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    Relocation of the 1936 Mojokerto skulldiscovery site near Perning, East Java

    O.F. Huffman a,*, Y. Zaim b, J. Kappelman a, D.R. Ruez Jr.c,J. de Vos d, Y. Rizal b, F. Aziz e, C. Hertler f

    a Department of Anthropology, The University of Texas at Austin, Austin, TX 78712-0254, USAb Department of Geology, Institut Teknologi Bandung, Jalan Ganesa 10, Bandung 40132, Indonesia

    c Department of Geological Sciences, The University of Texas at Austin, Austin, TX 78712-0254, USAd Naturalis, the National Museum of Natural History, P.O. Box 9517, 2300 RA Leiden, The Netherlands

    e Geological Research and Development Centre, Jalan Diponegoro 57, Bandung 40122, IndonesiafJ. W. Goethe University, Zoological Institute, Vertebrate Paleobiology, Siesmayerstr. 70, D-60054 Frankfurt, Germany

    Received 22 November 2004; accepted 3 November 2005

    Abstract

    The fossil calvaria known as the Mojokerto childs skull was discovered in 1936, but uncertainties have persisted about its paleoenvironmen-

    tal context and geological age because of difficulties in relocating the discovery site. Past relocation efforts were hindered by inaccuracies in old

    base maps, intensive post-1930s agricultural terracing, and new tree and brush growth. Fortunately geologic cross sections and site photographs

    from 1936-1938dnot fully utilized in past relocation fieldworkdclosely circumscribe site geography and geology. These documents match the

    conditions at just one sandstone outcrop. It is situated on the southern margin of a topographic nose at the upper end of a w18 m-wide gully

    (w0663760 m E, 9183430 m N, UTM Zone 49M), w15 m southeast of the Kumai et al. (1985) relocation. The relocated discovery bed

    is w3.3 m of fossiliferous pebbly sandstone, a river-channel deposit cut into tuffaceous mudstone. The sandstone and mudstone beds correspondto original site descriptions. Pebbly sandstone is also found within the skull.

    The calvaria is well-preserved and taphonomically similar to large and fragile specimens found among several hundred vertebrate fossils

    excavated from the sandstone in 2001-2002. Since no well-preserved fossils were found intact at the surface of the sandstone, the good condition

    of the Mojokerto skull suggests that it was buried fully when discovered. The relocated hominin bed is the uppermost fluvial sandstone of a ma-

    rine-deltaic sequence in the upper Pucangan Formation. The Mojokerto child probably died along the ancient seacoast, judging from the large

    extent of the deltaic facies and evidence that the calvaria experienced minimal transport. The relocated discovery bed is w20 m stratigraphically

    above the horizon from which the widely cited 1.81 0.04 Ma 40Ar/39Ar date for the skull (Swisher et al., 1994, Science 263, 1118) was

    obtained. Additional field and laboratory results will be required to determine the skulls age.

    2005 Elsevier Ltd. All rights reserved.

    Keywords: Homo erectus; Homo modjokertensis; Pithecanthropus; Indonesia.

    * Corresponding author. P.O. Box 548, Bastrop, TX 78602, USA. Tel.: 1 512 303 9501.

    E-mail addresses: [email protected] (O.F. Huffman), [email protected] (Y. Zaim), [email protected] (J. Kappelman), ruez@mail.

    utexas.edu (D.R. Ruez), [email protected] (J. de Vos), [email protected] (Y. Rizal), [email protected] (F. Aziz), [email protected]

    (C. Hertler).

    0047-2484/$ - see front matter 2005 Elsevier Ltd. All rights reserved.

    doi:10.1016/j.jhevol.2005.11.002

    Journal of Human Evolution 50 (2006) 431e451

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    Introduction

    The fossil calvaria found in 1936 near Perning, East Java,

    and known as the Mojokerto childs skull (Figs. 1 & 2) appears

    to indicate that Homo occupied a marine-coast habitat in the

    Plio-Pleistocene as early as w1.8 Ma. Such an age would

    make the Mojokerto one of the oldest non-African homininfossils discovered. The marine-coast setting is quite unlike

    habitats known for mid-Pleistocene and older hominin popula-

    tions outside Java.

    Efforts to confirm the geological age of the skull and ex-

    plore the implications of its paleoenvironment have been frus-

    trated in recent years by uncertainty over the bedrock origin of

    the fossil and field location of the discovery bed. The history

    of the find has been evaluated in detail recently (Huffman

    et al., 2005), reaffirming an in situ provenience for the homi-

    nin fossil. Here we document the field relocation of the

    discovery site and bed, and discuss the importance of the relo-

    cation for dating the skull and linking it to a seacoast habitat.

    Andoyo, a geological assistant with the Geological Survey

    of the Netherlands Indies (Survey), found the Mojokerto skull

    at a hillside exposure of the Pucangan Formation while search-

    ing systematically for vertebrate fossils (Huffman et al., 2005).

    Survey geologist Johan Duyfjes, who had mapped the areageologically, examined the site and emphatically asserted

    that the skull had been excavated from bedrock at a depth of

    1 m. Survey paleontologist G.H. Ralph von Koenigswald iden-

    tified the specimen as an early hominin. Quaternary geologist

    Helmut de Terra and archaeologist Hallam L. Movius visited

    the site with von Koenigswald in 1938 and concurred with

    Duyfjes geological assessment of the skulls context (de Terra

    et al., 1938).

    By WWII, scientists generally accepted the paleontological

    determination and considered the Mojokerto skull to be among

    Jetis

    m52

    50m

    m0

    m57

    0663000 m E

    0005819

    Nm0003819

    0001819

    0006819

    0004819

    00

    02819

    0665000

    0664000 m E

    MOJOKERTO

    PERNING

    Sumbersuko

    Sumbertengu

    Sambikerep

    Klagen

    m05

    m05

    C

    Sumberglagah

    m52

    Topographic

    Contour

    (CI=12.5 m)

    Road

    Trail

    Village

    Figure 2A

    Mojokerto skull

    discovery area

    (Figure 2B)

    UTM Grid

    (Zone 49M)

    0 1 km

    0663000 m

    B Java Sea

    1000M-

    Skull Site

    Pacific

    Ocean

    Indian Ocean

    EASTERN JAVA

    B

    A

    Fig. 1. Index (A, B) and topographic (C) maps of the Mojokerto skull discovery site near Perning, East Java, Indonesia (C is taken from the 1:25,000 digital to-

    pographic maps ofBakosurtanal, 1998, 1999).von Koenigswald (1936a,b)named a new species,Homo modjokertensis, on the basis of the Mojokerto childs skull

    (Huffman et al., 2005).Jacob (1973, 1975a,b; see alsoIndriati, 2004) designated the fossil Perning 1 (or I) and Modjokerto 1.Storm (1994)and Anton (1997)

    attributed it to Homo erectus. The specimen, also known as the Mojokerto child, has been variously characterized as a braincase, calvaria, cranium, skull and

    skullcap.

    432 O.F. Huffman et al. / Journal of Human Evolution 50 (2006) 431e451

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    the oldest hominin fossils known, based on publications that

    described the discovery location and geology (Duyfjes,

    1936, 1938b; von Koenigswald, 1936a,b, 1940; de Terra,

    1943). Regrettably, the Survey left no permanent marker at

    the discovery site, Duyfjes died during World War II, and

    von Koenigswald left Java permanently in 1946. As a conse-

    quence of these and other events, confusion arose over the

    geographic and stratigraphic position of the find.

    Interest in the Mojokerto skull surged following the 1994

    publication of a 1.81 0.04 Ma 40Ar/39Ar date on horn-

    blende from samples of the Pucangan Formation collected

    in the discovery area (Swisher et al., 1994, 2000). However,

    questions remain about this date, with the field identification

    of the discovery site and the relationship of the hominin bed

    to the dated rock being in doubt. Uncertainties such as these

    have clouded paleoanthropological interpretation of the

    Mojokerto fossil for years (de Vos, 1985, 1994; Semah,

    1986; Theunissen et al., 1990; Hyodo et al., 1993; Walker

    and Shipman, 1996; Lewin, 1998; Klein, 1999; Wolpoff,

    1999; Langbroek and Roebroeks, 2000; Swisher et al.,

    2000; Shipman, 2001; Huffman, 2001a; Klein and Edgar,

    2002; Morwood et al., 2003).

