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Introduction to Pacific Northwest Ecosystems Introduction to PNW Ecosystems 1. Where are we? : global / continental position 2. An overview of our place: regional geography & landforms 3. How are landforms created? 4 Th i t f l t lti l l I. Physical & Chemical (Abiotic) Environment of WA 4. The importance of geology at multiple scales 5. Climate 1. Ecoregions 2. Ecoregions: a virtual field trip 3. Environmental determinants of ecoregions II. Ecological Zones of WA The Ecology of Washington 1. Global / Continental Position 2. Regional Geography & Landforms 3. Forces Behind Landforms I. Abiotic Environment of WA 3. Forces Behind Landforms 4. Geology 5. Climate I. Abiotic Environment of WA 1. Global / Continental Position A) Where are we ? Hokkaido Frankfurt Newfoundland Latitude : Bothell °N 1. Global / Continental Position A) Where is WA ? Continental Position : Coastal Maritime I. Abiotic Environment of WA Global / Continental Position B) What are the ecological implications of our position? I. Present-day Climate 1) Precipitation & Temperature It affects our 2) Daily & Seasonal Changes II. Past Environment 1) Past Climate 2) Geological history (and hence present day geology)

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•1

Introduction to

Pacific Northwest Ecosystems

Introduction to PNW Ecosystems

1. Where are we? : global / continental position

2. An overview of our place: regional geography & landforms

3. How are landforms created?

4 Th i t f l t lti l l

I. Physical & Chemical (Abiotic) Environment of WA

4. The importance of geology at multiple scales

5. Climate

1. Ecoregions

2. Ecoregions: a virtual field trip

3. Environmental determinants of ecoregions

II. Ecological Zones of WA

The Ecology of Washington

1. Global / Continental Position

2. Regional Geography & Landforms

3. Forces Behind Landforms

I. Abiotic Environment of WA

3. Forces Behind Landforms

4. Geology

5. Climate

I. Abiotic Environment of WA

1. Global / Continental Position A) Where are we ?

Hokkaido

FrankfurtNewfoundland

Latitude:

Bothell

°N

1. Global / Continental PositionA) Where is WA ?

Continental Position: Coastal Maritime

I. Abiotic Environment of WA Global / Continental PositionB) What are the ecological implications of our position?

I. Present-day Climate1) Precipitation & Temperature

2) D il & S l Ch

It affects our

2) Daily & Seasonal Changes

II. Past Environment 1) Past Climate

2) Geological history(and hence present day geology)

•2

Global / Continental PositionB) What are the ecological implications of our position?

I. Present-day Climate1) Precipitation & Temperature

A. Atmospheric circulationB Oceanic circulation

It affects our

B. Oceanic circulationC. Maritime influences

2) Daily & Seasonal Changes

II. Past Environment 1) Past Climate

2) Geological history

Atmospheric Circulation is a major determinant of global precipitation & temperature patterns

1. Sunlight energy greatest near equator

2. Results in warm, rising air at low latitudes

Campbell (2001)

2) Present-day Climate: Atmospheric Circulation

3. Rising air cools & rain falls abundantly at low latitudes

Atmospheric Circulation is a major determinant of global precipitation & temperature patterns

Campbell (2001)

Hadley Cell

2) Present-day Climate: Atmospheric CirculationAtmospheric Circulation is a major determinant of global precipitation & temperature patterns

4. Rising air leaves low pressure area behind.

Surface air from N and S flow into area

Campbell (2001)

into area.

Results in large-scale circular flow of air masses (Hadley Cells)

5. Hadley Cells create dry latitudes of descending air at about 30 °N & S

WA

WA sits at the

2) Present-day Climate: Atmospheric CirculationAtmospheric Circulation is a major determinant of global precipitation & temperature patterns

WA sits at the edge of another rising air mass region – hence

the tendency for higher levels of precipitation.

Campbell (2001)

2) Present-day Climate: Oceanic Circulation

Ocean currents determine the temperature of surface waters.

