ANNALES HISTORICO-NATURALES MUSEI NATIONALIS HUNGARICI Volume 95 Budapest, 2003 pp. 19-35.

Notes on Dermestidae (Coleoptera) with description of eight new species*

J. HÁVA

Brönicke 13, CZ-147 00 Praha 4, Czech Republic e-mail: jh.dermestidae@volny.cz

Abstract - Anthrenus (Anthrenodes) endroedyi sp. n. (Ghana), Anthrenus (Anthrenus) merkli sp. n. (Tanzania), Anthrenus (Anthrenus) kantneri sp. n. (Zimbabwe, Malawi), Attagenus endroedyi sp. n. (Ghana), Attagenus incognitus sp. n. (Spain), Attagenus globosus sp. n. (Panama), Orphinus (Orphi-nus) apicebrunneus sp. n. (Ghana) and Phradonoma endroedyi sp. n. (Ghana) are described, illus­trated and compared with closely related species. Attagenus kocheri nom. n. is proposed as replace­ment name for Attagenus antennatus ESCALERA, 1914, nec C A S T E L N A U , F. L . N. C. D E L A P O R T E ,

1840. New records extending the known geographical distribution are published for the following species: Anthrenus (Anthrenodes) umbellatarum C H O B A U T , 1898 (Egypt); Anthrenus (Anthrenus) rotundulus R E I T T E R , 1889 (Afghanistan); Anthrenus (Nathrenus) basilewskyi K Á L I K , 1965 (Zam­bia); Anthrenus (Nathrenus) verbasci ( L I N N A E U S , 1767) (Chile); AttagenusatricolorWC, 1931 (Ma­lawi); Attagenus lobatus R O S E N H A U E R , 1856 (Romania); Attagenus pellio ( L I N N A E U S , 1758) (Ghana); Attagenus similaris M U L S A N T et R E Y , 1868 (Tunisia); Dermestes (Dermestes) bicolor F A B R I C I U S , 1781 (Iraq); Dermestes (Dermestes) schneiden H Á V A , 2002 (Mongolia); Dermestes (Dermestinus) maculants D E G E B R . 1774 (Ghana); Dermestes (Dermestinus) szekessyi K A L Í K , 1950 (Mongolia); Globicornis (Dearthrus) bodemeyeri ( G A N G L B A U E R , in B O D E M E Y E R , 1900) (Iran); Orphinus (Orphinus) congoanus (PlC, 1950) (Ghana); Globicornis (Globicornis) rufoguttata P I C , 1899 (Israel); Phradonoma nobile ( R E I T T E R , 1881) (Jordánia); Trogoderma angustum ( S O L I E R ,

1849) (Argentina). With 37 figures.

Key words - Coleoptera, Dermestidae, taxonomy, new species, nomenclature, distribution.

I N T R O D U C T I O N

During the determination of material coming from the following collections, I

found eight new species of the family Dermestidae and several other species repre­

senting new country records.

This paper is dedicated to the well-known entomologist and my friend, the late Dr. S E B A S T I A N

E N D R Ő D Y - Y O U N G A (1934-1999) who collected an important part of the material investigated between 1967 and 1970 in Ghana.

Label data of the type specimens are put in quotation marks. Type specimens are tagged with a red, printed label with the following text: "HOLOTYPE [or PARATYPE] species name sp. n. Jiff Hava det. 2003".

The following abbreviations refer to the collections in which the examined material is depos­ited: AHCD = collection of ANDREAS, HERRMANN, Stade, Germany; JHAC = collection of JlRÍ HÁVA, Praha, Czech Republic; JVAC = collection of JlRÍ VÁVRA, Ostrava, Czech Republic; HNHM = Hun­garian Natural History Museum, Budapest, (O. MERKL), Hungary; MRAC = Royal Museum for Central Africa, Tervuren (M. de MEYER), Belgium; NHMI = Natural History Museum, Tel Aviv (V. CHIKATUNOV), Israel; Z M A N = Zoologisch Museum, Amsterdam (B. BRUGGE), The Netherlands.

N O M E N C L A T O R I A L C H A N G E

Attagenus kocheri nom. n.

Attagenus antennatus ESCALERA, 1914:137, nec CASTELNAU, F. L. N. C. de LAPORTE, 1840: 36. Attagenus escalerai KOCHER, 1956:25, nec MADER, 1954: 53.

Remarks -Attagenus antennatus ESCALERA, 1914 is a junior secondary hom­

onym of Attagenus antennatus CASTELNAU, F. L . N . C. DE LAPORTE, 1840. Its re­placement name, Attagenus escalerai KOCHER, 1956 is a primary homonym of

Attagenus escalerai MADER, 1954, so another replacement name, Attagenus kocheri nom. n. is proposed hereby.

FAUNISTICS

Anthrenus (Anthrenodes) umbellatarum CHOBAUT, 1898

Material examined - "Aegyptus, Cairo", J. HÁVA, det. (6 specimens, HNHM, JHAC); "Aegyptus, Faggala", J. HÁVA det. (4 specimens, HNHM, JHAC).

