stand dynamics sept 9 cp

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    Disturbance and Stand Development

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    Disturbance

    any relatively discrete event in time that

    disrupts ecosystem, community or population

    structure, and changes resources, substrate

    availability or the physical environment

    White and Pickett 1985

    biotic or abiotic

    human-caused or natural

    single-tree, gap-forming or stand-replacing

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    Examples of forest disturbances

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    Recognizing forest disturbances

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    Stand Development

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    landscape

    Stand spatially contiguous group of trees and associated vegetation having

    similar structures and growing under similar soil and climatic conditions

    - relatively homogeneous

    Forest a heterogeneous mosaic of forest stands

    Landscape a heterogeneous mosaic of forest stands and non-forest ecosystems

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    Stand Development

    4 main stages of development of even-aged

    stands following stand-replacing disturbance

    Cohort: a group of trees regenerating after a

    single disturbance until growing space is filled

    Cohorts may range in age from years to

    decades

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    Stand initiation

    Stem exclusionUnderstory re-initiation

    Old growth

    Stages of stand development

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    Stand Initiation Stage

    Re-colonizing the site

    Available growing space and resources

    Open growth

    Single cohort

    Many species invade

    Competition with non-tree species

    Foliage to the ground

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    Stem Exclusion Stage

    no available growing space

    increasing competition among trees

    above ground (light)

    below ground (water, nutrients)

    crown closure

    mortality of smaller trees

    self-thinning no regeneration

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    Understory Re-initiation Stage

    Canopy begins to open

    some overstory trees die

    crowns thin as trees get older

    branches die, ripped off by wind

    more light reaches the understory

    understory trees often different species

    advanced regeneration

    at least two cohorts

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    Old-growth Stage

    trees of all ages and sizes

    diversity of tree and non-tree species

    gaps, clumps

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    Crown classes

    Dominant

    Co-dominant

    Intermediate

    Suppressed

    Why differentiation?

    age

    micro-sites

    competition

    genetically superior

    http://www.dnr.state.md.us/education/envirothon/forestry/silviculture.html

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    Gap Dynamics

    Gaps:

    openings of any size in the overstory

    death of one or several trees

    late-successional species perpetuatethemselves

    usually little disturbance of forest floor

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    Stand initiation

    Stem exclusionUnderstory re-initiation

    Old growth

    Stages of stand development

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    Stand initiation

    Stem exclusionUnderstory re-initiation

    Old growth

    Stand-replacing disturbance

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    Stand initiation

    Stem exclusionUnderstory re-initiation

    Old growth

    Hastening stand development