polychaetous annelids from the south china sea · paxton & chou: polychaetous annelids from the...

THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8: 209-232© National University of Singapore

POLYCHAETOUS ANNELIDS FROM THE SOUTH CHINA SEA

Hannelore PaxtonDepartment of Biological Sciences. Macquarie University. NS. W 2109, Australia

L. M. ChouDepartment of Biological Sciences, National University of Singapore, Kent Ridge 119260,

Republic of Singapore

ABSTRACT. - A checklist of polychaetous anneIids from the South China Sea isgenerated from available literature. On record are 661 species belonging to 54 families.Biodiversity distribution appears linked to collecting efforts rather than a reflectionof the real situation. The potential for discovery of new species remains high.

INTRODUCTION

This check-list, based mainly on literature reports, lists 661 species of polychaetous annelidsfrom the South China Sea. The apparently highest biodiversity ranked as mainland China,Vietnam, Hong Kong, Taiwan, Singapore, Philippines, Thailand, Malaysia, is due mainlyto the collecting efforts of residing specialists or expeditions rather than true biodiversity.

The polychaetes listed belong to 54 families. The families are grouped according to Hartman(1959), except for the Eunicidae and related families. We are emphasising that this list isprimarily based on literature rather than personal knowledge and are presenting referencesfor each record. This list is only a baseline; we expect it to change considerably with theremoval of names as families are revised, and the addition of new species, as the areas ofthe South China Sea become studied more extensively. This is publication number 300 ofthe Commonwealth Key Centre for Biodiversity and Bioresources, Macquarie University.

NAMEAphroditidae

Aphrodita talpa

Hermonia hystrix

Pontogenia indica

Pontogenia lichaucoi

AUTHOR

Quatrefages, 1865

Hartman, 1959

Grube, 1878

Pillai, 1965

209

COUNTRY

China

PhilippinesChina

PhilippinesSingapore

Philippines

REFERENCE

Yang & Sun 1988:133

Pillai 1965:112Yang & Sun 1988: 133

Grube 1878:19Tan & Chou 1993:28I

Pillai 1965:112

Paxton & Chou: Polychaetous annelids from the South China Sea

NAME AUTHOR COUNTRY REFERENCE

Pontogenia nuda Horst, 1917 China Yang & Sun 1988:135

Polynoidae

Eulagisca sp. Hong Kong Mak 1982:599

Gastrolepidia clavigera Sehmarda, 1861 China Wu et al. 1980:114

Halosydna brevisetosa Kinberg, 1855 Hong Kong Mak 1982:599

Harmothoe ampullijera (Grube, 1878) Philippines Pillai 1965: 117

Harmothoe dictyophora (Grube, 1878) S. Vietnam Gallardo 1968:47China Yang & Sun 1988:148

Harmothoe extenuata (Grube, 1840) Philippines Natividad & Palpal-Latoe 1986:296

Harmothoe imbricata (Linnaeus, 1767) Hong Kong Shin 1982: 162

Harmothoe minuta (Potts, 1910) China Wu et al. 1980:114

Harmothoe nigricans Horst, 1915 China Wu et al. 1980:114

Harmothoe (Lagisca) waahli Kinberg, 1855 Philippines Natividad & Palpa1-Latoe 1986:294

Hermenia acantholepis (Grube, 1876) China Wu et al. 1980:114

Hololapidella commensalis Willey,1905 China Wu et al. 1980:114

Intoshella aphthalma Gallardo, 1968 S. Vietnam Gallardo 1968:47

?Intoshella holothuricola (Izuka, 1912) S. Vietnam Gallardo 1968:48

Iphione muricata (Savigny, 1818) China Wu et al. 1980:114Hong Kong Mak 1982:599Singapore Tan & Chou 1993:281

Lepidonotus carinulatus Grube, 1870 Philippines Pillai 1965:115

Lepidonotus cristatus (Grube, 1876) China Wu et al. 1980:114

Lepidonotus hedleyi Benham, 1915 Philippines Natividad & Palpal-Latoe 1986:290Lepidonotus jacksoni Kinberg, 1855 Philippines Natividad & Palpal-Latoe 1986:286Lepidonotus squamatus (Linnaeus, 1767) Hong Kong Mak 1980:599

Lepidonotus tenuisetosus (Gravier, 1901) Hong Kong Shin 1982:162

Lepidasthenia izukai Imajima & Hartman, 1964 S. Vietnam Gallardo 1968:46China Yang & Sun 1988:144

Lepidasthenia microlepis Potts, 1910 China Wu et al. 1980:114

Lucopia magnicirra Pillai, 1965 Philippines Pillai 1965: 117Parahalosydnopsis hartmanae Pettibone, 1977 Hong Kong Shin 1982:162

Paralepidonotus ampullijerus (Grube, 1878) Singapore Tan & Chou 1993:281Singapore Tan & Chou 1993:281

Scalisetosus jragilis (Claparede, 1868) China Wu & Wu 1987:79

Scalisetosus longicirrus (Sehmarda, 1861) China Wu et al. 1980:114

Thormora jukesii Baird, 1865 China Wu et al. 1980:114Philippines Natividad & Palpa1-Latoe 1986:285

Polyodontidae

Eupanthalis maculosa (Treadwell, 1931) China Wu et al. 1980:114

Neopolyodontes lepidus Shen & Wu, 1993 China Shen & Wu 1993:537Neopanthalis muricatus Shen & Wu, 1993 China Shen & Wu 1993:537Panthalis jogasimae Izuka, 1912 China Wu et al. 1980:114Polyodontes gracilis Pflugfelder, 1932 S. Vietnam Gallardo 1968:49Polyodontes maxillosus (Ranzani, 1817) China Wu et al. 1980:114Polyodontes melanonotus (Grube, 1876) Hong Kong Shin 1982:162

China Yang & Sun 1988:151

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THE RAFFLES BULLETIN OF ZOOLOGY 2000 Supplement No. 8


Sigalionidae

Ehlersileanira incisa (Grube, 1877) China Yang & Sun 1988:154

Euleanira ehlersi Horst, 1916 S. Vietnam Gallardo 1968:50Mayel/a sp. Taiwan Hsieh, pers. comm.

Psammolyce fijiensis Mclntosh, 1885 S. Vietnam Gallardo 1968:50

Psammolyce malayana Horst, 1913 Philippines Natividad & Pa1pal-Latoc 1986:301

Sigalion sp. Taiwan Hsieh, pers. comm.

Sthenelais boa (Johnston, 1839) Hong Kong Shin 1982: 162

Sthenelais mitsuii (Okuda, 1938) S. Vietnam Gallardo 1968:51

Sthenelais nami Gallardo, 1968 S. Vietnam Gallardo 1968:51Augener, 1927 S. Vietnam Gallardo 1968:52

Sthenelais japonica (Mclntosh, 1885) S. Vietnam Gallardo 1968:52

Sthenolepis izuensis (Takahashi, 1938) China Wu et a1. 1980:115

Sthenolepis tentaculata (Horst, 1917) China Wu et a1. 1980: 115

Thalenessa oculata (Peters, 1854) China Wu & Wu 1987:79

Thalenessa cf. spinosa asiatica Uschakov & Wu, 1959 Taiwan Hsieh, pers. comm.