Distribution - This species is known from Algeria, Morocco and Israel. First record from Egypt.

Anthrenus (Nathrenus) basilewskyi KALÍK, 1965

Material examined - "Zambia NE, 60 km N Mpika Mts., 6.x.2002, J. Haiada Igt.", J. HÁVA det. (2 males, 1 female, JHAC).

Distribution - This species is known from Tanzania, Malawi and Kenya. First record from Zambia.

Anthrenus (Anthrenus) rotundulus REITTER, 1 8 8 9

Material examined - "Afghanistan, Hari Rnd, 15 km SE Herat, 950 m, 21.v. 1974, No. 111, L . Papp leg.", J. HÁVA det. (2 males, HNHM, JHAC); "Afghanistan, Herat, Hari Rud river, about 8 km SW Herat, 950 m, 20.V.1974, No. 120, L. Papp leg.", J. HÁVA det. (2 males, 1 female, HNHM, JHAC).

Distribution - This species is known from Caucasus, Iran and Turkmenistan. First record from Afghanistan.

Anthrenus (Nathrenus) verbasci (LINNAEUS, 1767)

Material examined -"Chile, prov. Santiago, El Arrayan, 9.x. 1965, Hungarian Soil-Zool. Exp., Nr. P-B.40, Loksa Igt.", J. HÁVA det. (1 specimen, HNHM).

Distribution - Cosmopolitan species. First record from Chile.

Attagenus atricolor PIC, 1931

Material examined - "Malawi S, Balaka env., 5-6.Í.2002, J. Bezdëk Igt.", J. HÁVA det. (11 specimens, JHAC).

Distribution - This species is known from Botswana and Congo (MROCZKOWSKI 1968). First record from Malawi.

Attagenus lobatus ROSENHAUER, 1856

Material examined - "Romania centr., Bucuresti env., 20 km W, 26.vi. 1996, J. Vavra Igt.", J. HÁVA det. (1 female, JVAC).

Distribution - This species is known from South Europe, North Africa, U.S.A., Afghanistan, China, India, Iran, Iraq, Mongolia, Saudi Arabia, Tadzhikistan, Turkmenistan, Uzbekistan (MROCZ­KOWSKI 1968). First record from Romania.

Attagenuspellio (LINNAEUS, 1758)

Material examined - "Ghana, Takoradi, iv/xi. 1967, Besnard Igt.", J. H Á V A det. (2 males, 1 fe­male, MRAC).

Distribution - Cosmopolitan species. First record from Ghana.

Attagenus similaris M U L S A N T et REY, 1868

Material examined - "Tunisia, Tozeur prov., Degache Oasis, 3.iv. 1994, Nádai L. Igt.", J. HÁVA det. (12 specimens, JHAC, HNHM).

Distribution - This species is known from Algeria ( M R O C Z K O W S K I 1968). First record from Tunisia.

Dermestes (Dermestes) bicolor FABRICIUS, 1781

Material examined - "Iraq occ, Western desert, Wádí al Hazimi, 6.iv. 1979, J. Macek Igt.", J. HÁVA det. (1 female, JHAC).

Distribution - This species is known from the Palaearctic region. First record from Iraq.

Dermestes (Dermestes) schneiden HÁVA, 2002

Dermestes (Dermestes) schneiden HÁVA, 2002: 35 Dermestes (Dermestes) bicolor: ZHANTIEV, 1973: 187.

Material examined - "Mongolia, Siidgobi aimak, Bajan zag, 20 km NO von Somon Bulgan, 1200 m, Exp. Dr. Z. Kaszab, 1967, Nr. 895, 7.VÜ.1967", Dermestes bicolor Zhantiev det. 1973, J. HÁVA revid. (1 female, HNHM).

Remarks - This species was described on the basis of males; the female is

similar to the male but sternite 4 without median tuft of short erected setae. The

above-mentioned specimen was determined by ZHANTIEV, (1973) as Dermestes

bicolor FABRICIUS, 1781 representing new record to Mongolia. However, the

specimen belongs to Dermestes schneiden HÁVA. Dermestes bicolor is still un­

known from Mongolia.

Distribution - This species was described from Mongolia.

Dermestes (Dermestinus) maculatus D E GEER, 1774

Material examined - "Ghana, Ashanti, Kumasi, 330 m, N 6.43 W 1.36, light trap, 24.6.1967, no. 229, S. Endrődy-Younga Igt.", J. HÁVA det. (2 specimens, HNHM); "Ghana, Ashanti region, Kumasi, Nhiasu, 320 m, N 6.43 W 1.36, Dr. S. Endrődy-Younga Igt., Nr. 294, 13.Í.1968", J. HÁVA det. (3 specimens, HNHM).

Distribution - Cosmopolitan species. First record from Ghana.

Dermestes (Dermestinus) szekessyi KALÍK, 1950

Material examined - "Mongolia, Chovd Aimak, Charus Nur, N & W. Seite, 11-13.viii. 1978, L. & Zs. Peregi Igt.", J. HÁVA det. (1 male, HNHM).

