the breeder’s equation r = h 2 s r = the response to selection

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S ize ofchicken lips in a population ofchickens 0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.5 0 10 20 30 40 Thickness ofchicken lips (m m) # ofobservations

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The Breeder’s equation R = h 2 S R = The response to selection h 2 = heritability (in the narrow sense) S = the selection differential. Mean of population. Mean of selected individuals. 3.5. 1.5. The Breeder’s equation R = h 2 S - PowerPoint PPT Presentation

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Page 1: The Breeder’s equation                    R  = h 2  S R = The response to selection

Size of chicken lips in a population of chickens

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

Page 2: The Breeder’s equation                    R  = h 2  S R = The response to selection

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

The Breeder’s equation R = h2 SR = The response to selectionh2 = heritability (in the narrow sense)S = the selection differential

Page 3: The Breeder’s equation                    R  = h 2  S R = The response to selection

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

The Breeder’s equation R = h2 SR = The response to selectionh2 = heritability (in the narrow sense)S = the selection differential

3.5

Mean of population

1.5

Mean of selected individuals

Page 4: The Breeder’s equation                    R  = h 2  S R = The response to selection

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

The Breeder’s equation R = h2 SR = The response to selectionh2 = heritability (in the narrow sense)S = the selection differential

3.5

Mean of population (a)

1.5

Mean of selected individuals (b)

S = a – b = 2

Page 5: The Breeder’s equation                    R  = h 2  S R = The response to selection

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

The Breeder’s equation R = h2 SR = The response to selectionh2 = heritability (in the narrow sense)S = the selection differential

3.5

Mean of population (a)

1.5

Mean of selected individuals (b)

S = a – b = 2 if h2 = 0.5, what is the response to selection?

Page 6: The Breeder’s equation                    R  = h 2  S R = The response to selection

Size of chicken lips in offspring of selected chickens

0.0 0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

Size of chicken lips in a population of chickens

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

Original population mean

Mean of offspring

If h2 = 0.5

R Response to selection

Page 7: The Breeder’s equation                    R  = h 2  S R = The response to selection

Fitness function – the relationship between phenotype and the survival probability.

Fitn

ess (

w)

0

1

Gene frequency/genotype frequency/phenotype frequency

Page 8: The Breeder’s equation                    R  = h 2  S R = The response to selection

Fitn

ess (

w)

0

1

Gene frequency/genotype frequency/phenotype frequency

An example of heterozygote superiority (overdominance

Page 9: The Breeder’s equation                    R  = h 2  S R = The response to selection

Fitn

ess (

w)

0

1

Gene frequency/genotype frequency/phenotype frequency

An example of homozygote superiority (underdominance)

Page 10: The Breeder’s equation                    R  = h 2  S R = The response to selection

Truncated selection – the probability of survival is truncated abruptly

Fitn

ess (

w)

0

1

Gene frequency/genotype frequency/phenotype frequency

Page 11: The Breeder’s equation                    R  = h 2  S R = The response to selection

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

What you need to do for your Brassica breeding

Breed these to achieve your goal of changing the frequency of a trait in a population.

Page 12: The Breeder’s equation                    R  = h 2  S R = The response to selection

0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 4.5 5.0 5.5 6.0 6.50

10

20

30

40

Thickness of chicken lips (mm)

# of

obs

erva

tions

What you need to do for your Brassica breeding

Breed these to achieve your goal of changing the frequency of a

trait in a population.

Breed these plants to use in your calculations of

heritability (h2).

Page 13: The Breeder’s equation                    R  = h 2  S R = The response to selection

How do we find heritability?

Controlled experiments

h2 = slope of the best fit line

Heritability (in the narrow sense) reflects the degree to which offspring resemble their parents.

Or

The degree to which variation in the population (offspring) is due to variation in genes (parents).

Page 14: The Breeder’s equation                    R  = h 2  S R = The response to selection
Page 15: The Breeder’s equation                    R  = h 2  S R = The response to selection

Mark the stem of the flower as you pollinate them.