This has large i fl

Oceanic Circulation can be a major determinant of regional precipitation & temperature patterns

influences on coastal climates

Ricklefs (1997)

•3

ENSO EventsEl Niño (warm phase)

Warm surface water in the central & eastern Pacific

ENSO: El Nino – Southern Oscillation Events

La Niña (cool phase)

Cool surface water in the

UW CIG

ENSO PNW Climate ImpactsEl Niño (warm phase)

Warmer & drier winter

La Niña (cool phase)

Cooler & wetter winter

central & eastern Pacific

Events on “inter-annual” cycles

PDO: Pacific Decadal OscillationsSimilar to El Niño events, warm-phase PDO events show

cooling of north Pacific; warming of central, eastern Pacific

UW CIG

PDO – warm phase ENSO – warm phase

PDO: Pacific Decadal Oscillations

But PDO Oscillations are decades – long; rather than the annual variations of ENSO

2) Present-day Climate: Maritime Influences

Our coastal location allows large water body to moderate temperature extremes – a maritime climate

Our coastal position results in strong maritime influences on our regional precipitation & temperature patterns

Locations further from large water bodies experience larger seasonal temperature fluctuations –continental climates

Global / Continental PositionB) What are the ecological implications of our position?

I. Present-day Climate1) Precipitation & Temperature

2) Daily & Seasonal Changes

It affects our

2) Daily & Seasonal Changes

II. Past Environment 1) Past Climate

2) Geological history

Ecological implications

of seasonal variation in daylength

Daylength & Seasonality

# hours it is light

July: 17 hours of light

WA

Ricklefs (1997)

during 24-hour

periodCosta Rica

October: 12 hours of light

•4

Global / Continental PositionB) What are the ecological implications of our position?

I. Present-day Climate1) Precipitation & Temperature

2) Daily & Seasonal Changes

It affects our

2) Daily & Seasonal Changes

II. Past Environment 1) Past Climate

2) Geological history

White area:

Extent of glaciation about 18,000 YBP

Global Position

Past Climate

History of G l i l

Present Climate

Our location defines our past environments

Geological Processes

Past Organisms & Ecosystems: our biogeographical

template

Global / Continental PositionB) What are the ecological implications of our position?

I. Present-day Climate1) Precipitation & Temperature

2) Daily & Seasonal Changes

It affects our

2) Daily & Seasonal Changes

II. Past Environment 1) Past Climate

2) Geological history

Our Tectonic SettingOur location defines our past geological history

The Ecology of Washington

1. Global / Continental Position

2. Regional Geography & Landforms

3. Forces Behind Landforms

I. Abiotic Environment of WA

3. Forces Behind Landforms

4. Geology

5. Climate

A) Water Bodies:Marine

Regional Geography

Strait of Juan de FucaSan Juan IslandsStrait of Georgia

Pacific Ocean

Puget Sound

•5

A) Water Bodies:Freshwater

Major Streams of WA

Columbia

Nooksack

Skagit

SnohomishEast sideColumbia

Snake

West sideColumbiaCowlitz

Regional Geography

River

Snake River

Yakima River

Chehalis

Cowlitz

Nisqually

SnakeYakima

SpokaneOkanogan

CowlitzChehalisNisquallyPuyallupGreen

SnoqualmieSnohomish

StillaguamishSkagit

NooksackSkokomish

QuinaultHoh

A) Water Bodies:Marine / Freshwater

Major Estuaries of Western Washington

Grays Harbor(Chehalis River)

Will B

Regional Geography

Willapa Bay(Willapa & Naselle Rivers)

Nisqually RiverPuyallup RiverCedar / Green RiverSnohomish RiverStillaguamish RiverSkagit RiverNooksack RiverSkokomish River

1. WA Geography & Features

B) Landscape Units:Watersheds

I. Abiotic Environment of WA

WHAT IS A WATERSHED?

Watersheds

WATERSHED WHAT IS A WATERSHED?