Distribution - This species is known from Europe to Byelorussia, Ukraina, Caucasus, Kazakhstan, Russia (Novosibirskaya obi.) (MROCZKOWSKI 1968). First record from Mongolia.

Globicornis (Dearthrus) bodemeyeri (GANGLBAUER, i n BODEMEYER, 1900)

Material examined-"Iran, Mazandaran prov., 10 km SW of Reine, 2600 m, 14. v.2001, Bene­dek B. & Csorba G. leg.", J. HÁVA det. (1 female, HNHM).

Distribution - This species is known from Turkey and Syria. First record from Iran.

Globicornis (Globicornis) rufoguttata PIC, 1899

Material examined - "Israel, Mer. Golan, 6.v. 1979, D. Furth Igt.", J. HÁVA det. (1 female, JHAC); "Israel, Newe Ativ, 26.iv.1974, D. Furth Igt.", J. HÁVA det. (1 female, NHMI).

Distribution - This species is known from Syria. First record from Israel.

Orphinus (Orphinus) congoanus (PIC, 1950)

Trododerma (sic!) congoana Pic, 1950: 356. Orphinus congoanus: KALÍK, 1955: 319.

Material examined - "Congo, Brazzaville, ORSTOM park, Soil-Zoological Exp., 25.12.1963, No 513, light trap, Endrődy-Younga Igt.", J. HÁVA det. (1 female, HNHM); "Ghana, Ashanti, Kumasi, 330 m, N 6.43 W 1.36, light trap, 24.6.1967, no. 229, Endrődy-Younga Igt", J. HÁVA det. (2 males, 1 female, HNHM, JHAC); "Ghana, Western region, Busua, Dr. Endrődy-Younga leg., No. 59, geeinzelt, 14-16.ix. 1970", J. HÁVA det. (2 females, HNHM, JHAC).

Distribution - This species is known from Congo and Kenya (DELOBEL & TRAN, 1993). First records from Ghana.

Phradonoma nobile (REITTER, 1881)

Material examined - "O. Jordan, Wadi Schaib, 200 m, 3.10.1958, J. Klapperich Igt.", J. HÁVA det. (1 specimen, HNHM).

Distribution - This species is known from Cyprus, Greece, Portugal, Spain, Algeria, Egypt, Eritrea, Libya, Morocco, South Africa, Tunisia, Zimbabwe, Afghanistan, Caucasus, India, Iran, Iraq, Saudi Arabia, Syria, Tadzhikistan, Turkmenistan and Uzbekistan (MROCZKOWSKI 1968). First re­cord from Jordánia.

Trogoderma angustum (SOLIER, 1849)

Material examined - "Argentina, Misiones Eldorado, 20.xi. 1982, A. Kovács leg.", J. HÁVA det. (1 male, HNHM).

Distribution - This species is known from Europe, Chile and the U.S.A.. First record from Ar­gentina.

DESCRIPTIONS

Anthrenus (Anthrenodes) endroedyi sp. n. (Figs 1-3)

Type mate rial - Holotype (male): "Ghana, Ashanti Kwadaso, 320 m, N 6.42 W 1.39, flowering Citrus, 28. 2. 1967, no. 200, leg. Endrődy-Younga". Paratypes (3 females): same data as holotype. Holotype and 2 paratypes are deposited in HNHM, 1 paratype in JHAC.

Figs 1-5. 1-3. Anthrenus (Anthrenodes) endroedyi sp. n.: 1 = antenna of male, 2 = terminal antennái segment of female, 3 = male genitalia. 4-5. A. (A.) poggiiHÁVA, 2002: 4 = antennái club of male, 5 =

male genitalia

Description - Male, body length 2.3 mm, maximum width 1.4 mm; body oval, rust-brown. Dorsal surfaces covered by yellow and brown scales. Individual scales narrow. Head covered by in­termixed yellow and brown scales. Frontal median ocellus present. Antennae 11-segmented, brown, segment 9 black (Fig. 1). Eye with median margin entire. Pronotum covered by yellow scales on lat­eral portions, disc with brown scales. Each elytron with three bands of yellow scales; one on the hu­merus, one in the middle and one near the apex. Ventral surfaces covered by yellow scales; abdominal ventrites bearing small spots of brown scales at antero-lateral margins. Ventrites 1-4 without spots in the middle covered of brown scales. Prosternum with yellow scales only. Meta-ventrite with yellow scales, without large patch at lateral margins. Legs brown with yellow scales and yellow setae. Male genitalia: Fig. 3.

Female. Similar to male; antennái club: Fig. 2. Variability - Body length 2.3-2.7 mm, width 1.4-1.8 mm.

Etymology - The species is dedicated to the late Dr. S. ENDRŐDY-YOUNGA.

Differential diagnosis - The subgenus Anthrenodes CHOBAUT, 1898 includes 12 species or subspecies from the Palaearctic region; only one species, Anthrenus (Anthrenodes) poggii HÁVA, 2002 is known from the Afrotropical region (Soma­lia) (male antennái club: Fig. 4; male genitalia: Fig. 5). The new species differs from the latter in the form of the antenna and male genitalia.