    Regardless of the exact age of the Mojokerto skull, its

    paleoenvironmental setting will remain important to under-

    standing early human evolution. Java continues to be the ear-

    liest maritime terrain known to have been inhabited by early

    hominins, and marine-coast adaptations potentially played

    a significant, yet little-understood role in the early hominin

    lineage. The ancient landscape in the Homo erectus district

    of eastern Java is notable for its variety of seacoasts, as well

    as volcanic mountains, non-volcanic uplands, river valleys,

    and lake margins (Huffman, 1997, 1999).

    Moreover, during periods of low Plio-Pleistocene sea level

    when the Java Sea and other parts of the Sunda Shelf were par-

    tially exposed, eastern Java was on the southern rim of a broad

    terrain inferred to have contained coastal lowlands, lakes,

    B

    A B

    Fig. 2. Aerial photograph of the Mojokerto skull discovery area (B) and vicinity (A) with topographic contours from Fig. 1C. The discovery site is known from

    1936 maps (Andoyo, 1936; Duyfjes, 1936) to be located east of the tributary of Kali Klagen (Klagen Creek) that is shown in (B). The attributes of the discovery siteand bed, as indicated in 1936-1938 descriptions and documents (Figs. 3A &5-7), match the geographical and geological conditions at only one locality, our

    Mojokerto skull site relocation. The scene-center lines of three 1936-1938 photographs ( Figs. 5-7), shown on (A), cross at the relocated site (see Figs. 8-10

    and text for details). This point is w15 m southeast of the relocation ofKumai et al. (1985). The documentation for the discovery site does not fit the landscape

    and geology elsewhere in the discovery area (Table 1; a location referred to as Jacob location 1 is in the broad field [Fig. 2B], and Jacob location 2 is at a cliff face

    [Fig. 3B]). Strata of the Pucangan Formation form the only bedrock in the discovery area (Figs. 3 & 4). PRQ (A) is a quarry on the Perning-Sumbertengu road

    where the Monument Sandstone and underlying deltaic topset- and foreset-beds are exceptionally well exposed (Fig. 4; see photograph in Huffman and Zaim,

    2003). The aerial photograph was taken in October 1996; original prints were at w1:11,500.

    433O.F. Huffman et al. / Journal of Human Evolution 50 (2006) 431e451

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    discovery area (Fig. 2B), leaving little possibility that the shal-

    low discovery pit of the Mojokerto skull could have survived

    the last 60 years. Second, topographic contour lines on the

    1936 maps are too stylized to accurately reflect the landscape

    details indicated in the old documents. Third, no geographic

    co-ordinates, such as latitude and longitude, were included

    on Duyfjes map, nor were any coordinates reported for theMojokerto skull site during the 1930s. Fourth, several reloca-

    tion approaches were employed in the past, and their contra-

    dictory results are hard to evaluate due to incomplete

    documentation of the relocation efforts.

    In 1969 Teuku Jacob made the first reported attempt to relo-

    cate the point where the Mojokerto skull was found. Apparently

    without revealing the exact reasoning behind his choice, Jacob

    identified the discovery site as a meter-deep rectangular pit

    measuring four feet by eight feet . in the middle of a barren

    field. He went again to Perning in 1975, this time with Andoyo,

    who pointed out a different location for the find (Hyodo et al.,

    1993; Walker and Shipman, 1996; Swisher et al., 2000:42-43,

    reported the date of the Andoyo visit as 1969). It has recentlybecome clear that by 1990 Andoyos recollection of 1936 events

    was in error (Huffman et al., 2005). His memory of the location

    of the site may also have been unreliable in 1975.

    Indonesian and Japanese geologists reexamined the Pucan-

    gan Formation in the general discovery area in 1976-1978. Not

    knowing of Jacobs work (Hyodo et al., 1993), they selected

    another location and concluded:

    Although we were unable to determine exactly the site and

    horizon of theP. [Pithecanthropus] modjoketensis find, the

    locality which we assumed and showed in . [a sketch

    map] coincides with the locality drawn in Duyfjes

    (1936)geological map. The stratum at this locality contains

    many vertebrate fossils and resembles the lithofacies of

    the P. modjokertensis-bearing bed described by Duyfjes.

    (Kumai et al., 1985:61).

    Their locality, which we call the Kumai location, is in an ag-

    ricultural field on a topographic nose where they collected ver-

    tebrate fossils on the surface in September 1978 (Fig. 2B);

    team leader N. Watanabe, now deceased, selected the location

    based on information provided by local villagers, together

    with the distribution of vertebrate fossils in the discovery

    area, as determined by fieldwork (H. Kumai, pers. comm.,

    2000, 2002).

    In 1992, Jacob showed Garniss Curtis and Carl Swisher a 2-

    3 m high cliff-face outcrop of conglomeratic sandstone below

    a recently constructed monument commemorating the Mojo-

    kerto discovery (Swisher et al., 2000:42-43; see also Hyodo

    et al., 1993:180). Jacob indicated that the cliff was where An-

    doyo said the skull was found. The spot is w60 m from the

    location that Jacob identified as the discovery site in 1969.

    We call the bedrock exposed in this cliff and field the Mon-

    ument Sandstone (Huffman and Zaim, 2003), and refer to the

    1969 and 1992 localities as Jacob location 1 and Jacob loca-

    tion 2, respectively. Jacob 2 is w110 m south of the Kumai

    location. We do not know the exact position of Jacob location

    1, only that it must sit between his location 2 and the Kumai

    Fig. 4. Stratigraphy of the Mojokerto skull discovery sequence. DB is the re-

    located discovery bed; MS, the Monument Sandstone; MM II, Mollusk Mem-

    ber II; MM III, Mollusk Member III. MM II and III contain shallow-marine

    mollusks and are regional mapping marker units in the Pucangan Formation

    of Duyfjes (1938a,b). The figure is based upon detailed measured sections

    made by Djuhaeni, R.T. Buffler, O.F. Huffman and F.P. Wesselingh in 2001-

    2002 (Fig. 2A shows the section traverses; Huffman and Zaim, 2003). The

    sandstone and conglomerate is volcaniclastic and frequently cross bedded;

    most of the mudstone appears to be tuffaceous. The sequence between MM

    II and MM III comprises (from bottom to top): (a) interbedded delta-front

    sandstone and mudstone exhibiting bottomset, foreset and topset bedding (as

    shown diagrammatically in the lithofacies column); (b) deltaic sandstone

    and conglomerate with cross beds, MS; (c) lower delta flood-plain mudstone

    and distributary-channel sandstone, including DB and an overlying paleosol

    mudstone. DB is the uppermost fluvial sandstone in the delta-plain sequence,and is capped by an unconformity having at least local importance (noted by

    the heavier line in lithofacies column of the figure;Huffman and Zaim, 2003).

    Measured sections of this sequence also have been published by Duyfjes

    (1936), Sartono et al. (1981), Kumai et al. (1985), Semah (1986), Hyodo

    et al. (1992, 1993) and Morwood et al. (2003). Hyodo et al. (1992, 1993,

    2002) assigned MM II to the Olduvai subchron. For this and their other

    GPTS assignment to be correct, however, the beds below the Monument Sand-

    stone (MS) would have to have accumulated at an average depositional rate 1/

    70th of that in strata above this level; that is, the w35-m thick top-MM II to

    base-MS interval would equate to w0.7 Ma, or a 0.005 cm/yr average deposi-

    tional rate; the base-MS to base-MM III, w35 m thick, would represent

    w0.01 Ma and 0.35 cm/yr; the superjacent portions of the Pucangan Forma-

    tion and the Kabuh Formation, roughly 300 m according to Duyfjes

    (1938a,b), would comprise the post-Jaramillo period ofw0.99 Ma and equate

    to a 0.30 cm/yr average rate of deposition (see also Morwood et al., 2003).

    435O.F. Huffman et al. / Journal of Human Evolution 50 (2006) 431e451

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    location. The relationship of the three locations is most clearly

    illustrated inFig. 3B.