Thalenessa stylolepis Willey, 1905 S. Vietnam Gallardo 1968:53

Thalenessa tropica Hartman, 1954 S. Vietnam Gallardo 1968:53

Eulepethidae

Pareulepis malayana (Horst, 1913) S. Vietnam Gallardo 1968:54

Chrysopetalidae

Bhawania brevis Gallardo, 1968 S. Vietnam Gallardo 1968:54Hong Kong Shin 1982: 162

Bhawania cryptocephala Gravier, 1901 China Yang & Sun 1988:158

Bhawania goodei Webster, 1884 Hong Kong Mak 1982:599Philippines Natividad & Palpal-Latoc 1986:304China Wu & Wu 1987:79

Chrysopetalum ehlersi Gravier, 1901 China Yang & Sun 1988:160

Paleanotus bel/is (Johnson, 1897) Hong Kong Mak 1982:599

Paleanotus chrysolepis Schmarda, 1861 China Yang & Sun 1988:161

Paleanotus debilis (Grube, 1855) China Wu et a1. 1980:115

Amphinomidae

Amphinome pulchra Horst, 1912 China Wu et a1. 1980:115

Amphinome rostrata (Pallas, 1766) China Yang & Sun 1988:164Philippines Natividad & Pa1pal-Latoc 1986:321

Chloeia flava (Pallas, 1766) S. Vietnam Gallardo 1968:56Hong Kong Shin 1982:163Philippines Natividad & Palpal-Latoc 1986:321China Yang & Sun 1988:165Singapore Tan & Chou 1993:281

Chloeia fusca Mclntosh, 1885 China Yang & Sun 1988:165

Chloeia entypa Chamberlin, 1919 Philippines Natividad & Palpal-Latoc 1986:321

Chloeia inermis Quatrefages, 1865 China Wu & Wu 1987:79

Chloeia parva Baird, 1870 S. Vietnam Gallardo 1968:56China Yang & Sun 1988:165

Chloeia rosea Pous, 1909 Philippines Natividad & Pa1pa1-Latoc 1986:324

211



Chloeia violacea Horst, 1910 S. Vietnam Gallardo 1968:56China Wu et at. 1980:115

Eurythoe complanata (pallas, 1766) Philippines Pillai 1965:119China Wu et at. 1980:115Singapore Tan & Chou 1993:282

Eurythoe parvecarunculata Horst, 1912 China Wu et at. 1980:115

Unopherus hirsuta (Wesenberg-Lund, 1949) Hong Kong Shin 1982: 163

Notopygos gigas Horst, 1911 China Wu et at. 1980:115

Notopygos labiatus Fauvel, 1932 Philippines Natividad & Palpal-Latoe 1986:326

Notopygos ornata Grube, 1856 Philippines Natividad & Palpal-Latoe 1986:326

Notopygos sibogae Horst, 1911 China Wu et at. 1980:115

Notopygos supragigas Usehakov & Wu, 1962 China Wu et a.11980:115

Notopygos variabilis Grube, 1855 Philippines Pillai 1965: 120

Paramphinome indica Fauvel, 1932 China Wu et at. 1980:115

Pareurythoe borealis (Sars, 1862) China Yang & Sun 1988:170

Pherecardia striata (Kinberg, 1857) Philippines Pillai 1965:121China Wu et at. 1980:115

Pseudeurythoe hirsuta Wesenberg-Lund, 1949 S. Vietnam Gallardo 1968:57

Pseudeurythoe oligobranchia Wu et aI, China Wu et at. 1980:115

Euphrosinidae

Euphrosyne foliosa Audouin & M. Edw., 1833 China Sun et at. 1978:161

Euphrosine myrtosa Savigny, 1818 Hong Kong Mak 1982:599China Yang & Sun 1988:172

Phyllodocidae

Anaitides madeirensis (Langerhans, 1880) China Sun et at. 1978:135 as P.(Anait.)mad.Hong Kong Mak 1982:599Singapore Tan & Chou 1993:282

Anaitides tenuissima (Grube, 1878) Philippi nes Grube 1878:95

Anaitides ?tenuissima S. Vietnam Gallardo 1968:57

Eteone (Mysta) siphodonta (Delle Chiaje, 1822) Singapore Tan & Chou 1993:282

Eteone (Mysta) tchangsii Usehakov & Wu, 1959 China Yang & Sun 1988:109

Eulalia (Pterocirrus) magalhaensis Kinberg, 1866 Singapore Tan & Chou 1993:282

Eulalia viridis (Linnaeus, 1767) Hong Kong Mak 1982:599China Yang & Sun 1988: 105

Eumida sanguinea (Marenzeller, 1879) Hong Kong Mak 1982:599

Genetyllis castanea (Marenzeller, 1879) Hong Kong Mak 1982:599China Yang & Sun 1988: 108Philippines Natividad & Palpal-Latoe 1986:275

Genetyllis sp. Taiwan Hsieh, pers. eomm.Mysta ornata (Grube, 1878) S. Vietnam Gallardo 1968:58

China Yang & Sun 1988:109

Notophyllum splendens (Sehmarda, 1861) China Wu et at. 1980:115

Paranaitis zeylanica (Willey, 1905) S. Vietnam Gallardo 1968:58

Phyllodoce (Anaitides) dissotyla Willey, 1905 China Wu & Sun 1978:176Phyllodoce (Anait.) madeirensis Langerhans, 1880 China Wu et at. 1980:115

Phyllodoce fristedti Bergstriim, 1914 China Wu et at. 1980:115

212



Phyllodoce ?laminosa Savigny, 1818 S. Vietnam Gallardo 1968:58

Phyllodoce malmgreni Gravier, 1900 China Yang & Sun 1988:110

Pterocirrus macroceros (Grube, 1860) China Yang & Sun 1988:111

Alciopidae

Alciopina parasitica Claparecte & Panceri, 1867 China Shen 1978:215

Krohnia lepidota (Krohn, 1845) China Sun & Wu 1979:64

Naiades cantrainii Delle Chiaje, 1830 China Shen 1978:211

Plotohelmis capitata (Greeff, 1876) China Shen 1978:214

Rhynchonerella petersii (Langerhans, 1880) China Sun & Wu 1979:63

Rhynchonerella gracilis Costa, 1864 China Wu et at. 1980:116

Rhynchonerella xishaensis Shen, 1978 China Shen 1978:214

Vanadis crystallina Greeff, 1876 China Shen 1978:212

Vanadis juscapunctata Treadwell, 1906 China Shen 1978:212

Vanadis minuta Treadwell, 1906 China Shen 1978:212Singapore Tan & Chou 1993:282

Lopadorrhynchidae

Lopadorrynchus brevis Grube, 1855 China Shen 1978:203

Lopadorrynchus krohni (Claparecte, 1870) China Shen 1978:205

Lopadorrhynchus tuberculus Shen, 1978 China Shen 1978:205

Lopadorrhynchus uncinatus Fauvel, 1915 China Shen 1978:203

Maupasia caeca Vignier, 1886 China Shen 1978:207

Pedinosoma curtum Reibisch, 1895 China Shen 1978:207

Pelagobia longicirrata Greeff, 1879 China Shen 1978:206

Lacydoniidae

Paralacydonia paradoxa Fauvel, 1913 S. Vietnam Gal1ardo 1968:59China Wu et at. 1980:117Hong Kong Shin 1982: 163

Iospilidae

Phalacrophorus pietus Greeff, 1879 China Shen 1978:208

Phalocrophorus uniformis Reibisch, 1895 China Shen 1978:209

Pontodoridae

Pontodora pelagica Greef, 1879 China Shen 1978:209

Typhloscolecidae

Sagitella kowalevskii Wagner, 1872 China Shen 1978:222

Travisiopsis dubia Stop-Bowitz, 1948 China Wu et at. 1980:117

Travisiopsis levinseni Southern, 1910 China Shen & Wu 1978:105

Travisiopsis lobifera Levinsen, 1885 China Shen 1978:223

Typhloscolex muelleri Busch, 1851 China Shen 1978:221

Tomopteridae

Tomopteris apsteini Rosa, 1908 China Sun & Wu 1979:65

Tomopteris cavalii Rosa, 1908 China Shen 1978:218

Tomopteris duccii Rosa, 1908 China Shen 1978:220

213



Tomopteris dunckeri Rosa, 1908 China Wu et al. 1980: 117

Tomopteris elegans Chun, 1888 China Shen 1978:217

Tomopteris ligulata Rosa, 1908 China Shen 1978:220

Tomopteris mariana Greef, 1885 China Wu et al. 1980:117

Tomopteris planktonis Apstein, 1900 China Shen 1978:218

Tomopteris rolasi Greef,1885 China Shen & Wu 1978:103

Tomopteris septentrionalis Steenstrup, 1849 China Sun & Wu 1979:67

Hesionidae

Bonuania parva Pillai, 1965 Phillippines Pillai 1965: 123

Gyptis labatus (Hessle, 1925) China Yang & Sun 1988:80

Gyptis cf. maraunibinae Taiwan Hsieh, pers. eomm.