Anthrenus (Anthrenus) kantneri sp. n. (Figs 6-7)

Type material - Holotype (male): "Zimbabwe, Lake Kariba, Zebra Island, 22.iii.2000, R. Beenen Igt." (ZM AN). Paratypes: 5 specimens: same data as holotype (ZMAN, JHAC); 6 specimens: "Zimbabwe, road Binga-Karoi, 18.iii.2000, R. Beenen Igt." (ZMAN); 6 specimens: "Zimbabwe, Bubi river vail., 70 km N of Beitbridge, 8.XÜ.1998, F. Kantner Igt." (JHAC, MRAC); 5 specimens: "Zimbabwe, 50 km W of Chimanimani, 14-15.xii.1998, S. Beevár Igt." (JHAC).

Non-type material - "Malawi centr., Salima env., 5-6.i.2002, J. Bezdëk Igt.", J. HÁVA det. (7 specimens, JHAC); "Malawi Nkhotakota, lake Malawi, 2.Í.2002, J. Haiada Igt.", J. HÁVA det. (4 specimens, JHAC).

Description - Male. Body length 2.8-3.1 mm, width 1.7-2.1 mm; body oval, black. Dorsal sur­face covered by black, white and brown scales. Head covered by intermixed white and brown scales. Pronotum covered by white scales on lateral margins; disc covered by intermixed black and brown scales. Elytra covered by intermixed black and brown scales, white scales forming five small patches on each elytron. Individual scales with sides parallel or slightly diverging from proximal 1/3 or mid­dle. Antennae 11-segmented, brown, antennái club 3-segmented, compact (Fig. 6). Eye with median margin entire. Ventral surface covered with white and brown scales, abdominal ventrites bearing small spots of brown scales at antero-lateral margins. Ventrites I—IV without spots in the middle. Prosternum with brown scales only. Metaventrite with white and brown scales, without a large patch at lateral margins. Legs brown with brown scales and white setae. Male genitalia: Fig. 7. Female body length 2.8-3.1 mm, width 1.7-2.1 mm; similar to the male.

Variability - Arrangement of the brown and black scales of elytra is quite variable. White scales forming small patches are variable in size.

Etymology - Dedicated to my friend Mr. F R A N T I S E K K A N T N E R (Ceské Budejovice, Czech Re­public) who is one of the collectors of the new species.

Remarks - The new species is similar to Anthrenus merkli sp. n., but differs in the form of antennae and male genitalia. A. merkli sp. n. is covered on dorsal sur­

faces with grey and brown scales; A. kantneri sp. n. is covered on dorsal surfaces

with black, white and brown scales.

Anthrenus (Anthrenus) merkli sp. n. (Figs 8-9)

Type material - Holotype (male): "Tanganyika, 1.5 miles to Lake Sereri, E shore, 3150 feet"; "singled material, 11-27.viii.1965, coll. Dr. J. Szunyoghy" (HNHM). 23 Paratypes: same data as holotype (15, HNHM, 8, JHAC).

Description - Male. Body length 2.5-3.0 mm, width 1.4-1.8 mm; body oval, black. Dorsal sur­face covered by grey and brown scales. Head covered only grey scales. Scales of pronotum grey on lateral margins, brown on disc. Elytra with intermixed grey and brown scales; brown scales forming

Figs 6-9. 6-7. Anthrenus (Anthrenus) kantneri sp. n.: 6 = antennái club of male, 7 = male genitali 8-9. A. (A.) merkli sp. n.: 8 = antenna of male, 9 = male genitalia

spots on each elytron. Individual scales widest at about proximal 1/3, parallel-sided or slightly taper­ing toward apex. Antennae 11-segmented, brown, antennái club 3-segmented, compact (Fig. 8). Eye with median margin entire. Ventral surface covered with grey scales, abdominal ventrites bearing small spots of brown scales at antero-lateral margins. Ventrites I - I V without spots in the middle. Prosternum with grey scales only. Metaventrite with grey scales only, without a large patch at lateral margins. Legs brown with grey scales and white setae. Male genitalia: Fig. 9. Female body length 2.7-3.1 mm, width 1.4—1.8 mm; similar to the male, but differs in the form of antennae, terminal seg­ment being short and rounded.

Etymology - This species is dedicated to my friend Dr. OTTÓ MERKL (HNHM), a known spe­cialist in the family Tenebrionidae (Coleoptera).

Remarks - The new species is similar to Anthrenus flavipes L E C O N T E , 1854, but differs in the form of antennae and male genitalia. Dorsal surfaces are covered by grey and brown scales in A . merkli sp. n.; by white, yellow and black scales in A . flavipes.

Figs 10-15. 10-13. Attagenus incognitas sp. n.: 10 = antenna of male, 11 = antennái club of female, 12 = male genitalia in dorsal view, 13 = elytral pattern of white pubescence. 14-15. A. pantherinus

(AHRENS, 1814) antennae: 14 = male, 15 = female (both according to MROCZKOWSKI 1975).