    Jacob himself has not published details about his relocation

    efforts in the years since he took Andoyo to Perning. Concern

    about the relocation issue led Susan Anton, Garniss Curtis,

    Carl Swisher and Agus Suprijo (Jacobs assistant) to attempt

    to pinpoint the field location of the discovery from several1930s photographs. They did not find the site, but believed

    that it must be within a few meters, or perhaps a few tens

    of meters, distant from the Mojokerto monument (Swisher

    et al., 2000:98). This is near the Monument Sandstone outcrop

    where in 1992 they collected the rock sample used for dating

    purposes (Fig. 3B).

    Our initial fieldwork in 1999 and 2000 indicated that the

    stratigraphic difference between beds at the Kumai and Jacob

    2 locations was of potential significance in assessing the

    meaning of the published radioisotopic date for the skull

    (Huffman, 2001a). About the same time, Morwood et al.

    (2003)concluded that the find site was near Jacob 2, but did

    not publish the reasoning behind their choice. A more compre-hensive relocation effort was needed.

    Relocation of the discovery site and bed

    In 2001 and 2002, we relocated the discovery outcrop of the

    Mojokerto skull, and found numerous in situvertebrate fossils

    in the relocated hominin bed (Huffman et al., 2002). We also

    mapped the area around the relocated site with a Total Data

    Station, measured detailed stratigraphic sections through the

    discovery sequence, and determined the stratigraphic level of

    the relocated discovery bed (Figs.2B,3B,4 &8-10;Huffmanand Zaim, 2003). To achieve a reliable relocation, we com-

    pared old maps to new ones, and then far more importantly,

    matched 1930s photographs and descriptions to conditions

    on the ground.

    1936 & recent maps

    We first sought to constrain the relocation effort by compar-

    ing 1936 maps to recent maps and aerial photographs, which

    had not been available to previous investigators. The approach

    seemed promising, because Duyfjes (1936) published loca-

    tion is the same as the one that Andoyo (1936) inscribed on

    a 1:25,000 topographic map five days after the discovery,

    and Andoyo had apparently surveyed the point with a transit

    (Huffman et al., 2005). Insurmountable problems arose, how-

    ever, when we overlaid 1936 maps on a composite of the new

    digital topographic maps and high-resolution aerial photo-

    graphs (Figs. 1C &2A).

    There is a considerable difference in topographic contour-

    ing on the old and new maps. A tributary of Klagen Creek

    (Kali Klagen), which lies west of the discovery site (Figs.

    1C & 2B), is positioned too far to the east on the 1936

    maps. The geographic placement of roads and road intersec-

    tions also is off by 20-50 m at some points. These inaccuracies

    can be detected because an Army Map Service (1942/1943)

    chart of the area is available. It has road tracks and contouring

    similar to Duyfjes and Andoyos maps, and corresponds more

    closely to the aerial photograph than do the maps used by the

    discoverers. Drafting errors evidently were introduced when

    Survey personnel prepared the hand-drawn base maps on

    which Andoyo plotted the discovery location. The best result

    that our comparison of old and new maps achieved was a gen-eral match of hills, valleys, creeks and roads.

    Based upon this match, the discovery point taken from

    Duyfjes map is repositioned on the present-day landscape at

    a small ridge lying between the Kumai and the Jacob 2 loca-

    tions (Fig. 2B). This point is 80-100 m from the locations.

    When either location was forced to fit the point on the 1936

    maps, geographic features did not line up on the overlay. How-

    ever, the geographical accuracy of the point on Andoyos

    (hence Duyfjes) 1936 map cannot be verified because An-

    doyos survey notes have not been located. The starting place

    of his traverse on the day of the discovery may have been at

    one of the areas inaccurately drawn on the base map. Andoyo

    also might have made mistakes while surveying. Therefore,the map overlay exercise does not exclude or favor any loca-

    tion within the discovery area.

    1930s documentation

    The inconclusive results from the use of map overlays led

    us to compare information from other 1930s site documents

    with the landscape and geology in the discovery area. Fortu-

    nately the old documents allow for a precise reconstruction

    of the geography and geology around the site.

    The documentation indicates that the discovery was made

    at an outcrop of conglomeratic tuffaceous sandstone in theupper Pucangan Formation; the strata at the site dipped

    w10 along the northern flank of the Kedungwaru Anticline

    (Andoyo, 1936; Duyfjes, 1936, 1938a,b; von Koenigswald,

    1936a,b; de Terra, 1943; Movius, 1944:82, termed the litho-

    logy breccia). Terrace deposits were absent, and soil devel-

    opment was limited, so that near-surface deposits did not

    obscure the bedrock geology at the site (Fig. 3A; de Terra

    et al., 1938; de Terra, 1943; Movius, 1944; this situation

    has been confirmed by Kumai et al., 1985; Huffman, 2001a;

    Huffman and Zaim, 2003).

    Fragmentary vertebrate fossils occurred at the surface of

    the site; their presence is what prompted Andoyo to dig there

    in February 1936 (Duyfjes, 1936). Although in situ fossils

    were not abundant in the discovery sandstone (von Koenigs-

    wald, 1936b), Andoyo evidently found the skull after digging

    only 1 m deep into the bedrock (Huffman et al., 2005). His ex-

    cavation notched into the bedrock, and was no more than

    w2 m north-south and w7 m east-west, even after additional

    digging took place during April 1936 (Figs. 3A,5A & C,6A

    &7; see also North View 1 in Huffman et al., 2005, figure 10).

    The skull was found in the basal 0.5 m of the sandstone bed,

    which has a total stratigraphic thickness of at least 4 m on

    Duyfjes site cross section. The dip drawn on the cross section

    is, however, w14, not the w10 stated in written accounts.

    The bed therefore might be somewhat thinner than 4 m.

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    This site is north of the small ridge and close to, but not exactly

    at, the Kumai location.Figure 8illustrates the topographic set-

    ting north of the small ridge. The Kumai location sits

    atop a topographic nose. A flat-bottomed gully lies to the south

    of the nose, and a broad valley is to its northwest along the trib-

    utary of Kali Klagen. The nose merges eastward into a slope

    that rises northward into a large ridge. This is the main east-

    west ridge in the discovery area. In general form, the nose,

    gully, valley and ridge are identical to the principal features sur-

    rounding the site, as determined from 1930s records.

    The gully terminates eastward at a terrace wall (Fig. 6A).

    Approximately 18-m wide by w5-m deep, the gully is only

    B

    1

    D

    3

    3

    2

    1

    TREES &

    BUSHES

    (partial)

    BANANA PLANTAT MOJOKERTO

    SKULL SITE

    YOKE &

    BASKETS

    VON

    KOENIGSWALD

    2 GULLY

    1 NOSE

    RIDGE3

    A

    C

    Perning-Sumbertengu Road

    Fig. 5. Photographs taken on April 19, 1938, when H. de Terra, H.L. Movius, and G.H.R. von Koenigswald visited the discovery site. The main image (A, C), not

    published previously, is referred to as Southwest View because it looks southwestward across the site. The inset photograph (B, D) shows von Koenigswald near

    a pit in which a banana plant grew. de Terra (1940, 1943) indicated that photographs similar to (B) showed the Mojokerto skull discovery site ( Huffman et al.,

    2005). All known early photographs of the site show the same pit and banana plant (Figs. 5-7; see also,Huffman et al., 2005, figures 3 & 10). The Southwest View

    demonstrates that the skull site was located on a topographic nose (labelled #1 on C). The nose was north of the gully (labelled #2) that is seen on Duyfjes cross

    section (Fig. 3A) and in North View 2 (Fig. 6A). Persons standing near the site in the 1930s could look westward past the nose to the tributary to Kali Klagen and

    beyond to the ridge (labelled #3 on C) and Perning-Sumbertengu road. An image similar to (B) was published by Swisher et al. (2000)as an illustration of the use

    of old photographs in their efforts to relocate the site. The photographs presented in this figure, apparently filed with the papers of H. de Terra and H.L. Movius

    since 1938, are archived in Peabody Museum of Archaeology and Ethnography, Movius papers (PMAE), and are copyrighted by the President and Fellows of

    Harvard College (A, image #N34853 PMAE, and B, image #N39442 PMAE; photographers unidentified; published with permission).