Hesione genetta Grube, 1866 China Wu et al. 1980: 117

Hesione intertexta Grube, 1878 Philippines Grube 1878:China Wu et al. 1980:117

Hesione splendida Savigny, 1818 China Wu et al. 1980:117

Leocrates chinensis Kinberg, 1866 Hong Kong Mak 1982:599China Yang & Sun 1988:82

Leocrates claparedii (Costa in C1aparede, 1868) Singapore Tan & Chou 1993:282

Ophiodromus berrisfordi Day, 1967 China Wu & Wu 1987:79

Ophiodromus pugettensis (Johnson, 1901) Hong Kong Shin 1982:164

Podarke latifrons (Grube 1878) Singapore Tan & Chou 1993:282

Podarke cf. obscura (Verrill, 1837) China Yang & Sun, 1988:84

Sinohesione genitaliphora Westheide et aI., 1994 Thailand Westheide et al. 1994:95

Pilargidae

Ancistargis sp. Taiwan Hsieh, pers. eomm.

Loandalia indica Thomas, 1963 S. Vietnam Gallardo 1968:60

Pilargis mohri Gallardo, 1968 S. Vietnam Gallardo 1968:60

Pilargis verrucosa pacific Usehakov, 1955 China Yang & Sun 1988:77

Sigambra cf. bassi Taiwan Hsieh, pers. eomm.

Sigambra hanaokai (Kitamori, 1960) S. Vietnam Gallardo 1968:61

Sigambra tentaculata (Treadwell, 1941) Hong Kong Shin 1982:164

Synelmis albini (Langerhans, 1881) S. Vietnam Gallardo 1968:61China Wu et al. 1980:117Hong Kong Shin 1982: 164

Synelmis annamita Gallardo, 1968 S. Vietnam Gallardo 1968:62

Syllidae

Autolytus robustisetus Wu & Sun, 1978 China Sun et al. 1978:137

Autolytus setoensis 1majima, 1966 China Yang & Sun 1988:55

Autolytus sp. Taiwan Hsieh, pers. comm.

Ehlersia cornuta (Rathke, 1843) Hong Kong Shin 1982: 164China Yang & Sun 1988:65

Ehlersia ef. rosea Imajima, 1966 China Yang & Sun 1988:66

Eusyllis edenticulata Pillai, 1965 Philippines Pillai 1965:125

Eusyllis ef. irregulata Imajima, 1966 China Yang & Sun, 1988:57

214



Exogone verugera (Claparede, 1868) China Sun & Wu 1981:621

Haplosyllis spongicola (Grube, 1855) China Yang & Sun 1988:67

Haplosyllis spongicola tentaculata (Marion, 1879) China Yang & Sun 1988:67

Langerhansia cornuta (Rathke, 1843) S. Vietnam Gallardo 1968:62

Myrianida cf. pachycera (Augener, 1913) China Yang & Sun 1988:56

Odontosyllis gibba Claparede, 1863 China Sun et al. 1978:140

Odontosyllis rubrofasciata Grube, 1879 China Wu & Sun 1978:178

Parautolytus luzonensis Pillai, 1965 Philippines Pillai 1965:123

Sphaerosyllis hystrix Claparede, 1863 China Sun & Wu 1981:622

Streptosyllis baolingi Ding & Westheide, 1994 China Ding & Westheide 1994:304

Streptosyllis hainanensis Ding & Westheide, 1994 China Ding & Westheide 1994:308

Syllis amica Quatrefages, 1865 China Yang & Sun 1988:68

Syllis gracilis Grube, 1840 China Yang & Sun 1988:69Singapore Tan & Chou 1993:283

Trypanosyllis taeniaformis (Haswell, 1886) Hong Kong Mak 1982:599China Yang & Sun 1988:71 as

T. (Tryp. )taen.

Trypanosyllis zebra (Grube, 1860) China Wu et al. 1980:118

Typosyllis aciculata orientalis Imajima & Hartman, 1964 S. Vietnam Gallardo 1968:63Hong Kong Mak 1982:599

Typosyllis alternata (Moore, 1908) China Wu & Sun 1978:179

Typosyllis armillaris (Muller, 1771) China Wu et al. 1980:118

T)posyllis exilis Gravier, 1900 Philippines Natividad & Palpal-Latoc 1986:310

Typosyllis maculata Imajima, 1966 China Yang & Sun 1988:74

Typosyllis magnipectinis (Storch, 1967) China Licher et al. 1995: 107

Typosyllis monilata Imajima, 1966 China Sun et al. 1978:141

Typosyllis prolifera (Krohn, 1852) China Wu et al. 1980:118

Typosyllis variegata (Grube, J860) S. Vietnam Gallardo 1968:63China Sun et al. 1978: 140Hong Kong Mak 1982:599

NereididaeCeratonereis anchylochaeta (Hors!, 1924) China Wu et al. 1981:Table 1

Ceratonereis cf. burmensis Monro, 1937 China Sun et al. 1978: 148

Ceratonereis costae (Grube, 1840) China Sun et al. 1978:146Hong Kong Mak 1982:599

Ceratocephale fauveli (Pillai, 1961) S. Vietnam Gallardo 1968:63

Ceratonereis erythraeensis Fauvel, 1918 China Wu et al. 1981:Table 1

Ceratonereis hircinicola (Eisig, 1870) China Wu et al. 1981:Table I

Ceratonereis ?hircinicola Singapore Tan & Chou 1993:283

Ceratonereis japonica Imajima, 1972 China Sun et al. 1978:145

Ceratonereis marmorata Horst, 1924 China Wu et al. 1981:Table 1

Ceratonereis mirabilis Kinberg, 1866 China Sun et al. 1978:144Hong Kong Mak 1982:599Singapore Tan & Chou 1993:283

Ceratonereis pachychaeta Fauvel,1918 China Sun et al. 1978:147

Ceratonereis tripartita Horst, 1918 China Wu et al. 1980:118

215



Ceratonereis sp. Taiwan Hsieh, pers. comm.

Gymnonereis fauveli (Pillai, 1961) Thailand Hylleberg & Nateewathana 1988:8

Dendronereis pinnaticirris Grube, 1878 Philippines Pillai 1965:127China Wu et al. 1981:Tb1e 1Taiwan Hsieh, pers. comm.

Leonnates decipiens Fauve1, 1929 China Wu et al. 1981 :Tab1e 1Singapore Tan & Chou 1993:283

Leonnates decipiens manilensis Pillai, 1965 Philippines Pillai 1965:144

Leonnates jousseaumei Gravier, 1911 China Wu et al. 1981:Table 1Singapore Tan & Chou 1993:283

Leonnates nipponicus 1majima, 1972 Hong Kong Mak 1982:599

Leonnates persica Wesenberg-Lund, 1949 S. Vietnam Gallardo 1968:64China Wu et al. 1981 :Tab1e 1Hong Kong Shin 1982: 165

Leonnates sp. Taiwan Hsieh, pers. comm.

Namalycastis aibiuma (Muller in Grube, 1881) China Wu et al. 1981:Tab1e 1Taiwan Hsieh, pers. comm.

Namalycastis rigida Pillai, 1965 Philippines Pillai 1965:133

Neanthes arenaceodentata (Moore, 1903) S. Vietnam Gallardo 1968:65

Neanthes bongcoi Pillai, 1965 Philippines Pillai 1965:142

Neanthes diversicolor (Muller, 1776) Philippines Natividad & Pa1pa1-Latoc 1986:315

Neanthes glandicincta (Southern, 1921) China Wu et al. 198I:Tab1e 1Taiwan Hsieh, pers. comm.