Attagenus incognitus sp. n.

(Figs 10-13)

Type material - Holotype (male): "Spain, Madrid, I .vii. 1973, M . Tóth Igt." Paratypes, Nos 1-22: same data as holotype; Nos 23-26: Spain, Villovieco/ Palencia, Camino de Santiago, 13.vii. 1999, P. Bahillo Igt. Holotype and paratypes Nos 1 -16 are deposited in HNHM, Nos 17-22 in JHAC, Nos 23-26 in AHCD.

Description - Male. Body length 4.5 mm, maximum width 2.0 mm; cuticle black-brown on dorsal surfaces, black on ventral surfaces; generally large and elongate. Head finely punctate with long black pubescence. Palpi entirely brown; pubescence on mentum denser. Ocellus on front pres­ent. Antennae 11-segmented, antennái club 3-segmented (Fig. 10), dark brown. Pronotum finely punctate as head, with long black pubescence on disc and long white pubescence in posterior angles. Scutellum triangular, finely punctate as pronotum, with short brown pubescence. Elytra finely punctate; cuticle black-brown with three orange-red bands covered by long white pubescence; other parts with long brown pubescence (Fig. 13). Legs brown with yellow pubescence; tibiae with short brown thorns. Meso- and metaventrite with short white pubescence. Abdominal ventrites with long white pubescence. Male genitalia: Fig. 12. Female body length 4.5 mm, maximum width 2.1 mm; similar to male but differs in the form of antennae (Fig. 11).

Variability - Length of males 4.5-4.8 mm, width 2.0-2.4 mm; length of females 4.5^4.9 mm, width 2.1-2.5 mm.

Etymology - Latin incognitus (= unknown).

Remarks - Attagenus incognitus sp. n. is very similar to A. pantherinus (AHRENS, 1814). Differential diagnosis of these two species is in the following

key:

1(2) Antennae 11-segmented in both sexes; antennái club 3-segmented, dark brown (Figs 10-11); pronotum with black and white pubescence; each

elytron with 3 orange-red bands covered by white pubescence (Fig. 13);

male genitalia as in Fig. 12; Spain A. incognitus sp. n.

2(1) Antennae 11-segmented in both sexes; antennái club 3-segmented, black (Figs 14-15); pronotum with intermixed black, white and yellow pubes­cence; each elytron with 4 bands and several spots with white and yellow pu­bescence; Europe, Turkey, Caucasus A. pantherinus (AHRENS, 1814)

Attagenus endroedyi sp. n. (Figs 16-19)

Type material- Holotype (male): "Ghana, Ashanti region, Kumasi, Nhiasu, 320 m, N 6.43 W 1.36, Dr. S. Endrődy-Younga Igt."; "Nr. 294, beaten, 13.i. 1968". Paratypes (4 males, 1 female): same data as holotype; (2 males): "Ghana, Ashanti Kwadaso, 320 m, N 6.42 W 1.39"; "flowering Citrus,

28. 2. 1967, no. 200, leg. Endrődy-Younga". Holotype and 4 paratypes are deposited in HNHM, 3 paratypes in JHAC.

Description - Male. Body (Fig. 16) small, oval, convex. Cuticle bicoloured; head and pronotum black or very dark brown, scutellum and elytra unicolorous dark brown. Body length 2.8 mm, width 1.6 mm. Head finely punctate with long brown pubescence. Palpi entirely yellow; pubes­cence on mentum denser. Frontal median ocellus present. Eye large, strongly convex. Antennae brown, 11-segmented, antennái club 3- segmented, dark brown, apical antennái segment elongate (Fig. 17). Pronotum finely punctate as head with long brown pubescence. Scutellum triangular with short brown pubescence. Elytra finely punctate; cuticle of each elytron dark brown, without bands of pubescence; pubescence entirely yellowish-brown. Epipleuron in anterior part broad with dense punctures and terminating gradually before posterior margin of metepisternum. Legs brown with yel­low pubescence; anterior tibiae with short red-brown spines. Ventral surface with long yellow­ish-brown pubescence. Trochantin exposed; lateral regions of prosternum narrow, not forming knife-like carina. Abdominal ventrites with long yellow pubescence. Male genitalia: Fig. 19. Female. Body length 2.8 mm, width 1.8 mm; antenna! club: Fig. 18.

Variability - Body length 2.5-2.8 mm, width 1.3-1.8 mm.

Etymology - The species is dedicated to the late Dr. S. Endrődy-Younga.