    438 O.F. Huffman et al. / Journal of Human Evolution 50 (2006) 431e451

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    w2 m wider and shallower than the topographic profile on

    Duyfjes cross section (Fig. 3). The situation therefore fits

    the requirement based upon the old documents that the dis-

    covery site is north of the upper reaches of a small gully. Be-

    yond the terrace wall to the east, the agricultural fields

    widen, again matching conditions in an old site photograph

    (Fig. 7). The gully-nose combination no longer exists east

    of the wall, and thus the site could not have been located

    there (Fig. 8).

    The outcrop geology at the upper end of the gully also cor-

    responds to the old site documents, particularly Duyfjes cross

    section (Fig. 3A). Sandstone makes up the entire south wall

    and most of the north wall of the gully (Fig. 3B). A persistent

    light-colored tuffaceous mudstone bed is exposed in the north

    wall between the sandstone and the agricultural soil of the

    nose (Fig. 9). The mudstone averages w2.2 m thick and varies

    in thickness from w1 to 3 m. It is overlain by several meters

    of pebbly (volcaniclastic) sandstone.

    This sandstone is our candidate for the Mojokerto discovery

    bed. In addition to cropping out north and northeast of the

    upper end of the gully, the sandstone underlies the agricultural

    soil atop the nose (Fig. 9). The soil zone is whereKumai et al.

    (1985)made their surface collection of vertebrate fossils (H.

    Kumai, pers. comm., 2000).

    B

    B

    A

    Fig. 6. (A) North View 2, a 1936-1938 photograph of the site, and (B) an enlarged portion of the scene highlighting the banana plant and discovery pit (the top of

    which is marked by a gray line). The photographer (identity unknown) stood on the south wall of a gully, foreground of (A), looking northward towards the site.

    This alignment and location are the same as are depicted in Duyfjes site cross section (Fig. 3A). Based on the estimated height of the man, the pit measured w2 m

    north-south andw7 m east-west (w7 m3 with the volume approximated as a right-triangular prism). Sandstone formed the back walls of the pit, judging from old

    descriptions of the site and the appearance of the outcrop (erosion-resistant rock ledges are sandstone in the area today). A thin stratum of sandstone may have

    underlain the pit floor, but the lighter tone of the ground south of the pit in the photograph suggests that tuffaceous mudstone cropped out here. Therefore, only the

    basal w1 m of the sandstone bed was exposed in the pit, and the contact between the hominid-bearing sandstone and the mudstone was at or slightly below the pit

    floor (compare toFig. 3A). The hill slope that lay north of the pit rose steadily to form an east-west ridge (A). The gully floor (lower left and center of A) ended

    eastward at a terrace wall crowned with small plants (noted by inverted triangles). A field lies farther east on the upside of the wall. This field is seen more fully in

    the Northwest View (Fig. 7). The size of the banana plant in the pit suggests that its corm was planted 6-7 months before the photograph was taken; the banana

    leaves show wilting indicative of dry season conditions (S.R. Gowen, pers. comm., 2004; R.L. Swennen, pers. comm., 2004). The photograph thus appears to have

    been taken in the summer of 1936 or 1937 during an otherwise undocumented site visit. The identity of the standing man is uncertain. His appearance suggests he

    might be Andoyo (seeHuffman et al., 2005, figure 3A). North-View 2 is in the von Koenigswald archive, Research Institute Senckenberg, Frankfurt (published

    with permission), and apparently has been filed with von Koenigswalds papers since the 1930s. The photograph was published previously bySwisher et al. (2000).

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    The relocated discovery sandstone is a fluvial deposit. It has

    a broad channelized base, prominent cross-bed sets and other

    bedding indications of mid-channel river bars, including pre-

    served dune surfaces (mega-ripples) on the bars. During

    2001-2002 we opened the lower 2 m of the sandstone for ex-

    cavation, and recovered several hundred vertebrate fossils.

    These include terrestrial mammals, crocodile, turtle, fish and

    freshwater mollusks (Table 2; Huffman and Zaim, 2003;

    Zaim et al., 2003). The upper part of the sandstone was not

    exposed in our excavations, but must occur between nearby

    outcrops of fine-grained sandstone and mudstone. The full

    thickness of the sandstone is w3.3 m using a 7.5 dip

    (see below).

    The relocated discovery bed is overlain by 4-5 m of mud-

    stone on which a paleosol had developed. This paleosol mud-

    stone is unconformably overlain by sandstone and mudstone

    with burrows and marine mollusks. These beds are the basal

    strata of a marine unit capped by Mollusk Member III, as map-

    ped byDuyfjes (1934, 1936, 1938b). Our relocated discovery

    bed is therefore the uppermost fluvial sandstone of the coarsely

    clastic, rapidly prograding deltaic sequence lying between

    Mollusk Members II and III (Figs. 3B &4).

    Kumai et al. (1985:59) present a stratigraphic sequence at

    our relocation site that is similar to the one that we observed.

    They apparently attributed the Mojokerto discovery site to

    a 1.3 m medium- to coarse-grained pebbly sand bed, overlain

    by 1.5 m of medium-grained sand with granules (Hyodo et al.,

    1993, referred these sands as SG2). They found 1.1 m of silt

    below their 2.8 m sand (equivalent to our w3.3 m relocated

    discovery sandstone). They report 3.5 m of silt above the sand-

    stone. These two silt beds correspond to our tuffaceous mud-

    stone and paleosol mudstone, respectively.

    Determining the structural dip (and hence stratigraphic

    thickness) is problematic in the discovery area. The sedimen-

    tary succession consists of deltaic deposits with fore-set bed-

    ding, large-scale cross-bed sets, and cut-and-fill features that

    confound point observations of structural attitude (Fig. 4).

    We employ a 7.5 average structural dip. This is based on

    B

    A

    B

    C R O P P E D

    Fig. 7. Northwest View (A), taken on April 19, 1938, with the portion of the scene around the Mojokerto skull site enlarged (B). The photographer stood at the

    south edge of a field looking northwest toward the nose and discovery pit ( Fig. 10B highlights these features). The broad valley with the tributary of Kali Klagen

    was visible in the distance, as was the ridge with the Perning-Sumbertengu road. The gully seen in North View 2 ( Fig. 6A) sat to the west (left) of the photo-

    graphers location. The man in the pit, H. de Terra or possibly H.L. Movius, was at approximately the same spot that the men were positioned in North View1 (Huffman et al., 2005, figure 10) and North View 2 (Fig. 6B). This spot may have been known in the late 1930s to be the exact point of discovery of the Mo-

    jokerto skull. The terrace wall visible to the east of the pit (right in A) was probably an outcrop of the discovery sandstone. This outcrop, as modified by post-1930s

    terracing, was the location of our principal 2001-2002 excavation (Fig. 8). The photograph, which apparently remained in possession of H. de Terra and H.L.

    Movius, was filed with the Movius papers of Peabody Museum of Archaeology and Ethnography and is copyrighted by the President and Fellows of Harvard

    College (image #N34852 PMAE; photographer unidentified; published with permission).

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    numerous individual strike and dip measurements taken when

    measuring stratigraphic sections. Duyfjes report of a steeper

    dip at the site (10-14 , see above) was probably due to his hav-

    ing less stratigraphic control in this complex sedimentary sit-

    uation. We note, for example, that the closest strike and dip

    values (7-15 ) on his 1934 geologic map were recorded along

    the Perning-Sumbertengu road. This is w500 m from the dis-covery site and >30 m stratigraphically below the relocated

    hominin bed. The strata along the road preserve the foreset

    bedding with sedimentary and structural dip as high as 27 .

    Given the intensive agricultural terracing in the area, it is

    not surprising that the 1936-1938 excavation pit no longer ex-

    ists, and that the present-day micro topography differs in small

    ways from the conditions shown in Duyfjes site cross section

    and the old photographs. These differences are reconciled if

    we assume that, since the 1930s, farmers flattened the nose,

    cut back the gully walls a bit, raised the gully floor with

    w2 m of fill, rebuilt the gully-field transition, and terraced

    the hillside northeast of the site.

    Micro-topographic changes such are these are certainlywithin the farmers earth-moving capabilities. For example,

    the local men hired for our excavation crew readily dug

    back the sandstone bedrock with their normal agricultural

    tools and transported the debris to nearby fields in buckets.