Neanthes maculata Wu et aI., 1981 China Wu et al. 1981:66

Neanthes manatensis Pillai, 1965 Philippines Pillai 1965:139

Neanthes nanhaiensis Wu et aI., 1981 China Wu et al. 1981:68

Neanthes unifilSciata (Willey, 1905) China Wu et al. 1980:119Hong Kong Mak 1982:599

Nectoneanthes ijimai (Izuka, 1912 ) China Wu et al. 1981:Tab1e 1Nectoneanthes multignatha Wu et aI., 1981 China Wu et al. 1981:73Nectoneanthes oxypoda (Marenzeller, 1879) China Wu et al. 1981:Tab1e INectoneanthes cf. oxypoda Hong Kong Shin 1982: 164Nereis coutieri Gravier, 1899 China Wu et al. 1981 :Table INe reis dayana Sun & Shen, 1978 China Sun et al. 1978:155Nereis denhamensis Augener, 1913 China Sun et al. 1978:151Nereis falcaria (Willey, 1905) China Wu et al. 1980:119Nereis guangdongensis Wu et aI., 1981 China Wu et al. 1981:75

Nereis heteromorpha Horst, 1924 China Sun et al. 1978:157Nereis jacksoni Kinberg, 1866 China Wu et al. 1981:Tab1e INereis multignatha Imajima & Hartman, 1964 China Wu et al. 1981 :Tab1e 1Nereis nichollsi Kotl, 1951 China Wu et al. 1980:]]9Nereis pers;c'a Fauve1, 1911 China Wu et al. 1981:Table 1

Hong Kong Shin 1982: 165Nereis surugaense nanhaiensis Sun & Shen, 1978 China Sun et al. 1978:153Nereis trifasciata Grube, 1878 China Wu et al. 1980:119Nereis unifasciata Willey,1905 China Sun et al. 1978:154Nereis zhongshaensis Shen & Sun, 1978 China Sun et al. 1978:158

216



Nereis zonata Malmgren, 1867 China Wu et al. 1980:119

Nereis cf. zonata Taiwan Hsieh, pers. comm.

Nicon japonica 1majima, 1972 China Wu & Sun 1979:104

Nicon maculata Kinberg, 1866 S. Vietnam Gallardo 1968:65 as N. maculatusChina Wu & Sun 1979:101

Nicon sinica Wu & Sun, 1979 China Wu & Sun 1979:99

Paraleonnates uschakovi Chlebovitsch & Wu, 1962 China Wu et al. 1981:Table I

Perinereis aibuhitensis Grube, 1878 China Wu et al. 1981:Table 1Taiwan Hsieh, pers. comm.

Perinereis barbara Momo, 1926 Singapore Tan & Chou 1993:283

Perinereis camiguinoides Augener, 1922 China Wu et al. 1981:Table 1

Perinereis cavifrons Ehlers, 1920 China Wu et al. 1981:Table I

Perinereis cultrifera cultrifera (Grube, 1840) China Wu et al. 1981:Table IHong Kong Mak 1982:599Philippines Natividad & Palpal-Latoc 1986:313

Perinereis cultrifera jloridana Ehlers, 1868 China Wu et al. 1985:218

Perinereis cultrifera helleri Grube, 1878 China Wu et al. 1985:217

Perinereis cultrifera obfuscata Grube, 1878 China Wu et al. 1985:219

Perinereis cultrifera perspicillata Grube, 1878 Singapore Tan & Chou 1993:283

Perinereis cultrifera striolata Grube, 1878 Singapore Tan & Chou 1993:284

Perinereis nuntia (Savigny, 1818) China Wu et al. 1981:Table 1

Perinereis nuntia brevicirris (Grube, 1878) Singapore Tan & Chou 1993:284China Wu et al. 1985:208

Perinereis nuntia majungaensis Fauvel, 1918 China Wu et al. 1985:213

Perinereis nuntia vallata (Grube, 1857) China Wu et al. 1985:210

Perinereis quatrefagesi (Grube, 1878) Thailand Hylleberg et al. 1986:5

Perinereis rhombodonta Wu et aI., 1981 China Wu et al. 1981:83

Perinereis singaporiensis (Grube, 1878) Singapore Grube 1878:84

Perinereis suluana Horst, 1924 China Sun et al. 1978: 150

Perinereis vancaurica (Ehlers, 1868) China Wu et al. 1981:Table 1Singapore Tan & Chou 1993:284

Perinereis weijhouensis Wu et aI., 1981 China Wu et al. 1981:83

Perinereis cf. weijhouensis Taiwan Hsieh, pers. comm.

Platynereis abnormis (Horst, 1924) China Wu et al. 1980:119

Platynereis bicanaliculata (Baird, 1863) China Wu et al. 1981:Table 1

Platynereis dumerilli (Audouin & M. Edw., 1833) China Wu et al. 1980:119

Platynereis pulchella Gravier, 1902 S. Vietnam Gallardo 1968:65China Wu et al. 1981:Table 1

Platynereis sinica Sun & Shen, 1978 China Sun et al. 1978: 148

Pseudonereis anomala Gravier, 1902 China Wu et al. 1980:119

Pseudonereis gallapagensis Kinberg, 1866 China Wu et al. 198\ :Table 1

Pseudonereis variegata (Grube, 1857) China Wu et al. 1981:Table 1

Rullierinereis misakiensis (Imajima & Hayashi, 1972) China Wu & Sun 1979:106

Sinonereis heteropoda Wu & Sun, 1979 China Wu & Sun 1979:96

Tambalagamia fauveli Pillai, 1961 China Yang & Sun 1988:28Taiwan Hsieh, pers. comm.

Tylonereis bogoyawleskyi Fauvel, 1911 China Wu et al. 1981:Table 1

217



Tylonereis heterochaeta Tan & Chou, 1994 Singapore Tan & Chou 1994:665

Tylorrhynchus heterochaetus (Quatrefages, 1865) China Wu et al. 1981:Tab1e 1

Nephtyidae

Aglaophamus dibranchis (Gmbe, 1877) Singapore Tan & Chou 1993:284

Aglaophamus dicirroides Fauehald, 1968 S. Vietnam Faueha1d 1968: 10

Aglaophamus ef. jeffreysii (McIntosh, 1885) China Wu et al. 1980:120

Aglaophamus lyrochaeta (Fauve1, 1902) Hong Kong Shin 1982: 165

Aglaophamus munamaorii Gibbs, 1971 China Wu et al. 1980:120

Aglaophamus orientalis Faueha1d, 1968 S. Vietnam Faucha1d 1968:12

Aglaophamus sinensis (Fauve1, 1932) S. Vietnam Fauchald 1968:13China Yang & Sun 1988:94

Aglaophamus vietnamensis Faucha1d, 1968 S. Vietnam Faucha1d 1968:13

Aglaophamus sp. Taiwan Hsieh, pers. comm.

Inermonephtys gallardi Faueha1d, 1968 S. Vietnam Faucha1d 1968: 15

Inermonephtys inermis (EhIers, 1887) S. Vietnam FauchaId 1968:16China Wu et al. 1980:120

Micronephthys sphaerocirrata (Wesenberg-Lund, 1949) S. Vietnam Fauchald 1968: 18Singapore Tan & Chou 1993:284

Micronephthys sp. Taiwan Hsieh, pers. comm.

Nephtys cf. califomiensis Taiwan Hsieh, pers. comm.

Nephtys oligobranchia Southern, 1921 S. Vietnam Faucha1d 1968: 18

Nephtys polybranchia Southern, 1921 S. Vietnam FauchaId 1968:19China Yang & Sun 1988:98

Sphaerodoridae

Sphaerodorum sp. S. Vietnam Gallardo 1968:67

Glyceridae

Glycera alba (Muller, 1788) S. Vietnam Gallardo 1968:68Hong Kong Shin 1982:166China Yang & Sun 1988:87

Glycera ?capitata Oersted, 1843 S. Vietnam Gallardo 1968:68

Glycera chirori lzuka, 1912 China Yang & Sun 1988:87

Glycera convoluta Keferstein, 1862 Singapore Tan & Chou 1993:284

Glycera cf. convoluta Taiwan Hsieh, pers. comm.