Figs 16-28. 16-19. Attagenus endroedyi sp. n.: 16 = body outline, 17 = antenna of male, 18 = anten­nái club o f female, 19 = male genitalia. 20-28. Antennae of Attagenus spp. (according to HALSTEAD 1981). A. insidiosus HALSTEAD, 1981: 20 = antenna of male, 21 = antenna o f female; A. nigripennis ARROW, 1915: 22 = antenna o f male, 23 = antenna of female; A atricolor?\C, 1931: 24 = antennái club o f female; A. smirnovi ZHANTIEV, 1973: 25 = antenna of male, 26 = antenna of female; A.

unicolor unicolor (BRAHM, 1791): 27 = antenna of male, 28 = antenna of female

Remarks - The new species is very similar to Attagenus smirnovi ZHANTIEV, 1973 known from the Afrotropical and Palearctic regions (HALSTEAD 1981). Main morphological differences are shown in the following key:

1 (8) Trochantin more or less concealed by an auricle-like lobe of hypomeron; lat­eral regions of prosternum broad, slightly or very strongly raised in front of procoxae (in latter case forming knife-like carinae).

2 (7) Lateral regions of prosternum strongly raised (appearing narrow) and form­ing knife-like carinae in front of coxae.

3 (4) Protibia more elongate, protarsus with 2nd segment about half as long as 5th; dorsal pubescence unicolorous black; antenna: Figs 20-21; female an­tenna with apical segment more elongate; mesoventrite elongate; length 3.0-4.3 mm; Kenya, Ethiopia Attagenus insidiosus HALSTEAD, 1981

4 (3) Protibia less elongate, protarsus with 2nd segment about 1/3 as long as 5th; dorsal pubescence unicolorous dark brown or with lighter golden setae pres­ent on pronotum and elytra; female antenna with apical segment less elon­gate; mesosternum quadrate.

5 (6) Apical segment of antenna (of both sexes) smaller, obviously narrower than previous segment, very small in male (Figs 22-23); pronotum with lighter setae laterally (at lateroposterior angles only or covering larger area), a few to many scattered along basal margin and sometimes forming inconspicous patches at each side near base of disc; elytra may have a few lighter setae near humeral angles and scutellum; length 3.0-3.6 mm; Ghana, Congo

Attagenus nigripennis A R R O W , 1915

6 (5) Apical segment of antenna larger, about as broad as previous segment (Fig. 24); dorsal pubescence unicolorous dark brown to black; length 3.3-3.5 mm; Congo Attagenus atricolorPiC, 1931

7 (2) Lateral regions of prosternum slightly raised (appearing broad) not forming knife-like carinae; dorsal pubescence black or dark brown; antenna: Figs 27-28; cuticle unicolorous black to dark brown; cosmopolitan

Attagenus unicolor unicolor (BRAHM, 1791)

8(1) Trochantin exposed; lateral regions of prosternum narrow, not forming knife-like carina.

9(10) Apical segment of male antenna very elongate (Fig. 25), that of female small and less elongate (Fig. 26); dorsal cuticle bicoloured: head, pronotum and

scutellum black or very dark brown, elytra unicolorous reddish or yellowish brown; Afrotropical and Palaearctic regions

Attagenus smirnovi ZHANTIEV, 1973

10 (9) Apical segment of male antenna less elongate (Fig. 17), that of female also less elongate (Fig. 18); dorsal cuticle bicoloured: head and pronotum black or very dark brown, scutellum and elytra unicolorous dark brown; Ghana

Attagenus endroedyi sp. n.

Attagenus globosus sp. n. (Figs 29-30)

Type material - Holotype (female): "Panama, Cordillera de San Bias, El Liano env., 1978" (JHAC).

Description ofholotype -Female. Body (Fig. 29) small, oval, convex. Cuticle bicoloured; head and pronotum red-brown, elytra light brown. Body length 2.9 mm, width 1.8 mm. Head finely punctate with long yellow pubescence. Palpi entirely yellow; pubescence on mentum denser. Ocellus on front presented. Eye large, strongly convex. Antennae yellow, 11-segmented, antennái club 3-segmented, apical antennái segment oblong (Fig. 30). Pronotum finely punctate as head, with long yellow pubescence. Scutellum triangular, with short yellow pubescence. Elytra finely punctate; elytra light brown, without bands of pubescence; pubescence entirely yellow. Epipleuron in anterior part broad with dense punctures and terminating gradually before posterior margin of metepisternum. Legs light brown with yellow pubescence; anterior tibiae with short red-brown thorns. Ventral sur­faces with long yellow pubescence. Lateral regions of prosternum strongly raised (appearing narrow) and forming knife-like carinae in front of coxae. Abdominal ventrites with long yellow pubescence. Male unknown.

Etymology - Latin globosus (= round).

Remarks - Seven species of the genus Attagenus are known from South America. Attagenus globosus sp. n. is similar to the two introduced species, A. unicolor (BRAHM, 1791) and A. cyphonoides REITTER, 1881:

1 (4) Lateral regions of prosternum strongly raised (appearing narrow) and form­ing knife-like carinae in front of coxae; apical segment of male antenna ( i f known) elongate.

2 (3) Antenna of female with elongate apical segment; antenna of male with api­cal segment about five times as long as the previous two combined; body elongate; cuticle yellow-brown to almost black

Attagenus cyphonoides REITTER, 1881

3 (2) Antenna of female with oblong apical segment; (male antenna unknown); body oval; cuticle bicoloured: head and pronotum red-brown, elytra light brown Attagenus globosus sp. n.