    Moreover the changes that we infer to have occurred around

    the site are within the range of modification evident many pla-

    ces in the Perning area, as the little-exploited 1930s landscape

    was developed into the intensely farmed terrain of today.

    Because thegeology andgeographyimmediately north of the

    upper end of the gullyare,withthe exceptions noted, to the same

    as conditions documented for the discovery site in 1936-1938,

    we conclude that our relocation site and the sandstone that weexcavated are the Mojokerto skull site and the discovery bed.

    1936-38 & 2001 photographs

    In order to test the validity of our field relocation efforts and

    fix the position of the discovery site as precisely as possible, we

    overlaid the old site photographs on images made in 2001. The

    new photographs were taken from vantage points that were as

    close as possible to those used in 1936-1938. The overlays

    match well (Figs. 9 & 10). We also plotted the vantage points

    and scene-center lines of the old photographs onto an aerial pho-

    tograph and a map. This more comprehensive form of compar-

    ison also verifies the relocation (Figs.2A &8).

    Overlays were made for the 1936-1938 photographs that we

    call North View 2 and Northwest View. The new north view

    photograph (Fig. 9) was taken from a spot high on the south

    gully wall. From this vantage point, the gully, gully-field tran-

    sition, nose, and main ridge to the north are visible in the same

    relative positions as seen in North View 2. A new northwest

    view photograph was taken using similar criteria (Fig. 10).

    We are more confident in relocating the point from which

    the North View 2 photograph was taken than the location at

    which Northwest View was photographed.

    The overlays for North View 2 and Northwest View were

    made independently of one another. Each overlay was judged

    to be correct when the ridge lines, gully and field boundaries,

    and other topographic features on the old images corresponded

    to those seen in the modern photographs (Huffman et al.,

    2002). Once this result was achieved, the 1936-1938 discovery

    pit was transferred to the new north view and northwest view

    images (Figs. 9 & 10). In both cases the pit transfers to the

    field relocation of the discovery site. The transferred point issomewhat lower in elevation and farther to the south on the

    modern north view than on the northwest view. The topo-

    graphic features used to align the new and old photographs

    begin to mismatch when the old images are shifted laterally

    on the modern images in an amount that is equivalent to

    only a few meters on the ground.

    It was not possible to complete a Southwest View overlay

    comparison. Numerous trees now line the north edge of the

    gully and obscure the view beyond the nose when looking

    southwestward across the relocation from the point used for

    Southwest View in 1938 (Figs. 5A &8). It is also exception-

    ally difficult to stand near this point because the ground has

    been cut away by agricultural terracing.A map-view comparison of the 1936-1938 photographs and

    the modern landscape provides an even more comprehensive

    test of our relocation effort. Because trees do not hamper the

    map-view approach, we were able to include the Southwest

    View along with the two other views. This method makes

    use of the fact that the discovery pit lies at the approximate

    center of the three old photographs. In each case, the site falls

    along a map line (the scene-center line of the photograph) that

    extends from the vantage point through the pit and across the

    terrain beyond. The intersection of the three scene-center

    lines, once they are correctly positioned on the map, precisely

    defines the map location of the discovery site.To achieve this result, we constrained the vantage points of

    the North View 2, Northwest View and Southwest View to the

    small map areas that fieldwork indicated were acceptable. We

    then forced the scene-center lines to intersect at the relocation

    north of the upper end of the gully, but not necessarily at the

    best field estimate of the relocation. We limited the pit to mod-

    ern locations where the pit floor would lie at or slightly below

    the projected contact between the tuffaceous mudstone and re-

    located hominin sandstone (Fig. 6). We allowed this position

    to fall above the present-day land surface, where necessary,

    because of the likelihood that the rock around the Mojokerto

    skull was dug away completely in 1936 and the pit destroyed

    by farming activities since 1938. We assumed that the place

    where the man stood in the North View 2 photograph was

    the actual discovery point of the calvaria (Fig. 6).

    The final positioning of the relocation site and vantage points

    produces a close correspondence between the features seen in

    the old photographs and the topographic elements of the modern

    terrain, both as represented on the map and aerial photograph

    (Figs. 2 & 8) and determined by fieldwork. For example, the

    Southwest View scene-center line aligns along the nose

    (Fig. 8), and intersects the ridge with the Perning-Sumbertengu

    road where it rises to the south (Figs. 2A, 5A & C). Additionally,

    the intersection of the scene-center line of the Northwest View

    and the road ridge closely approximates that found in the old

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    photograph. Even the distinctive shape and high point of the

    ridge line seen in 1938 are recognizable today.

    The intersection of the scene-center lines in Fig. 8

    matches closely our best field estimate of the point where

    the man stood in North View 2. The comparison of 1936-

    1938 and 2001 photographs therefore substantiates our field

    relocation.

    This point is w3 m south of the nearest place in our exca-

    vations that we dug into the relocated discovery bed (w6 m

    south of a concrete pillar inscribed ITB August 2002 that

    we placed at the west end of our long excavation face at

    the terrace wall; Fig. 8). We made a TDS measurement at

    best field-estimate of the relocation, and also took 943 GPS

    (Garmin eTrex Vista set for WGS 84) readings at a single

    station in the TDS grid. The average of the readings

    and TDS measurements combine to give UTM (Zone 49M)

    co-ordinates for the relocated site of 0663760 m E and

    9183430 m N.

    Discussion

    Bed & matrix lithology

    The validity of our relocation might be tested further by

    comparing the lithology and fossil taphonomy of the relocated

    discovery bed to the matrix in the Mojokerto specimen and its

    state of bone preservation. The lithological comparison is

    addressed first.

    In 1936, von Koenigswald (1936b:1001dtranslated)

    described the matrix as greenish, slightly conglomeratic tuff-

    aceous sandstone. The greenish color presumably indicates

    that the rock was little weathered, because weathering and

    soil formation in the discovery area turns rock to brownish

    and reddish hues. Good quality photographs of the specimen

    taken in 1936 reveal fill that is consolidated, medium- to

    coarse-grained, and apparently pebbly (Huffman and Zaim,

    2003; Huffman et al., 2005).

    GULLY

    FIELD

    SMALL RIDGE

    MAIN RIDGE

    NOSE

    BROAD

    VALLEY

    WEIVWS 5.gi

    F

    WEIV

    WN

    01&7

    .giF

    WEI

    VN

    9&6

    .giF

    STEEP SLOPES

    WITH TREES

    & BRUSH

    OLD PHOTOGRAPH

    VANTAGE POINT &

    SCENE CENTERLINE

    DUYFJES 36/38B

    CROSS SECTION

    (FIGURE 3A)

    DUYFJES

    1936 MAP

    LOCATION(APPROX.)

    KUMAI ET AL.

    1985 HOMINIDSITE

    mN

    TERRACED

    FIELDS IN

    VALLEYS

    TERRACED

    FIELDS ON

    HILL SLOPES

    WATER

    COURSE

    2001-2002

    EXCAVATIONS

    MOJOKERTO

    SKULL SITE

    RELOCATION

    0 20 40M

    RELOCATIONS

    Tr

    ibu

    t

    ar

    yof

    ne

    gal

    Ki

    la

    K

    Fig. 8. Map of the area surrounding the relocated Mojokerto skull-discovery site (based on Total Data Station measurements and an aerial photograph, Fig. 2B).

    The relocated hominin site is on the hill slope north of the upper reaches of the gully. The site lies at the intersection of the scene-center lines of three 1936-1938

    photographs that were taken from vantage points east, south and southwest of the site ( Figs. 5-7). The location proposed byKumai et al. (1985)was positioned on

    this map using the outline of the gully and nose that are shown on their sketch map.

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    Duyfjes (1936, 1938bdtranslated) characterized the dis-

    covery bed as conglomeratic sandstone composed of sand

    and gravel of volcanic origin (volcaniclastic). Before visiting

    the Mojokerto discovery field site, de Terra (1938;

    1943:443) studied the mineral composition of the fill withinthe skull and rock cleaned from it; after the fieldwork in

    the laboratory in Bandoeng, [de Terra] compared the material

    from the pit and found it to be identical with the matrix. No

    more detailed data from his comparison have been located.