Glycera lancadivae Schmarda, 1861 China Sun et al. 1978:136

Glycera nicobarica Grube, 1868 S. Vietnam Gallardo 1968:69

Glycera onomichiensis Izuka, 1912 S. Vietnam Gallardo 1968:69China Wu et al. 1980:120

Glycera posterobranchia Hoagland, 1920 Philippines Hoag1and 1920:620S. Vietnam Gallardo 1968:70

Glycera parashadi Fauve1, 1932 China Wu et al. 1980:120

Glycera rouxii Audouin & M. Edw., 1833 China Wu & Wu 1987:79Singapore Tan & Chou 1993:285

Glycera cf. subaenea Taiwan Hsieh, pers. comm.

Glyera cf. tenuis Taiwan Hsieh, pers. comm.

Glycera tesselata Gmbe, 1863 S. Vietnam Gallardo 1968:70China Wu et al. 1980:119

218



GoniadidaeGlyeinde oligodon Southern, 1921 S. Vietnam Gallardo 1968:71

Goniada annulata Moore, 1905 China Wu et al. 1980:119

Goniada emerita Audouin & M. Edw., 1833 China Wu et al. 1980:119

Goniada cf. emerita Taiwan Hsieh, pers. comm.

Goniada cf. maeulata Oersted, 1843 China Wu et al. 1980:119Singapore Tan & Chou 1993:285Taiwan Hsieh, pers. comm.

Goniada multidentata Arwidsson, 1899 S. Vietnam Gallardo 1968:72

Goniada paueidens Grube, 1878 Hong Kong Mak 1982:599

Goniada tridens Gallardo, 1968 S. Vietnam Gallardo 1968:73

Goniada uneinigera EhJers, 1901 S. Vietnam Gallardo 1968:73

Goniada cf. uneinigera Taiwan Hsieh, pers. comm.

Goniadopsis sp. Taiwan Hsieh, pers. comm.

DorvilleidaeDorvillea australiensis (McIntosh, 1885) China Wu et al. 1980:121 as Papilliod.aust.

Sehistomeringos ineerta (Schmarda, 1861) S. Vietnam Gallardo 1968:91 as Dorvillea ine.Hong Kong Shin 1982: 167

EunicidaeEuniee afra Peters, 1854 China Wu & Wu 1987:79

Euniee antennata (Savigny, 1820) Philippines Pi1lai 1965: 148China Wu et al. 1980:121Hong Kong Mak 1982:599Singapore Tan & Chou 1993:285

Euniee aphroditois (Pallas, 1788) China Wu & Wu 1987:79Singapore Tan & Chou 1993:285

Euniee ?australis Quatrefages, 1865 S. Vietnam Gallardo 1968:77

Euniee australis Hong Kong Mak 1982:599China Wu & Wu 1987:79

Euniee eineta (Kinberg, 1865) Hong Kong Mak 1982:599

Euniee eoeeinea Grube, 1878 China Wu et al. 1980:121

Euniee eoeeinioides Augener, 1922 Singapore Tan & Chou 1993:285

Euniee grubei Gravier, 1900 Singapore Tan & Chou 1993:286

Euniee hirsehi Fauchald, 1992 Singapore Tan & Chou 1993:286

Euniee indiea Kinberg, 1865 S. Vietnam Gallardo 1968:78Philippines Gallardo 1968:78China Wu et al. 1980:121

Euniee lueei Grube, 1856 Singapore Tan & Chou 1993:286

Euniee marenzelleri Gravier, 1900 China Wu et al. 1980:121

Euniee medicina Moore, 1903 S. Vietnam Gallardo 1968:78

Euniee nesiotes (Chamberlin, 1919) Singapore Tan & Chou 1993:286

Euniphysa aeuleata Wesenberg-Lund, 1949 China Shen & Wu 1991:131Singapore Tan & Chou 1993:286

Euniphysa oeulata Wu & Sun in Wu et aI., 1979 China Wu et al. 1979:89

Euniphysa unieusa Shen & Wu 1991 China Shen & Wu 1991:131

Heterophysa tridontesa Shen & Wu, 1991 China Shen & Wu 1991:138

219



Lysidice collaris Grube, 1870 S. Vietnam Gallardo 1968:79Philippines Gallardo 1968:79China Sun et al. 1978: 162 as L.nin.collarisHong Kong Mak 1982:599Singapore Tan & Chou 1993:287

Lysidice ninetta Audouin & M. Edw., 1833 Hong Kong Mak 1982:599China Yang & Sun 1988:184

Marphysa adenensis Gravier, 1900 Hong Kong Mak 1982:600

Marphysa bellii (Aud. & M. Edwards,1833) S. Vietnam Gallardo 1968:80Thailand Gallardo 1968:80Hong Kong Shin 1982: 166

Marphysa bifurcata Kott, 1951 S. Vietnam Gallardo 1968:80

Marphysa depressa (Schmarda, 1861) China Yang & Sun 1988:186

Marphysa disjuncta Hartman, 1961 Singapore Tan & Chou 1993:286

Marphysa gravelyi Southern, 1921 Philippines Pillai 1965: 150

Marphysa macintoshi Crossland, 1903 China Wu et al. 1980:121Singapore Tan & Chou 1993:286

Marphysa mossambica (Peters, 1854) China Wu & Wu 1987:79Singapore Tan & Chou 1993:287

Marphysa sanguinea (Montagu, 1815) Hong Kong Shin 1982:166China Yang & Sun 1988:186

Marphysa stragulum (Grube, 1878) China Wu et al. 1980:121

Nauphanta sp. Tan & Chou, in press Singapore Tan & Chou, in press

Nematonereis unicornis (Grube, 1840) S. Vietnam Gallardo 1968:81China Wu et al. 1980:121

Palola siciliensis (Grube, 1840) China Wu et al. 1980:121Hong Kong Mak 1982:600

Paraeuniphysa falciseta Shen & Wu, 1991 China Shen & Wu 1991:134

Paraeuniphysa spinea (Miura, 1977) China Shen & Wu 1991:136

Paraeuniphysa taiwanensis Wu & He, 1988 Taiwan Shen & Wu 1991:134

Paramarphysa sp. Hong Kong Mak 1982:600

Lumbrineridae

Lumbrineris acutiformis Gallardo, 1968 S. Vietnam Gallardo 1968:82

Lumbrineris ? amboinensis (Grube, 1877) S. Vietnam Gallardo 1968:82

Lumbrineris bifurcata (Mclntosh, 1885) China Wu et al. 1980:121

Lumbrineris caudaensis Gallardo, 1968 S. Vietnam Gallardo 1968:83

Lumbrineris cf. cruzensis Hartman, 1944 China Wu et al. 1980:121Taiwan Hsieh, pers. comm.