Murdoch & Cheo (1999)

1. WA Geography & Features

B) Landscape Units:Watersheds

l bi

Nooksack Watershed

Snohomish Watershed

Major Watersheds of Washington

Columbia

Chehallis

I. Abiotic Environment of WA

Columbia River

Watershed

Willapa & Naselle

Nisqually

Puyallup

Cedar / Green

Snohomish

Stillaguamish

Skagit

Nooksack

Skokomish

1. WA Geography & Features

B) Landscape Units: Physiographic Regions

I. Abiotic Environment of WA

Note: These are arbitrary physiographic divisions for use in our class. Many different schemes exist.

•6

Physiographic Regions of WA

Olympic Mts

1-8,000 ft

San Juan / Puget SdIslands

Cascade Mountains4-9,000 ft

(volcanoes up to

14 000 ft)

Palouse Hills

Okanogan Plateau

6 – 8,000 ftSelkirk

Mts6-7,000 ft

Coastal Lowlands

Willapa Hills

(1-2,000 ft)

Puget Lowlands

14,000 ft)

Columbia Gorge

Columbia Basin

< 2,000 ft

< 2,000 ft

Blue Mts6 - 7,000 ft

1. Global / Continental Position

2. Regional Geography & Landforms

3. Forces Behind Landforms

I. Abiotic Environment of WA

3. Forces Behind Landforms

4. Geology

5. Climate

1. Building the Landscape: Tectonic ProcessesA) Terrane accretion

B) Folding & uplift

C) Volcanism

3. Forces Behind Landforms

2. Processes Reshaping the LandscapeA) Continental Ice

B) Mountain Glaciers

C) Water

D) Wind

Forces That Shape Our Land1. Building the Landscape: Tectonic Processes

Our Tectonic Setting

Subduction – the Pacific Plate is being forced down under the North American plate as it pushes eastward

Forces that Shape our Land1. Tectonic Processes

Alt & Hyndman (1995)

Protruding pieces of crust on the Pacific Plate are scraped off and accrete onto the shoreline (“terranes”)

(A) Creating Landforms: terrane accretion

Tectonic terrane:A piece of crustal rocks

Forces that Shape our Land1. Tectonic Processes

PNW coast2-300 MYBP

Terrane accretion

Complex geology

A piece of crustal rocks separated from its original plate

Alt & Hyndman (1995)

•7

North American

Plate

(B) Creating Landforms: folding & uplift

Forces that Shape our Land1. Tectonic Processes

Pacific Plate

Montgomery (1997)

Juan de Fuca Plate

Old Basin & Range Basalt Flows: 13 - 16 MYBP

Modern Cascade Volcanoes: 3 – 500,000 YBP

(C) Creating Landforms: volcanism

Forces that Shape our Land1. Tectonic Processes

Alt & Hyndman (1994)

(A) Continental Ice

Forces that Shape our Land2. Processes Reshaping the Land

Glacier 18,000 YBP

Seattle

Olympia

Kruckeberg (1991)

• Mountain carving

• Moraines

(B) Mountain Glaciers

Forces that Shape our Land2. Processes Reshaping the Land

1) Hill & valley local topography

Water & Ice interact with geology to create unique landscapes

2) Mountain valley topography

(C) Water

Forces that Shape our Land2. Processes Reshaping the Land Erosional forces interact with geology to define habitat diversity

in the Olympic MountainsOlympic Peninsula

Easily erodable

Siltstones & shales eroded away leaving gently angled sandstones to dominate. Topography gentle –

Core Sedimentary Rocks Volcanic Crescent Formation

Forces that Shape our Land

low habitat complexity

Resistant to erosion

Steeply angled basalts result in more rugged topography -different habitats & habitat complexity

Sedimentary Core Rocks

Volcanic Crescent Formation

McNulty (1996)

•8

1) Hill & valley local topography

3) Eastern WA scablands & coulees

2) Mountain valley topography

(C) Water

Forces that Shape our Land2. Processes Reshaping the Land

Great Floods reshaping the lands of Eastern WA (C) Water

Forces that Shape our Land2. Processes Reshaping the Land

Lake Missoula Floods15,000 YBP

Alt & Hyndman (1995)