    The sandstone in the skull was covered with paint after de

    Terras examination, thus limiting direct observation and de-

    scription. During a hands-on examination of the fossil in

    1992, however, Swisher scraped away some of the paint and

    found a light-colored pumice pebble in the matrix ( Swisher

    et al., 2000:48, 87, photograph). Jacob later gave Swisher

    and colleagues a fragment of the pumice pebble. It played

    a key role in their dating studies (Swisher et al., 1994, 2000).

    Images of the matrix produced from computed tomography

    (CT) scanning have recently become available (Coqueugniot

    et al., 2004; Balzeau et al., 2005). Balzeau et al. (2005)find

    that the matrix is heterogeneous. It contains a coarse layer

    in the antero-superior portion of the cranial cavity, and much

    finer-grained material elsewhere. Based upon images they

    published, the coarser layer is very-coarse-grained sandstone

    with granules and pebbles.

    Our excavations reveal substantial vertical and horizontal

    grain-size variations in the relocated discovery bed. Medium-

    to very-coarse-grained, conglomeratic sandstone predominated.

    Granule-pebble conglomerate and fine-grained sandstone were

    present in lesser volumes. Only one thin lens of mudstone and

    very-fine-grained sandstone was encountered within the portion

    of the bed that we exposed. The material immediately surround-

    ing our excavated fossils varied from coarse-grained sandstone

    to pebble conglomerate. Only one fossil was encased in rock as

    fine as medium-grained sandstone.The bed is gray when fresh, and reddish where affected by

    iron-oxide staining. The sandstone consists of subangular to

    subrounded volcanic-rock fragments, feldspar, quartz and

    mafic minerals. Well-rounded, light-colored pebbles and small

    cobbles of pumice with pyroxene and hornblende phenocrysts

    are a prominent gravel component (Huffman and Zaim, 2003).

    Most of the gravel consists of rounded, darker and denser vol-

    canic rock types, what Duyfjes (1938b) characterized as an-

    desite. The largest pumice clasts we found were 5-10 cm in

    greatest dimension. This is several times larger than the typical

    darker and denser volcanic pebbles.

    The pebbly coarse-grained fill in the skull appears to be

    closely similar, if not identical, to the typical lithology of

    the relocated hominin bed. The skull matrix is consistent in

    this way with the lithology of the bed. However, the Monu-

    ment Sandstone and some other units in the sequence contain

    pebbly coarse-grained sandstone also. The pebbly matrix

    therefore does not tie the skull to the relocated discovery

    bed unequivocally. The statement of Swisher et al.

    (1994:1119) that only one stratum in the discovery area (our

    Monument Sandstone) contains pumice and volcanic ma-

    trix similar to the infilling of the Mojokerto skull is at odds

    with our field observations.

    Although the fine matrix in the skull is finer-grained than

    the bulk of the relocated hominin bed, the difference does

    Kumai Locality Discovery SiteFrom North View 2

    Agricultural Fill

    WATER COURSE

    (hidden)

    ESRU

    OC

    RETA

    W

    FillMudstone

    Sandstone

    Field in Gully

    Gully North Wall

    Fig. 9. North view of the relocated Mojokerto skull site taken in 2001 from a point high on the gully south wall (Fig. 8). The scene shows the sandstone and

    ruffaceous mudstone exposed in the gully north wall. Even after removing brush, visibility was obscured far more than was the case in the 1930s (compare to

    Fig. 6A). To minimize the interference of trees, this photograph was taken from a point somewhat lower on the gully wall and farther west than was the case

    in 1936-1938 when North View 2 was made (Fig. 6A). The new and old images were overlaid by aligning gully boundaries (north and east edges) and the crest

    line of the main ridge north of the site. The 1936-1938 pit, man and banana plant were then transferred to this image. The old pit lies at our relocated site position.

    Tuffaceous mudstone crops out just below the feet of the man taken from the 1936-1938 photograph. The mudstone is overlain by the vertebrate-bearing sandstone

    that we identify as the Mojokerto skull bed. The sandstone, which cannot be seen in this photograph, crops out immediately north of the relocated 1936-1938 pit,

    underlies the field at theKumai et al. (1985) relocation site, and is well exposed in a nearby terrace wall where we excavated it in 2001-2002 ( Figs. 7 & 8).

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    not eliminate this stratum as the source of the Mojokerto hom-

    inin fossil. The bed where we excavated it might have con-

    tained less fine-grained sandstone than it did at the point of

    discovery, located several meters away. The calvaria also

    might have lain on a river bottom in a position that affected

    the sediment size coming into the cranial cavity. The fill might

    have been finer (or coarser) than the surrounding matrix. Fur-thermore, the grain size of the river sediment might have

    changed over time in ways preserved within the calvaria but

    not fully represented by the deposit. The matrix-bed compar-

    ison therefore is of limited value as a test for our relocation

    because of insufficient lithological information on the fill

    and other uncertainties.

    Mojokerto skull & 2001-2002 fossils

    It would be valuable to compare the state of preservation of

    the hominin specimen and the vertebrate remains found with it

    in 1936 to the fossils that we excavated from the relocated

    hominin bed in 2001-2002. Regrettably, the 1936 non-hominin

    fossils appear to have been lost, and von Koenigswald did not

    publish a comparison of the fossilized condition of the childs

    skull versus that of the associated fossil remains (Huffman

    et al., 2005). Furthermore, we were not permitted to examine

    the original Mojokerto skull.Our comparison of the preservational states therefore was

    limited to assessing the condition of our excavated fossils

    from direct observation and the skull from casts, photographs,

    drawings and recently published analyses (Storm, 1994; An-

    ton, 1997; Huffman and Zaim, 2003; Coqueugniot et al.,

    2004; Balzeau et al., 2005; Krovitz and Shipman, in press).

    This comparison is nonetheless valuable. In addition to help-

    ing to evaluate our relocation results, it provides a basis for

    testing the conclusion that skull was buried in sandstone

    when discovered and for proposing that the hominin died on

    the Mojokerto coastal plain.

    B

    B

    RIDGEI G

    NOSEOS

    FIELDI L

    Site

    April 1938

    August 2001

    A

    Perning-Sumbertengu Road

    Fig. 10. (A) Northwest view of the relocated site taken in 2001; (B) an annotated 1938 Northwest View (Fig. 7) for comparison. The photographs look northwest-

    ward across the nose and broad valley to the distant ridge where the Perning-Sumbertengu road is located. Trees presently obscure the valley and ridge from points

    east of the site, so (A) was taken from the place nearest the 1938 vantage point that gave a good view of the valley and ridge. A wide-angle lens was used to produce

    an image that encompassed the entire 1938 scene (B), the outline of which is indicated on (A). The field in the foreground is the same one seen in the lower right of

    North View photographs (Figs.6A &9). The 1938 and modern photographs were aligned by matching the upper boundary lines of the field (green), nose (yellow),

    and ridge (blue) on the two photographs. The lines from the 1938 photograph are shown on (A). The pit (with man and banana plant), transferred as part of this

    process, indicates the relocated Mojokerto discovery site.

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    evidence indicates that he unearthed the juvenile hominin

    calvaria from sandstone having a modest density of vertebrate

    remains. And his discovery excavation may have been less

    volumetrically than 1 m3 and was probably less than w7 m3

    in volume (Fig. 6).

    Second, because fragile fossils degrade severely when they

    weather out of the relocated discovery sandstone, a specimenas well preserved as the hominin calvaria was unlikely to have

    been exposed to any significant surface weathering before it

    was found by Andoyo.

    Only fragmentary fossil remains are mentioned in reports

    from the 1930s as having been seen at the surface in the area

    around the discovery site; there was no report of surface homi-

    nin material (Huffman et al., 2005). We found isolated teeth and

    small fragments of long bone and antler loose on the surface

    near the relocated site. Similar fossils were surface collected

    byKumai et al. (1985), including cervid and bovid teeth, mam-

    mal bone fragments, and a turtle shell (Aimi and Aziz, 1985).

    However, since the surface specimens found in the last 30 years

    are at least as likely to be exposed by agricultural activity as bynatural weathering of bedrock, they do not help us to determine

    how fossils weather out of the relocated hominin bed.