Lumbrineris hartmani (Day, 1953) Singapore Tan & Chou 1993:287

Lumbrineris heteropoda (Marenzeller, 1879) S. Vietnam Gal1ardo 1968:84China Yang & Sun 1988:192

Lumbrineris inflata Moore, 1911 Hong Kong Mak 1982:600

Lumbrineris japonica (Marenzeller, 1879) S. Vietnam Gallardo 1968:84

Lumbrineris latreilli Audouin & M. Edw., 1834 China Wu et al. 1980:121Singapore Tan & Chou 1993:287

Lumbrineris mucronata (Ehlers, 1908) S. Vietnam Gallardo 1968:85

Lumbrineris nagae Gallardo, 1968 S. Vietnam Gallardo 1968:85China Yang & Sun 1988:191

220



Lumbrineris ?notocirrata (Fauvel, 1932) S. Vietnam Gallardo 1968:86

Lumbrineris pterignatha Gallardo, 1968 S. Vietnam Gallardo 1968:86

Lumbrineris shiinoi Gallardo, 1968 S. Vietnam Gallardo 1968:87Hong Kong Shin 1982: 167

Lumbrineris simplex Southern, 1921 China Wuetal.1980:121

Lumbrineris sphaerocephala (Sehmarda, 1861) China Wu et al. 1980: 121

Ninoe bruuni Gallardo, 1968 S. Vietnam Gallardo 1968:88

Ninoe mando (Crossland, 1924) S. Vietnam Gallardo 1968:88

Oenonidae

Arabella iricolor (Montague, 1804) Singapore Tan & Chou 1993:287

Arabella mutans (Chamberlin, 1919) China Wu & Wu 1987:79

Arabella novecrinita Crossland, 1924 S. Vietnam Gallardo 1968:89

Drilonereis logani Crossland, 1924 S. Vietnam Gallardo 1968:89China Wu et al. 1980:121

Drilonereis ef. logani Taiwan Hsieh, pers. eomm.

Drilonereis ?paucidentata Treadwell, 1922 S. Vietnam Gallardo 1968:90

Notocirrus biaciculus Sehmarda, 1861 S. Vietnam Gallardo 1968:90

Notocirrus japonicus (Okuda, 1939) S. Vietnam Gallardo 1968:91

Oenone julgida (Savigny, 1818) Hong Kong Mak 1982:600Singapore Tan & Chou 1993:287

Onuphidae

Diopatra bulohensis Tan & Chou, 1996 Singapore Tan & Chou 1996:359

Diopatra neapolitana Delle Chiaje, 1841 Singapore Tan & Chou 1993:285

Diopatra sugokai lzuka, 1907 China Yang & Sun 1988:175as D. chiliensis

Taiwan Hsieh & Chang 1991 :331as D. bilobata

Diopatra variabilis Southern, 1921 S. Vietnam Gallardo 1968:74China Wu et al. 1980:120Hong Kong Shin 1982:166

Hyalinoecia tubicola (MUller, 1788) China Wu et al. 1980: 120

Onuphis punggolensis Tan & Chou, in press Singapore Tan & Chou, in press

Onuphis sp. Taiwan Hsieh, pers. eomm.

Nothria conchylega (Sars, 1835) S. Vietnam Gallardo 1968:75

Nothria holobranchiata (Marenzeller, 1879) S. Vietnam Gallardo 1968:75

Nothria nr. atlantisa Hartman, 1965 S. Vietnam Gallardo 1968:75

Onuphis eremita Audouin & M. Edw. 1833 S. Vietnam Gallardo 1968:76China Wu et al. 1980:120

Onuphis pseudodibranchiata Gallardo, 1968 S. Vietnam Gallardo 1968:76China Wu et al. 1980:120

Onuphis variolata Shen, 1987 China Shen 1987:222

Paradiopatra willemoesii (Mclntosh, 1885) China Wu et al. 1980:120

Orbiniidae

Haploscoloplos elongatus (Johnson, 1901) S. Vietnam Gallardo 1968:92China Yang & Sun 1988:204

Haploscoloplos ef. jragilis (Verrill, 1873) S. Vietnam Gallardo 1968:92

221



Leitoscoloplos elongatus Taiwan Hsieh, pers. eomm.

Leitoscoloplos ef. kerguelensis (McIntosh, 1885) Hong Kong Shin 1982:167

Naineris hainanensis Q. WU China Wu & Wu 1987:79

Orbinia ef. dicrochaeta Taiwan Hsieh, pers. eomm.

Orbinia vietnamensis Gallardo, 1968 S. Vietnam Gallardo 1968:93China Yang & Sun 1988:205

Phylo kupfferi (Eh1ers, 1875) Hong Kong Shin 1982: 167

Phylo ornatus (Verrill, 1873) China Yang & Sun 1988:206

Scoloplos cf. acmeceps Taiwan Hsieh, pers. eomm.

Scoloplos ef. armiger (Muller, 1776) China Wu et al. 1980:122

Scoloplos (Leodamas) ?dubia Tebble, 1955 S. Vietnam Gallardo 1968:94

Scoloplos (L.) gracilis Pillai, 1961 S. Vietnam Gallardo 1968:94

Scoloplos (L.) rubra (Webster, 1879) China Yang & Sun 1988:208

Scoloplos (L.) rubra orientalis Gallardo, 1968 S. Vietnam Gallardo 1968:95

Scoloplos cf. rubra Taiwan Hsieh, pers. eomm.

Scoloplos (Scoloplos) marsupialis Southern, 1921 China Yang & Sun 1988:209

Scoloplos (Scoloplos) spiniferus Gallardo, 1968 S. Vietnam Gallardo 1968:95

Scoloplos tumidus Maekie, 1991 Hong Kong Maekie 1991:38S. Vietnam Gallardo 1968:95 as S. marsupialis

Scoloplos cf. tumidus Taiwan Hsieh, pers. eomm.

Paraonidae

Aricidea cf. fragilis caeca Taiwan Hsieh, pers. eomm.

Paradoneis lyra (Southern, 1914) S. Vietnam Gallardo 1968:96

Spionidae

Aonides sp. China Wu & Sun 1978:187

Aonides ef. oxycephala Taiwan Hsieh, pers. eomm.

Dispio ef. uncinata Taiwan Hsieh, pers. eomm.

Laonice cirrata (M. Sars, 1851) S. Vietnam Gallardo 1968:97China Wu & Sun 1978:192Hong Kong Shin 1982:167

Malacoceros (Rh.) areninc.asiaticus (Ch1ebovitseh, 1959 ) China Wu & Sun 1978:189

Malacoceros indicus (Fauve1, 1928) China Wu et al. 1980:122Taiwan Hsieh, pers. eomm.

Microspio sp. China Wu & Sun 1978:191

Minuspio cirrifera (Wiren, 1883) Hong Kong Shin 1982:167

Nerine faughni Gallardo, 1968 S. Vietnam Gallardo 1968:97

Paraprionospio pinnata (Ehlers, 1901) Hong Kong Shin 1982: 167China Yang & Sun 1988:218

Paraprionospio sp. Taiwan Hsieh, pers. eomm.

Polydorella novaegeorgiae Gibbs, 1971 Hong Kong Shin 1982: 168

Polydora cf. pilikia Ward, 1981 China Yang & Sun 1988:213

Polydora cf. socialis (Sehmarda, 1861) Hong Kong Shin 1982: 167

Polydora ef. tentaculata Blake & Kudenov,1978 Hong Kong Shin 1982: 168Taiwan Hsieh, pers. eomm.

Polydora cavitensis Pillai, 1965 Philippines Pillai 1965:152

222



Polydora glycymerica Radashevsky, 1993 Manchenko & Radashevsky1994:767

Polydora ligni Webster, 1879 Taiwan Hsieh, pers. comm.

Polydora vulgaris Mohammad, 1972 Manchenko & Radashevsky1994:767

Prionmpio africana Augener, 1918 S. Vietnam Gallardo 1968:98

Prionospio depauperata Taiwan Hsieh, pers. comm.

Prionospio ehlersi Fauvel, 1928 S. Vietnam Gallardo 1968:98China Wu et al. 1980:122Hong Kong Shin 1982: 168

Prionospio japonica Taiwan Hsieh, pers. comm.

Prionospio komaeti Hylleberg & Nateew., 1991 Singapore Tan & Chou 1993:288

Prionospio malmgreni Claparede, 1870 China Wu & Sun 1978:189Hong Kong Shin 1982: 168

Prionospio malayensis Caullery, 1914 S. Vietnam Gallardo 1968:99Singapore Tan & Chou 1993:288

Prionospio paradisea Taiwan Hsieh, pers. comm.

Prionospio pinnata Eh1ers, 1901 S. Vietnam Gallardo 1968:99

Prionospio cf. pulchra Taiwan Hsieh, pers. comm.