4 (1 ) Lateral regions of prosternum slightly raised (appearing broad), not forming knife-like carinae; apical segment of male antenna less elongate; cuticle black to reddish brown, unicolorous to bicolored (pronotum may be slightly to obviously darker than elytra). Attagenus unicolor ( B R A H M , 1791)

Orphinus (Orphinus) apicebrunneus sp. n. (Figs 31-33)

Type material - Holotype (male): "Ghana, Ashanti region, Kwadaso, 320 m, N 6.42 W 1.39, Dr. S. Endrődy-Younga Igt."; "Nr. 324, mixed light, 1 l.iii.1969". Paratype (male): same data as holotype. Holotype is deposited in HNHM, paratype in JHAC.

Figs 29-36. 29-30. Attagenus globosus sp. n.: 29 = body outline, 30 = antenna of female. 31-33. Orphinus (O.) apicebrunneus sp. n.: 31 = dorsal view (dotted area indicating brown cuticle), 32 = an­tenna of male, 33 = male genitalia. 34-35. Phradonoma endroedyi sp. n.: 34 = body outline, 35 = an­

tenna of male, 36 = male genitalia

Description - Male. Body black on dorsal and ventral surfaces; generally small and ovate (Fig. 31). Body length 1.8 mm, width 1.0 mm. Head finely punctate with short black pubescence. Palpi brown; pubescence on mentum denser. Frontal median ocellus present. Antennae yellow, 11 -segmented, antennái club 3-segmented, terminal antennái segment small, circular (Fig. 32). Pronotum punctate as head on disc, densely foveolate, with short black pubescence on posterior part. Scutellum triangular with short black pubescence. Elytra densely foveolate; cuticle black with black pubescence, apical part brown, with black pubescence. Legs brown with yellow pubescence. Mesoventrite coarsely punctate on lateral parts, elsewhere finely punctate. Abdominal ventrites with short light brown pubescence. Male genitalia: Fig. 33. Female unknown.

Etymology - Latin apice-brunneus, referring to the brown apical part of elytra.

Remarks - The new species is very similar to Orphinus (Orphinus) africanus PIC, 1927. Differential diagnosis of both species is in the following key:

1 (2) Body ovate, cuticle of elytra black, with apex brown, with black pubescence throughout; length 1.7-1.8 mm; Ghana

Orphinus (Orphinus) apicebrunneus sp. n.

2( 1 ) Body elongate, cuticle of elytra black, with apex red, covered by yellow pu­

bescence; each elytron wi th a broad transverse band with yellowish pubes­cence; length 2.5-2.8 mm; Congo, Haut-Niger.

Orphinus (Orphinus) africanus PIC, 1927

Phradonoma endroedyi sp. n. (Figs 34-36)

Type material - Holotype (male): "Ghana, North. Reg. Nakpanduri, 75 km S von Bawku, Dr. Endrődy-Younga Igt."; "No. 13, lichtfalle, 11.vii. 1970". Holotype is deposited in HNHM.

Description - Male. Body oval (Fig. 34), bicolorous. Body length 2.1 mm, width 1.0 mm. Head coarsely punctate with erect yellow pubescence Maxillary palpi brown. Eyes large with microsetae. Antennae brown, 11 -segmented, with 3-segmenled club (Fig. 35). Antenna! fossa broad. Frontal me­dian ocellus present. Pronotum black, finely punctate with long, yellow erect pubescence, lateral margins not visible from above. Scutellum black, triangular, without pubescence. Cuticle of elytra brown (except black humeri and anterior part near scutellum), without fasciae, covered with yellow erect pubescence only. Each elytron with minute bump on humeri; anterior part densely foveolate, other parts finely punctate. Prosternum and metaventrite with long yellow pubescence. Legs brown, anterior tibia with black spines along shaft. Abdominal ventrites finely punctate with long yellow pu­bescence. Male genitalia: Fig. 36. Female unknown.

Etymology - The species is dedicated to the late Dr. S. Endrődy-Younga.

Remarks - The genus Phradonoma J ACQUELIN DU V A L , 1859 includes 7 spe­cies from the Afrotropical region. The new species is very similar to P. uninotata Pic, 1942 from "East Africa" (elytra black with red spot on the disc), but differs from it in the much smaller size of body, antennae and male genitalia.

Anthrenus (Anthrenus) cf. senegalensis PIC, 1927

Material examined - "Ghana, Western region, Elmina, 20 m, N 5.05 W 1.20, Dr. S. Endrődy-Younga Igt."; "Nr.291, beaten, 31.xii.1967", J. H Á V A det. (1 female, HNHM).

Remarks - This female specimen agrees in all characters with Anthrenus (Anthrenus) senegalensis Pic, 1927, but differs in the colour pattern of scales on elytra. The status of this specimen remains doubtful until a more extensive material can be studied.

Phradonoma cf. endroedyi sp. n.