    We found only one fossil at the surface embedded in the

    sandstone prior to excavation. The specimen was a highly

    weathered and fragmented turtle carapace (Huffman and

    Zaim, 2003). Some of the larger excavated fossils, such as

    the antler and radius, showed evidence of natural in situbreak-

    age. This fracturing would help explain severe fragmentation

    of fossils upon exposure to weathering at the surface.

    Noin situvertebrate fossils were observed at the surface of

    other sandstone beds in the discovery area. Moreover, few fos-

    sils were seen embedded in natural or terrace outcrops of sand-stone anywhere in the portion of the Perning district that we

    surveyed. None of these fossils was as fragile and well pre-

    served as the hominin skull.

    Given conditions in the Perning district, it is difficult to

    imagine circumstances that would have permitted the Mojo-

    kerto specimen to erode out of sandstone bedrock with the

    calvaria largely intact and filled with sandstone. The well-

    preserved condition of the Mojokerto calvaria therefore argues

    strongly for an in situ discovery of a buried specimen, just as

    the discoverers stated unequivocally was the case.

    Finally, given the taphonomic evidence at hand, we con-

    clude that the Mojokerto child is likely to have died within

    the ancient Mojokerto Delta. Fluvial transport of the calvaria

    was apparently limited. Sedimentary facies relationships in

    the Pucangan Formation indicate that the Mojokerto Delta

    was at least 10 km across; the river feeding the Delta originat-

    ed in highlands many tens of kilometers upstream (Duyfjes,

    1938a,b; Huffman, 2001a,b; Huffman and Zaim, 2003). It

    seems likely therefore that the child lived in and died on the

    ancient delta plain near Perning, where its remains were sub-

    jected to defleshing and disarticulation.

    Less likely is the possibility that the childs body was carried

    intact by flood waters from a long distance upstream, exposed to

    weathering in the Delta, and then reincorporated into the river

    briefly before burial in the relocated discovery bed. Long-

    distance transportation of carcasses, even live animals, is noted

    in the historical record of Java as having occurred during lahar-

    related floods (volcanic mudflows; Carthaus, 1911:27-28; de

    Terra, 1943:449, in part quoting Franz Junghuhn). However,

    the terrestrial-vertebrate assemblage in the relocated hominin

    bed lacks the taphonomic hallmarks of a mass-kill event and

    provides no specific support for long-distance transport bylahars or other floods. Lahar breccias do occur in the

    Pucangan Formation of the Mojokerto area, but they are found

    west of Perning and stratigraphically above and below the level

    of the hominin stratum (Duyfjes, 1934, 1938a,b).

    Implications for the age of the skull

    The widely cited 40Ar/39Ar date for the Mojokerto skull,

    1.81 0.04 Ma (Swisher et al., 1994, 2000), was obtained

    from material collected w20 m stratigraphically below the re-

    located hominin bed (Figs. 3 & 4). The hominin fossil is there-

    fore w1.8 Ma or younger. At least five other sources of

    ambiguity surround the age of the skull, which consequentlycannot be determined accurately at this time.

    First, Morwood et al. (2003) conclude that the skull

    is less than w1.49 Ma, having obtained 1.49 0.13 and

    1.43 0.15 Ma fission-track dates on pumice-clast zircons

    from the Monument Sandstone and relocated hominin bed,

    respectively (their Pumice Horizon 5 sample is from the

    Perning road quarry, and Pumice Horizon 6 from our relo-

    cated discovery site, judging from their location descriptions;

    Fig. 2).

    Second, the age determinations of bothSwisher et al. (1994)

    andMorwood et al. (2003)are subject to the questionable but

    common assumption in studies of the hominin-bearing forma-tions of Java that the dated pumice was erupted shortly before

    deposition of the sampledbed. The risks in this line of reasoning

    are highlighted by recent radioisotopic dates from a single lahar

    unit at Sangiran Dome.Bettis et al. (2004)obtained 40Ar/39Ar

    plateau ages spanning nearly a million years on hornblende sep-

    arates from various pumice clasts collected from this lahar. The

    study leaves no doubt that substantial supporting evidence is re-

    quired to justify accepting as the age of deposition a date from

    a pumice clast or a set of clasts. Reworked pumice is possible

    whether it is in a lahar deposit or a conglomerate that is less

    clearly related to a volcanic mudflow.

    Swisher et al. (1994, 2000) dated hornblende from pumice

    gravel and a sandy matrix found in what appeared to be a tuff

    layer of the Monument Sandstone (Fig. 3). However, the evi-

    dence they provide is not sufficient to accept the layer as a pri-

    mary volcanic deposit and the 1.81 0.04 Ma date as the

    depositional age (de Vos, 1994; de Vos and Sondaar, 1994;

    Swisher, 1994). Even if active volcanism was involved, there

    are other sources of ambiguity.Swisher et al. (1994)apparent-

    ly combined clasts which were not necessarily the same age to

    obtain hornblende from the pumice, and did not have a means

    of eliminating epiclastic hornblende from the matrix sample.

    The mixing of hornblende could produce an average40Ar/39Ar date that is significantly older than the youngest vol-

    canic product in the bed.

    446 O.F. Huffman et al. / Journal of Human Evolution 50 (2006) 431e451

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    Morwood et al. (2003)also combined pumice clasts in their

    dating samples and had concerns over reworked material. The

    clasts from the relocated-hominin bed were selected from grav-

    el composed of diverse rock types. The Monument Sandstone

    sample, on the other hand, was collected from a layer contain-

    ing virtually all-pumice gravel of various clast sizes. This situ-

    ation improves the chance that the pumice was introduced

    directly into the river system as lapilli and bombs by volcanic

    eruption (see alsoHuffman and Zaim, 2003). Even in the case

    of the Monument Sandstone sample, however, the fission-track

    analyses of individual grains give widely scattered results, and

    mixed-eruption ages are potentially involved.

    Therefore, while the abundance of fresh labile volcaniclas-

    tic detritus in the Perning section argues for active volcanism

    (Huffman, 2001;Huffman and Zaim, 2003), the radioisotopic

    determinations published so far do not necessarily date the

    deposition of the sedimentary sequence.

    Third, available magnetostratigraphic studies (Semah,

    1986; Hyodo et al., 1992, 1993) do not uniquely constrain

    the age of the relocated hominin bed. The data are difficult

    to interpret because of scattered results in the hominin-bearing

    part of the section. Moreover, the paleomagnetic sampling did

    not cover the entire Pucangan sequence exposed in the Perning

    district. This significantly limits the opportunity to tie the sec-tion to the Geomagnetic Polarity Time Scale (GPTS). There

    also are no complimentary studies elsewhere in the Pucangan

    and Kabuh outcrop belt of the greater Mojokerto area that are

    available to substantiate the Perning determinations.

    Semah (1986:386) obtained very scattered results strati-

    graphically above Monument Sandstone, continuing as high as

    the base of a zone of normal polarity overlying the Pucangan

    Formation in the Kabuh Formation. His readings below the

    Monument Sandstone were predominantly reversed polarities,

    which leads him to conclude that the lower part of the Perning

    section is older than the Brunhes Chron (>0.78 Ma).

    Hyodo et al. (1992, 1993) also obtained predominantly

    intermediate polarities together with some normal polarities inthe Monument Sandstone and in the section as high

    stratigraphically as the base of Mollusk Member III (Fig. 3).

    These workers assign the interval to the Jaramillo subchron

    (0.99-1.07 Ma). They conclude that the Olduvai subchron

    (1.77-1.95 Ma) is in Mollusk Member II, and that intermediate

    polaritiesbetween Mollusk Member II and the Monument Sand-

    stone correlate to Hyodos Sangiran excursion in the section at

    Sangiran Dome (w180 km to the west). However, their assign-

    ments of the Perning sequence to the GPTS imply a dramatic and

    unexplained change in the rate of deposition or significant un-

    recognized stratigraphic gaps within the section (Fig. 4). Their

    correlation of the marine-nonmarine transition at Perning toa similartransition at Sangiran Domedused to support their dat-

    ing of the Mojokerto skulldfails to take into account strati-

    graphic and facies variations known to occur in the

    intervening Pucangan outcrop belt (Duyfjes, 1936, 1938a,b).