Prionospio saccifera Mackie & H. 1990 Hong Kong Mackie & Hart1ey 1990:366

Prionospio sishaensis Wu & Chen, 1964 China Wu & Wu 1987:79

Pseudopolydora antennata Claparede, 1870 China Wu & Sun 1978:183

Pseudopolydora diopatra Hsieh, 1992 Taiwan Hsieh 1992:630

Pseudopolydora cf. kempi Taiwan Hsieh, pers. comm.

Pygospio elegans C1aparede, 1863 China Wu & Sun 1978:186

Scolelepis ? indica Fauvel, 1928 S. Vietnam Gallardo 1968:100

Scolelepis sp. Taiwan Hsieh, pers. comm.

Spiophanes bombyx (C1aparede, 1870) China Wu & Sun 1978:185

Spiophanes malayensis Caullery, 1915 China Wu et al. 1980:122

Spiophanes sp. Taiwan Hsieh, pers. comm.

Heterospionidae

Heterospio sinica Wu & Chen, 1966 China Yang & Sun 1988:221

Magelonidae

Magelona cf. cincta Taiwan Hsieh, pers. comm.

Magelona crenulifrons Gallardo, 1968 S. Vietnam Gallardo 1968:100Hong Kong Shin 1982: 168

Magelona japonica Okuda, 1937 S. Vietnam Gallardo 1968:101China Wu et a1 1980:122

Magelone koreana Okuda, 1937 S. Vietnam Gallardo 1968:101

Magelona lenticulata Gallardo, 1968 S. Vietnam Gallardo 1968: 101

Magelona cf. pacifica Taiwan Hsieh, pers. comm.

Magelona sp. China Wu & Sun 1978:195

Cossuridae

Cossura coasta Kitamori, 1960 Hong Kong Shin 1982: 169

Cossurella cf. diamorpha Taiwan Hsieh, pers. comm.

223



TrochochaetidaeTrochochaeta diverapoda (Hoagland, 1920) Hong Kong Mackie 1990:338

Trochochaeta cf. orissae (Fauvel, 1932) Hong Kong Shin 1982:168

PoecilochaetidaePoecilochaetus gallardoi Pilato & Cantone, 1976 S. Vietnam Pilato & Cantone 1976:43

S. Vietnam Gallardo 1968: 103 as P. sp. A

Poecilochaetus hystricosus Mackie, 1990 Hong Kong Mackie 1990:351

Poecilochaetus paratropicus Gallardo, 1968 S. Vietnam Gallardo 1968:102

Poecilochaetus serpens Alien, 1904 China Wu et aI 1980:122Singapore Tan & Chou 1993:288

Poecilochaetus spinulosus Mackie, 1990 Hong Kong Mackie 1990:342

Poecilochaetus tricirratus Mackie, 1990 Hong Kong Mackie 1990:346

Poecilochaetus tropicus Okuda, 1937 China Wu & Sun 1978:180

Poecilochaetus vietnamita Gallardo, 1968 S. Vietnam Gallardo 1968: 103

Poecilochaetus sp. Taiwan Hsieh, pers. comm.

Chaetopteridae

Chaetopterus variopedatus (Renier, 1804) Singapore Tan & Chou 1993:288China Yang & Sun 1988:224

Sasekumaria se/angora Rullier, 1976 Malaysia Ng & Sasekumar 1993:467

Spiochaetopterus costarum (Claparecte, 1868) Hong Kong Shin 1982: 168costarum

CirratulidaeCaulleriella glabra Gallardo, 1968 S. Vietnam Gallardo 1968: 104

Caulleriella cf. typhlops (Willey, 1905) China Sun et al 1978: 163

Chaetozone flagellifera Gallardo, 1968 S. Vietnam Gallardo 1968: 105

Chaetozone maotienae Gallardo, 1968 S. Vietnam Gallardo 1968:105

Cirratulus annamensis Gallardo, 1968 S. Vietnam Gallardo 1968: 105

Cirratulus cirratulus Muller, 1776 Singapore Tan & Chou 1993:288

Cirratulus filiformis Keferstein, 1862 China Yang & Sun 1988:234

Cirriformia chrysodermoides Pillai, 1965 Philippines Pillai 1965:156

Cirriformia cf. filiformis Taiwan Hsieh, pers. comm.

Cirriformia jiligera (Delle Chiaje, 1828) China Wu 1987:79Cirriformia punctata (Grube, 1859) China Wu et al. 1980:123

Cirriformia semicincta (Ehlers, 1905) S. Vietnam Gallardo 1968:106

Cirriformia tentaculata (Montagu, 1808) China Yang & Sun 1988:235Dodecaceria jewkesi Berkeley & Berk., 1954 China Yang & Sun 1988:236Tharyx filibranchia Day, 1961 Hong Kong Shin 1982: 169

Tharyx marioni (Saint-Joseph, 1894) S. Vietnam Gallardo 1968:106Tharyx sp. Taiwan Hsieh, pers. comm.

FlabelligeridaeBrada jerruginea Gallardo, 1968 S. Vietnam Gallardo 1968:107Diplocirrus erythroporus Gallardo, 1968 S. Vietnam Gallardo 1968: 108

Flabelliderma sp. Taiwan Hsieh, pers. comm.

224



Pherusa parmata (Grube, 1877) Hong Kong Mak 1982:600China Yang & Sun 1988:241

Pherusa curvisetis (Caullery, 1944) Philippines Gallardo 1968:108S. Vietnam Gallardo 1968:108

Pherusa eruca indica (Fauvel, 1928) S. Vietnam Gallardo 1968:109

Pherusa monroi (Day, 1957) Singapore Tan & Chou 1993:289

Piromis congoensis (Grube, 1877) S. Vietnam Gallardo 1968:109

Scalibregmidae

Scalibregma inflatum Rathke, 1843 S. Vietnam Gallardo 1968:110China Yang & Sun 1988:271

Opheliidae

Ammotrypane aulogaster Rathke, 1843 Singapore Tan & Chou 1993:289Ammotrypane ?dubia Caullery, 1944 S. Vietnam Gallardo 1968: 111

Ammotrypane ?ehlersi Horst, 1919 S. Vietnam Gallardo 1968:111Ammotrypane ?fauveli Caullery, 1944 S. Vietnam Gallardo 1968:111

Ammotrypane filobranchiata Gallardo, 1968 S. Vietnam Gallardo 1968:112Ammotrypane grandis Pillai, 1961 S. Vietnam Gallardo 1968:112

Armandia ?intermedia Fauvel, 1902 S. Vietnam Gallardo 1968: 113Armandia intermedia China Yang & Sun 1988:246Armandia lanceolata Willey, 1905 S. Vietnam Gallardo 1968: 113Armandia leptocirris (Grube, 1878) Philippines Grube 1878:194

S. Vietnam Gallardo 1968: 113China Wu & Wu 1987:79

Armandia ef. leptocirris Taiwan Hsieh, pers. eomm.Armandia longicaudata (Caullery, 1944) S. Vietnam Gallardo 1968:114Ophelina acuminata Oersted, 1843 China Yang & Sun 1988:248Ophelina ef. acuminata Taiwan Hsieh, pers. eomm.Polyophthalmus pictus (Dujardin, 1839) Philippines Pillai 1965:157

China Sun et al. 1978: 163

Sternaspidae

Sternaspis scutata (Renier, 1807) S. Vietnam Gallardo 1968:114China Wu et al. 1980:123Hong Kong Shin 1982:169Taiwan Hsieh, pers. eomm.

Capitellidae

Capitella capitata (Fabrieius, 1780) Singapore Tan & Chou 1993:289Capitella sp. Taiwan Hsieh, pers. eomm.Capitellides sp. Taiwan Hsieh, pers. eomm.Capitellethus branchiferus Gallardo, 1968 S. Vietnam Gallardo 1968:115Dasybranchus caducus (Grube, 1846) S. Vietnam Gallardo 1968:116

China Wu & Wu 1987:79Heteromastus filiformis (Claparede, 1864) China Yang & Sun 1988:254Heteromastus similis Southern, 1921 S. Vietnam Gallardo 1968: 117Mastobranchus loii Gallardo, 1968 S. Vietnam Gallardo 1968:118Mediomastus sp. Taiwan Hsieh, pers. eomm.