1) Hill & valley local topography

3) Eastern WA scablands & coulees

2) Mountain valley topography

(C) Water

Forces that Shape our Land2. Processes Reshaping the Land

4) Columbia River gorge

5) River deltas (estuaries): rivers bring in and take away sedimenttides take away sediment

Wind-blown deposits (loess) & the Palouse Prairie(D) Wind

Forces that Shape our Land2. Processes Reshaping the Land

Loess

Rolling prairie from loess deposition over old basalt flows

1. Global / Continental Position

2. Regional Geography & Landforms

3. Forces Behind Landforms

I. Abiotic Environment of WA

4. Geology: a multi-scale perspective

5. Climate

I. Abiotic Environment 4. Geology: influences ecological systems at different

spatial scales

Large scale: Tectonics

Medium scale: Regional

Landform creation

Landform modification

Groundwater surface

Small scale: microhabitats

Modified from Montgomery (1997)

Groundwater – surface water connections

Surface rock diversity weathers into diverse soils

Boulders create unique microsites

Influences on erosion

Soil chemistry

•9

1. Global / Continental Position

2. Regional Geography & Landforms

3. Forces Behind Landforms

4 G l

I. Abiotic Environment of WA

4. Geology

5. Climate

Washington Climate1) Climate diagrams

Pre

cie (

°C)

1. Temperature

2. Precipitation

3. Drought

Jan Dec

ipita

tion

(mm

)Te

mp

era

ture

Temp > Precip

3 g

WA State: Mediterranean Climate

Washington Climate2) Climate

patterns Western WA: Maritime Climate

Eastern WA: Continental Climate

Washington Climate2) Climate patterns

Why are these different ?Kruckeberg (1991)

Spatial patterns in precipitation –

Washington Climate2) Climate patterns

precipitation –across WA State

Prevailing Storm Track in F ll Wi t &

Washington Climate2) Climate patterns

Fall, Winter & Spring

Kruckeberg (1991)

Inches of annual

precipitation

•10

R i h d R i h d

Washington Climate2) Climate patterns

Rainshadow Rainshadow

Topo

Cascade Crest

Washington Climate

2) Climate patterns

Precip

Precipitation Patterns are actually highly complex across mountains

What causes these rainshadows?

Washington Climate2) Climate patterns

Campbell (2001)

Prevailing Storm Track in

Washington Climate

3) Local variations in climate patterns

Storm Track in Fall, Winter &

Spring

Kruckeberg (1991)

cm annual precipitation

South North

1520 m1523 m

Summer precipitation at 6 meteorological stations (1993)Olympic Mountain Study – Andrea Woodward

54 mm168 mm

Washington Climate3) Local variations in climate patterns

1520 m

1463 m

1404 m

1503 m

1453 m100 mm

61 mm 91 mm

84 mm

Bottom Line: beware of using general weather station for specific sites, especially in mountainous terrain

The Ecology of Washington

1. Where are we? : Global / Continental Position

2. An Overview of our Place: Regional Geography & Landforms

3. How are Landforms Created?

4 Th I t f G l t M lti l S l

I. Physical & Chemical (Abiotic) Environment of WA

4. The Importance of Geology at Multiple Scales

5. Climate

II. Ecological Zones of WA

•11

II. Ecological Zones of WA

1. Ecoregions

2. Ecoregions: a virtual field trip

3. Environmental Determinants of Ecoregions

II. Ecosystems of WA

1. Ecoregions

1. Marine Shoreline

2. Sitka Spruce

3. Western Hemlock

4. Silver Fir

5. Mountain Hemlock

6. Subalpine Fir

7. Alpine

8. Douglas-fir / Grand Fir

9. Ponderosa Pine

10. Shrub Steppe

11. Palouse Prairie

11 Ecoregions of Washington State

Washington State Ecoregions

West-sideMontane

ToAlpine

Marine Shoreline

West-side Montane – Alpine: Silver fir; Mountain hemlock; Subalpine fir; Alpine

Sitka Spruce

Western Hemlock

WESTEAST

Mountain hemlock

Sub alpine fir

Alpine

Washington State Ecoregions

Montane to Alpine Ecoregions

SeattleYakima

Silver fir

Washington State Ecoregions

West-sideMontane

ToAlpine

Douglas-fir /Grand fir

Marine Shoreline

Palouse Prairie

Shrub Steppe

West-side Montane – Alpine: Silver fir; Mountain hemlock; Subalpine fir; Alpine

Sitka Spruce

Western Hemlock

Ponderosa Pine

Washington State Natural Regions

Similar scheme to that we are using.

