard]: Mill Brook, III-27-2007 (6,net).


Pycnopsyche scabripennis (Rambur, 1842): [Martha’s Vineyard]: Roaring Brook, IX-13-2007 (2, net). Mill Brook, IV-30-2007 (1, net). Unnamed Priester’s Pond Feeder,VII-9-2007 (1, kick). [Southeastern mainland rivers]: Mashpee River, IV-20-2007(1, net), V-23-2007 (2, net). Red Brook, III-28-2007 (2, net), V-8-2007 (1, net).

Notes: Collected on MV in April, July, and September, and in SMA from March-May.

Odontoceridae

Psilotreta frontalis (Banks, 1899): [Martha’s Vineyard]: Roaring Brook, III-13-2007 (1,net), III-27-2007 (1, kick; 5, net), IV-10-2007 (1, kick; 1, net), VII-25-2007 (1, net),VIII-9-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-10-2007 (1, net), V-31-2007 (1, kick; 1, net), VI-15-2007 (2, net), VII-9-2007 (3, kick; 2, net), VIII-22-2007 (2, net), X-15-2007 (2, net). Paint Mill Brook, X-15-2007 (7, net).

Notes: Collected on MV from March-October.

455GREG D. WHITMORE

Philopotamidae

Chimarra obscura (Walker, 1852): [Martha’s Vineyard]: Roaring Brook, IV-10-2007 (1,kick). Tiasquam River, III-27-2007 (21, kick; 2, net), IV-10-2007 (15, kick), IV-30-2007 (13, kick), V-16-2007 (8, kick; 3, net), VI-15-2007 (26, kick), VII-9-2007 (17,kick; 37, net), VII-25-2007 (19, kick; 36, net), VIII-22-2007 (8, kick; 1, net), IX-13-2007 (14, kick; 1, net), X-15-2007 (17, kick; 9, net). Mill Brook, III-27-2007 (13,kick; 5, net), IV-10-2007 (9, kick; 1, net), IV-30-2007 (4, kick), V-16-2007 (2, kick;3, net), V-31-2007 (1, kick), VI-15-2007 (17, kick; 1, net), VII-25-2007 (2, kick),VIII-22-2007 (8, kick), IX-13-2007 (4, kick), X-15-2007 (10, kick; 3, net).Unnamed Priester’s Pond Feeder, VII-25-2007 (2, kick). [Southeastern mainlandrivers]: Mashpee River, III-28-2007 (11, kick), IV-20-2007 (10, kick), V-8-2007 (2,kick), V-23-2007 (2, kick), VI-15-2007 (5, kick), VIII-13-2007 (4, kick), IX-28-2007 (3, kick).

Notes: Collected on MV from March-October and in SMA from March-September.Common on MV and in Mashpee River.

Wormaldia moesta (Banks, 1914): [Martha’s Vineyard]: Paint Mill Brook, IV-1-2008 (5,net).

Notes: Collected on MV in April.

Phryganeidae

Oligostomis ocelligera (Walker, 1852): [Martha’s Vineyard]: Mill Brook, X-15-2007 (1,net). Paint Mill Brook, III-17-2008 (1, kick).

Notes: Collected on MV in March and October.

Oligostomis pardalis (Walker, 1852): [Martha’s Vineyard]: Roaring Brook, III-13-2007(1, net). Menemsha Hills Creek, III-19-2007 (1, net), IV-10-2007 (1, net). TiasquamRiver, IV-10-2007 (1, net). Unnamed Priester’s Pond Feeder, IV-30-2007 (1, net),X-15-2007 (1, net). Witch Brook, III-17-2008 (1, kick). Paint Mill Brook, X-15-2007 (2, net). [Southeastern mainland rivers]: Red Brook Feeder, III-28-2007 (1,net).

Notes: Collected on MV from March-April and in October, and in SMA in March.

Polycentropodidae

Cernotina spicata (Ross, 1938): [Martha’s Vineyard]: Roaring Brook, III-27-2007 (1,kick; 5, net), IX-13-2007 (1, net). Menemsha Hills Creek, V-31-2007 (1, net).Tiasquam River, III-27-2007 (2, kick; 5, net). Mill Brook, VII-25-2007 (2, net).Unnamed Priester’s Pond Feeder, IV-30-2007 (11, kick), VIII-22-2007 (1, kick).[Southeastern mainland rivers]: Mashpee River, V-8-2007 (1, net), V-23-2007 (1,net). Red Brook, V-23-2007 (1, kick).

Notes: Collected on MV from March-September, and in SMA in May.

*Nyctiophylax uncus (Ross, 1944): [Martha’s Vineyard]: Witch Brook, III-17-2008 (1,kick). Paint Mill Brook, III-17-2008 (1, net).



Polycentropus sp.: [Martha’s Vineyard]: Roaring Brook, IV-10-2007 (2, net). TiasquamRiver, IV-30-2007 (2, net), VI-15-2007 (1, net), VII-25-2007 (1, net). UnnamedPriester’s Pond Feeder, V-16-2007 (1, kick; 4, net), VIII-22-2007 (4, net), IX-13-2007 (2, kick), X-15-2007 (12, net).