225



Branchiomma cf. boholense (Grube, 1878) Taiwan Hsieh, pers. comm.

Branchiomma cingulata (Grube, 1870) Philippines Pillai 1965:163Hong Kong Mak 1982:600China Yang & Sun 1988:299

Branchiomma cf. conspersa Ehlers, 1887) Taiwan Hsieh, pers. comm.

Chone ?infundibuliformis Krayer, 1856 S. Vietnam Gallardo 1968:130

Chone trilobata Gallardo, 1968 S. Vietnam Gallardo 1968:130

Hypsicomus phaeotaenia (Schmarda, 1861) China Wu et al. 1980:124Singapore Tan & Chou 1993:290

lasmineira caducibranchiata Willey, 1905 S. Vietnam Gallardo 1968:131

Laonome albicingillum Hsieh, 1995 Taiwan Hsieh 1995:131

Megalomma intermedium (Beddard, 1889) Philippines Pillai 1965: 164

Megalomma vesiculosa (Montagu, 1815) Hong Kong Mak 1982:600China Yang & Sun 1988:301

Megalomma sp. Taiwan Hsieh, pers. comm.

Myxicola infundibulum (Renier, 1804) Hong Kong Mak 1982:600

Notaulax cL pyrroghaster Taiwan Hsieh, pers. comm.

Potamilla leptochaeta Southern, 1921 Singapore Tan & Chou 1993:290

Potamilla ocalea Pillai, 1965 Philippines Pillai 1965:164

Potamilla torelli Ma1mgren, 1866 Hong Kong Shin 1982: 170

Pseudobranchiomma sp. Taiwan Hsieh, pers. comm.

Pseudopotamilla reniformis (Muller, 1771) Hong Kong Mak 1982:600Singapore Tan & Chou 1993;600

Pseudopotamilla sp. Taiwan Hsieh, pers. comm.

Sabella penicillus Linneus, 1767 China Wu et al. 1980:124

Sabella sp. Hong Kong Mak 1982:600

Sabellastarte indica (Savigny, 1818) S. Vietnam Gallardo 1968:131Singapore Tan & Chou 1993:290China Yang & Sun 1988:305Taiwan Hsieh, pers. comm.

Sabellastarte cf. samoensis (Grube, 1870) Taiwan Hsieh, pers. comm.

Sabellastarte sanctijosephi (Gravier, 1906) China Wu et a1 1980:124

Serpulidae

Apomatus enosimae Marenzeller, 1884 China Yang & Sun 1988:308

Circeis spirillum (Linnaeus, 1758) China Yang & Sun 1988:318

Dexiospira foraminosus Busch, 1904 China Yang & Sun 1988:319

Dexiospira sp. Taiwan Cheng & Chen 1994:

Hydroides albiceps Grube, 1869 Hong Kong Mak 1982:601China Yang & Sun 1988:311

Hydroides centrospina Wu & Chen, 1981 China Wu & Chen 1981:354

Hydroides dirampha March, 1863 Hong Kong Mak 1982:602

Hydroides elegans (Haswell, 1883) Hong Kong Mak 1982:602China Yang & Sun 1988:311

Hydroides cf. elegans Taiwan Hsieh, pers. comm.

Hydroides exaltata (Marenzeller, 1884) China Wu et al. 1980: 124Hong Kong Mak 1982:603

Hydroides fusca Imajima, 1976 China Wu et al. 1980:124

228



Hydroides cf. jusicola March, 1863 Hong Kong Mak 1982:603

Hydroides grubei Pillai, 1965 Philippines Pillai 1965:166

Hydroides inornata Pillai, 1960 Hong Kong Mak 1982:605

Hydroides longistylaris Chen & Wu, 1980 China Chen & Wu 1980:249

Hydroides nanhaiensis Wu & Chen, 1981 China Wu & Chen 1981:355

Hydroides norvegica Gunnereus, 1768 Philippines Pillai 1965: 168

Hydroides rhombobulus Chen & Wu,1980 China Chen & Wu 1980:247Hong Kong Mak 1982:605

Hydroides trilobulus Wu & Chen, China Wu et al. 1980:125

Metavermilia acanthophora (Augener, 1914) China Wu et al. 1980:125Hong Kong Mak 1982:606

Metavermilia annobonensis Zibrowius, 1973 China Wu et al. 1980:125

Metavermilia cf. inflata 1majima, 1977 Hong Kong Mak 1982:607

Metavermilia cf. taenia Zibrowius, 1973 China Wu et al. 1980:125

Neodexiospira cf. joraminosa Taiwan Hsieh, pers. comm.

Neodexiospira cf. preacuta (Bush, 1904) Taiwan Hsieh, pers. comm.

Neodexiospira cf. knightjonesi Taiwan Hsieh, pers. comm.

Neodexiospira cf. steueri (Sterzinger, 1909) Taiwan Hsieh, pers. comm.

Neopomatus uschakovi Pillai, 1965 Philippines Pillai 1965:170lingayanensis

Pileolaria cf. militaris C1aparecte, 1870 Taiwan Hsieh, pers. comm.

Pileolaria cf. pettibonei Taiwan Hsieh, pers. comm.

Pileospira koehleri (Caullery & Mesni1, 1897) China Wu et al. 1980: 125

Pomatoleios kraussii (Baird, 1865) Hong Kong Mak 1982:607Thailand Becker 1993

Pomatoleios kraussii manilensis Pillai, 1965 Philippines Pillai 1965:169

Pomatostegus stellatus (Abi1dgaard, 1789) Hong Kong Mak 1982:608

Protohydroides elegans (Haswell, 1883) Hong Kong Shin 1982: 170

Protula tubularia tubularia (Montagu, 1803) China Wu et al. 1980:125

Protula sp. Taiwan Hsieh, pers. comm.

Salmacina cf. dysteri (Hyx1ey, 1955) Taiwan Hsieh, pers. comm.

Serpula (Paraserpula) sinica Wu & Chen in Wu et al.,1979 China Wu et al. 1979:92

Serpula vermicularis Linnaeus, 1767 Hong Kong Mak 1982:608China Yang & Sun 1988:312

Serpula watsoni Willey, 1905 Hong Kong Mak 1982:609

Simplicaria pseudomilitaris (Thiriot-Quievreux, 1965) Taiwan Hsieh, pers. comm.

Spirobranchus giganteus (Pallas, 1766) China Wu et al. 1980: 125

Spirobranch. giganteus corniculatus ten Hove Taiwan Dai & Yang 1995:

Spirobranchus jousseaumei (Gravier, 1908) China Yang & Sun 1988:314

Spirobranchus latiscapus (Marenzeller, 1884) China Wu et al. 1980:125

Spirobranchus maldivensis Pixell, 1913 China Yang & Sun 1988:315

Spirobranchus semperi March, 1863 Hong Kong Mak 1982:609China Yang & Sun 1988:315

Spirobranchus tricornis (March, 1863) Hong Kong Mak 1982:609China Yang & Sun 1988:317

Spirorbis (Dexiospira) treadwelli Pillai, 1965 Philippines Pillai 1965: 172

Vermiliopsis glandigerous Gravier, 1908 China Wu et al. 1980:125

229



Vermiliopsis infundibulum (Linneus, 1788) China Wu et al. 1980:125

Vermiliopsis infundibul. glandigera Imajima, 1967 Hong Kong Mak 1982:610

Vermiliopsis pygidialis (Willey, 1905) China Wu et al. 1980:125

Nerillidae

Leptonerilla diplocirrata Westheide & Purschke 1996 China

LITERATURE CITED

Westheide & Purschke 1996:587

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