Note some prominent differences:

• Eastern WA prairie / shrub-steppe distribution

• Distinction of prairie

from “Our Changing Nature”

WA Dept. of Natural Resources 1998

• Distinction of prairie woodland mosaic in western WA

• Discontinuity of high elevation forests/alpine

•12

The Ecology of Washington

1. Where are we? : Global / Continental Position

2. An Overview of our Place: Regional Geography & Landforms

3. How are Landforms Created?

4 Th I t f G l t M lti l S l

I. Physical & Chemical (Abiotic) Environment of WA

4. The Importance of Geology at Multiple Scales

5. Climate

1. Ecoregions

2. Ecoregions: a virtual field trip

II. Ecological Zones of WA

The Ecology of Washington

1. Where are we? : Global / Continental Position

2. An Overview of our Place: Regional Geography & Landforms

3. How are Landforms Created?

4 Th I t f G l t M lti l S l

I. Physical & Chemical (Abiotic) Environment of WA

4. The Importance of Geology at Multiple Scales

5. Climate

1. Ecoregions

2. Ecoregions: a virtual field trip

3. Environmental Determinants of Ecoregions (terrestrial)

II. Ecological Zones of WA

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Bottom Line

Major determinants of ecoregion distribution:

I. Precipitation• Amount

Ti i• Timing

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Bottom Line

Major determinants of ecoregion distribution:

I. Precipitation• Amount

• Timing

II. Temperature• Direct effectsBEWARE OF MEAN VALUES!

•13

II. Ecosystems of WA2. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm

II. Ecosystems of WA2. Environmental Determinants of Terrestrial Ecoregions

Bottom Line

Major determinants of ecoregion distribution:

I. Precipitation• Amount

• Timing

III. Interactive Effects of Temperature & Moisture

M i t ff t bilit tII. Temperature

• Direct effects

• Moisture effects ability to cope with temperature

II. Ecosystems of WA2. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm

II. Ecosystems of WA2. Environmental Determinants of Terrestrial Ecoregions

Bottom Line

Major determinants of ecoregion distribution:

I. Precipitation• Amount

• Timing

III. Interactive Effects of Temperature & Moisture

• Moisture effects ability to cope with t t

II. Temperature• Direct effects

temperature

• Temperature effects moisture availability

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm

•14

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Major determinants of ecoregion distribution:

I. Precipitation• Amount

• Timing

II. Temperature

III. Interactive Effects of Temperature & Moisture

• Moisture effects ability to cope with temperature

• Temperature effects moisture• Direct effects

Temperature effects moisture availability

temp water use water available

temp evaporation from soil water available

temp precip as snow water available

II. Ecosystems of WA3. Environmental Determinants of Terrestrial Ecoregions

Ecoregion Elevation Range (ft.)

Avg. Annual Temp (°F)

Avg annual precip(cm)

(Seattle) for reference 0 53 86

Sitka Spruce 0 – 500 52 200 – 300

Western Hemlock 0 – 2500 47 70 – 300

Silver Fir 1900 – 4200 42 220 – 280

M t i H l k 4200 5900 39 160 280Mountain Hemlock 4200 – 5900 39 160 - 280

Subalpine Fir 4200 - 5800 39 100 - 150

Alpine >5000 - >7000 37.5* 46*

Douglas-fir/Grand Fir 2000 – 5000 46 60 – 110

Ponderosa Pine 2000 – 4000 47 40 – 70

Shrub Steppe 150 – 2000 50 15 – 25

Palouse Prairie < 3000 48 40 – 70

* Data from Paradise R.S. on Mt. Rainier (subalpine zone) /

precip includes average snowfall of 256 cm