Notes: Collected on MV from April-October. Larvae cannot be identified to species usingcurrent keys.

Psychomyiidae

Lype diversa (Banks, 1914): [Martha’s Vineyard]: Unnamed Priester’s Pond Feeder, IX-13-2007 (1, net). [Southeastern mainland rivers]: Mashpee River, III-28-2007 (1,net).

Notes: Collected on MV in September and in SMA in March.

Psychomyia flavida (Hagen 1861): [Southeastern mainland rivers]: Mashpee River, III-28-2007 (2, kick), V-23-2007 (1, net).

Notes: Collected in SMA in March and May.

Rhyacophila carolina (Banks, 1911): [Martha’s Vineyard]: Roaring Brook, III-27-2007(2, kick), IV-10-2007 (2, kick), V-16-2007 (1, kick), V-31-2007 (3, kick), VII-9-2007 (1, kick). Tiasquam River, IV-10-2007 (1, kick). Witch Brook, III-17-2008 (1,kick). [Southeastern mainland rivers]: Mashpee River, III-28-2007 (1, kick), V-8-2007 (3, kick), VI-15-2007 (1, net).

Notes: Collected on MV from March-July, and in SMA from March-June.

Rhyacophila fuscula (Walker, 1852): [Martha’s Vineyard]: Roaring Brook, III-13-2007(2, kick), III-27-2007 (3, kick), IV-10-2007 (1, net), IV-30-2007 (7, kick), V-31-2007 (2, kick), VI-15-2007 (2, kick; 2, net).

Notes: Collected on MV from March-June.

Rhyacophila glaberrima (Ulmer, 1907): [Martha’s Vineyard]: Unnamed Priester’s PondFeeder, IV-30-2007 (1, kick). [Southeastern mainland rivers]: Mashpee River, IV-20-2007 (5, net).

Notes: Collected on MV and in SMA in April.

Rhyacophila sp.: [Martha’s Vineyard]: Mill Brook, IX-13-2007 (2, net).Notes: Collected on MV in September. Does not conform to any species found in current

larval keys.

Uenoidae

Neophylax concinnus (MacLachlan, 1871): [Martha’s Vineyard]: Unnamed Priester’sPond Feeder, VII-25-2007 (1, kick). [Southeastern mainland rivers]: Red BrookFeeder, V-8-2007 (1, net).

Notes: Collected on MV in July and in SMA in May.

Neophylax fuscus (Banks, 1903): [Southeastern mainland rivers]: Red Brook, III-28-2007 (5, net), IV-20-2007 (20, kick), V-8-2007 (1, kick).

Notes: Collected in SMA from March-May.

Neophylax oligius (Ross, 1936): [Martha’s Vineyard]: Tiasquam River, III-27-2007 (2,kick; 2, net), IV-30-2007 (6, kick; 3, net), V-16-2007 (5, kick; 4, net), V-31-2007 (2,kick), VII-25-2007 (3, net). Mill Brook, IV-10-2007 (1, kick). Unnamed Priester’sPond Feeder, X-15-2007 (1, net). [Southeastern mainland rivers]: Mashpee River,

457GREG D. WHITMORE

III-28-2007 (1, kick; 5, net), IV-20-2007 (9, kick; 5, net), V-8-2007 (2, net), V-23-2007 (28, kick; 11, net), VI-15-2007 (4, kick; 2, net).

Notes: Collected on MV from March-July and in October, and in SMA from March-June.

ACKNOWLEDGEMENTS

This study was funded entirely by The Trustees of Reservations. Meredith Kopespent endless hours collecting and sorting specimens. Don Chandler (University of NewHampshire) provided invaluable expertise in identification of difficult groups. John Burger(University of New Hampshire) identified blackfly larvae. Tim Simmons (MassachusettsNatural Heritage and Endangered Species Program) provided species lists. MarjorieWhitmore once again pretended to be interested.

LITERATURE CITED

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Chandler, M., Loose, J.† 2001.† Draft checklist of the aquatic macroinvertebrates of Mas-sachusetts and adjoining states.† Massachusetts Division of Fisheries and Wildlife,Natural Heritage and Endangered Species Program.

Fiorentino, J. 2002. Appendix A- Islands 2000 Biological Assessment: Technical Memo-randum TM-97-1 Islands (Martha’s Vineyard) 2000 Biological Assessment. Massa-chusetts Department of Environmental Protection, Division of Watershed Manage-ment, Worcester MA.

Jones, F.M., Kimball, C.P. 1943. The Lepidoptera of Nantucket and Martha’s VineyardIslands, Massachusetts. Publications of the Nantucket Maria Mitchell Association,Vol. IV. Nantucket, MA. 217p.

Needham, J.G., Westfall JR, M.J., May, M.L. 2000. Dragonflies of North America. Scien-tific Publishers, Gainesville, FL. xv + 940pp.

Nikula, B., Loose, J.L., Burne, M.R. 2003. A Field Guide to the Dragonflies and Damself-lies of Massachusetts. Massachusetts Natural Heritage and Endangered Species Pro-gram. 197 pp.

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Publications. Yarmouth Port. 208 pp.Stark, B.P., Baumann, R.W. 2006. Plecoptera Society of North America: Valid Names for

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