Material examined - "Ghana, Northern region, Yapei, 50 km SW of Tamale, Dr. Endrődy-Younga Igt."; "Nr. 409, beaten, 15.xii.1969", J. H Á V A det. (1 female, JHAC).

Remarks - This female specimen is similar to P. endroedyi sp. n., but differs in the elongate form of body (length 2.5 mm). The cuticle of elytra is bicolorous, i . e. anterior half black, postrior half red, covered by yellow short pubescence. The sta­tus of this specimen remains doubtful until a more extensive material can be studied.

Acknowledgements - Thanks are due to all the collectors and curators mentioned in the Intro­duction for loaning and/or providing me with the Dermestidae material.

REFERENCES

CASTELNAU, F. L . N. C. DE LAPORTE, (1840): Histoire naturelle des Insectes Coléoptères. Tome deuxième. P. Duménil, Paris, 564 pp.

DELOBEL, A. & T R A N , M . (1993): Les Coléoptères des denrées alimentaires entreposées dans les régions chaudes. Faune tropicale X X X I I . ORSTOM/CTA, Paris, 425 pp.

ESCALERA, M . M. DE LA, (1914): Los coleopteros de Marruecos. - Trahajos del Museo National de Ciencias Naturales (Madrid), Zoológia 11: 1-553.

H A L S T E A D , D. G . H . (1981): Taxonomic notes on some Attagenus spp. associated with stored prod­ucts. Including a new black species from Africa (Coleoptera: Dermestidae). - Journal of Stored Products Research 17: 91-99.

H Á V A , J. (2002): Five new species of Dermestidae (Coleoptera), with notes on synonymy and distri­bution of the family. - Acta Musei Moraviae, Scientiae hiologicae 87: 29^10.

K A L Í K , V . (1955): Dermestidae deraethiopischen Region aus der Sammlungen des Königl. Museum von Belgisch Kongo in Tervuren. (Mit Bemerkungen über verwandte Arten). -Revue Zoologie Botanique Africaines 52: 294-322.

K O C H E R , L. (1956): Catalogue commenté des Coléoptères du Maroc. Fase. IV - Clavicornes et groupes voisins (Dermestidae pp. 19-30). - Travaux de L'Institut Scientifique Chérijien 11: 1-136.

M A D E R , L. (1954): Bemerkungen und Richtigstellungen zu einigen paläarktischen Coleopteren. -Entomologische Nachrichten 1: 48-53.

M R O C Z K O W S K I , M. (1968): Distribution of the Dermestidae (Coleoptera) of the World with a cata­logue of all known species. - Annales Zoologici 26: 15-191.

M R O C Z K O W S K I , M . (1975): Dermestidae, Skórnikovate (Insecta: Coleoptera). Fauna Polski, Tom 4. - Polska Akadémia Nauk, Institut Zoologii, Warszawa, 162 pp.

P I C , M . (1950): Coléoptères nouveaux du Congo de diverses familles. - Revue Zoologie Botanique Africaines 43: 356-362.

Z H A N T I E V , R. D. (1973): Ergebnisse der zoologischen Forschungen von Dr. Z. Kaszab in der Mon­golei. Nr. 321. Dermestidae (Coleoptera). - Annales historico-naturales Musei nationalis hungarici 65: 187-194.

Diaria itinerum Pauli Kitaibelii III.

1805-1817 Edited by László Lőkös

Paul Kitaibel, the renown Hungarian botanist completed 17 longer and several shorter journeys in the 18-19th century during which he wrote a very detailed travel diary. Leaving Pest or Buda, from the boundary of the town he kept a scientifically very thorough itinary making it as detailed as possible. His observations covered the entire geological and natural geographical aspects of the land­scape, laying special emphasis on rocks, minerals, soil, running and still waters. He specifically examined mineral waters, the first analy­ses of which were made on the spot w i th his travelling laboratory kit. The most painstaking and thorough observations were made on the plants. Wi th profound thoughtfulness he made his notes also on the animals he saw. Besides the known plants he reported the discovery of numerous new plant species in his diary. He complemented the species list wi th phytogeographical, ecological, cenological observa­tions still valid today, wel l preceding the scientific character of his age. He jotted down everything he experienced.

His small note-books of 12-16 pages each bui ld up his diary, which is a valuable storehouse of information on the economy, agri­culture, industrial production, mining, trade and traffic of the day. Ethnography d id not fall far from his interest either, so he wrote colourful reports on wedding customs, popular life and rural feats.

Kitaibel's travel diary was written in German and Latin. The text is written wi th the orthography used by Kitaibel, and the same holds also for the species names. The illegible, consequently un-printed parts are indicated by a row of dots (...), while the uncertain readings of some words are (?) question marked.

The journeys are also shown on clear maps to show the com­pleted routes. The book closes wi th 55 pages of register containing in­dex of taxa and index of geographical names.

Published in 2001. Hard bound 460 pages. Price: 40 Euros plus p. and p.

ISBN 963 7093 729

Orders should be sent to: The Library, Hungarian Natural History Museum

H-1088 Budapest, Baross u. 13, Hungary Fax: (36-1) 3171669