    Neither Semah (1986) nor Hyodo (1998, 1999, 2002)

    appear to have identified unequivocally reversed polarities in

    or above the Monument Sandstone. Swisher et al. (1994; Fig-

    ure 3) found that two clay layers in the Monument Sandstone

    have normal remanent magnetism. If the radioisotopic material

    they dated was reworked from significantly older bedrock,

    the normal polarity interval could represent the older portion

    of the Brunhes chron (

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    Horizons II and III (Fig. 4). The vertical sequence of lithofacies

    is consistent with an unbroken progression of depositional envi-

    ronments from delta slope to flood plain, and the localized pro-

    gradation of a delta lobe into a shallow water embayment

    (Huffman and Zaim, 2003; see also:Morwood et al., 2003).

    Such a lobe might have been active for only a small fraction

    of the time represented by the entire lateral and vertical accumu-lation of the Pucangan Formation between Mollusk Horizons II

    and III, a stratigraphic interval of deltaic sedimentation that

    Duyfjes (1938a,b)traced along many tens of kilometers of out-

    crop in the greater Mojokerto area. The present-day delta of the

    Brantas River (Porong delta lobe), which liesw50 km southeast

    of Perning and is roughly the size of the ancient Perning lobe,

    prograded more than 5 km since 1880 (Hoekstra, 1989). This

    high rate of deposition underscores the possibility that the Pern-

    ing delta lobe represents100 m

    of exposed Pucangan strata and is overlain by w300 m of Pu-

    cangan and Kabuh formations (Duyfjes, 1934, 1938a,b). Thesebeds are folded and unconformably overlain by many tens of

    meters of flat lying Brantas-River fill near Mojokerto (Duyfjes,

    1935). Geologists working over many decades have consid-

    ered the Pucungan beds to be mid-Pleistocene or older on

    the basis of these geological relations.

    The percent of modern mollusk species in Mollusk Mem-

    bers II and III was once used to date the strata in which the

    Mojokerto skull was found (Cosijn, 1931, 1932; van Es,

    1931;Martin, 1932; Duyfjes, 1936). von Koenigswald (1934,

    1936a)recognized a fossil mammalian fauna in the upper Pu-

    cangan near Perningdthe Jetis fauna. He believed that the

    fauna indicated an early Pleistocene age for the Mojokerto

    skull. However, the exact stratigraphic positions of only

    a few of the critical fossils (Duyfjes, 1938b) were reported.

    Most of them apparently were surface finds. Also the upper

    Pucangan in the area includes strata above and below the

    hominin-bearing horizon. This complicates determining the

    stratigraphic relationship of the Mojokerto skull to the re-

    ported species. And the Jetis assemblage may be much youn-

    ger Pleistocene than von Koenigswald thought it was ( de Vos

    et al., 1982, 1994; de Vos, 1985, 1994). The vertebrate fossils

    known from the Kabuh Formation in the Perning district

    (Duyfjes, 1938a,b) are not age diagnostic, and therefore do

    not help in establishing a minimum age for the Mojokerto

    skull.

    The fossils excavated from the relocated hominin bed

    (Table 2) include four taxadHexaprotodon sivalensis, Axis

    lydekkeri, Rusa sp., and Duboisia santengdthat have been

    used to distinguish various extinct mammalian faunas in Java

    (de Vos et al., 1982, 1994). The assemblage in the relocated

    hominin bed clearly does not correlate with the oldest and youn-

    gest of these faunas (Satir, Punungand WajakFaunas).The largecervidRusa is not present in the Trinil Fauna from thePithecan-

    thropus erectusbed at Trinil and the hominin-bearing Bapang

    Formation of Sangiran Dome. The excavated material therefore

    appears to represent oneof three faunas that are older or younger

    than the Trinil Fauna: the Ci Saat Fauna (older), which occurs in

    the Sangiran Formation of Sangiran Dome and was defined on

    the basis of West Java localities; the Kedung Brubus Fauna

    (younger), which is recognized on the basis of sites near Kedung

    Brubus village located along the Pucangan-Kabuh outcrop belt

    between Perning and Trinil, and also is known from the Bapang

    Formationof Sangiran Dome; or the Ngandong Fauna (younger)

    found in the hominin-bearing Solo River terrace deposit at

    Ngandong, north of Trinil.Sartono et al. (1981)and Zaim (1981) list planktic forami-

    nifera from an unspecified site(s) in the clay facies of the

    Pucangan Formation. This facies underlies the hominin-

    bearing sequence, and forms a lateral facies equivalent to it

    north and east of the Perning district (Duyfjes, 1934,

    1938a,b). Neither the original sample(s) nor illustrations of

    specimens are available, further complicating age interpretation

    of the assemblage. Several of the listed species are valuable

    biostratigraphically in the Plio-Pleistocene time frame (e.g.,

    Globigerinoides jistulosus, Globorotalia tosaensis and Globi-

    gerinoides extremus; Gradstein et al., 2004; R.M. Leckie,

    pers. communication, 2005). Marine microfossils may ultimatelyhelp to determine the age of the skull, but do not do so now.

    In summary, additional field and analytical resultsare needed

    to date the Mojokerto fossil more exactly than latest Pliocene or

    early-mid Pleistocene in age. The w0.3 Ma difference between

    the 40Ar/39Ar and fission-track age determinations must be re-

    solved. Foranyof these radioisotopic dates to be considered other

    than a maximum age, better evidence must be advanced to show

    that the dated material was erupted shortly before deposition at

    Perning. Additional paleontological and magnetostratigraphic

    control and radioisotopic dating would seem to be required.

    Geochronological conclusions have to be evaluated further in

    terms of the potential for temporal stratigraphic breaks in the

    section, rates of deposition, and the regional stratigraphic

    (including sequence stratigraphic) context.

    Conclusions

    Although intensive agricultural activity over the last 60

    years has changed the appearance of the discovery area, infor-

    mation from maps, reports, and photographs dating from the

    1930s has been used to relocate the discovery site of the Mo-

    jokerto childs skull. Other potential sites, including those

    proposed in the past as probable discovery locations, were

    excluded by comparing the topography and geology observed

    in the field to the same documentary evidence ( Table 1).

    448 O.F. Huffman et al. / Journal of Human Evolution 50 (2006) 431e451

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    The relocated site lies on the south slope of a topographic

    nose situated near the upper end of a small gully and is

    w15 m southeast of the location proposed by Kumai et al.

    (1985). The main topographic features surrounding the relo-

    cated site match those seen in three photographs dating from

    the 1930s (Figs. 5-7). The relocation of the discovery site is

    substantiated further by new site photographs (Figs. 9 & 10),a detailed site map (Fig. 8), and an aerial photograph of the

    area (Fig. 2). The relocated site has UTM (Zone 49M) co-

    ordinates of 0663760 m E and 9183430 m N.

    The relocated discovery horizon is in the lower portion of

    a w3.3 m thick, vertebrate-bearing conglomeratic sandstone

    bed. The lithology of the bed is consistent with what is known

    about the matrix within the hominin calvaria. The bed is the

    uppermost fluvial sandstone body in the prograding deltaic se-

    quence lying between Pucangan Formation Mollusk Members

    II and III (Fig. 4). The relocated discovery stratum is w20 m

    stratigraphically above the bed thatSwisher et al. (1994)dated

    by a 40Ar/39Ar method at 1.81 0.04 Ma (Fig. 3B). The date

    is therefore a maximum age for the Mojokerto skull. The datealso conflicts with the fission-track age determinations re-

    ported byMorwood et al. (2003).

    The Mojokerto fossil is remarkably well preserved given

    that it was an immature calvaria deposited with gravelly

    sand in a swift flowing river. Fossils recovered by excavations

    in the relocated discovery sandstone include Panthera tigris,

    Proboscidea,Sus sp., Hexaprotodon sivalensis, Axis lydekkiri,

    Rusasp.,Duboisia santeng, large-bodied Bovidae,Crocodilus

    sp., Gavialis sp., Trionyxsp., Siluridae, and fresh-water mol-

    lusks (Table 2). The Mojokerto childs skull is consistent

    taphonomically with the few relatively large, fragile terrestri-

    al-vertebrate remains in this assemblage.Well-preserved fossils such as the Mojokerto skull were not

    found on the outcropping surfaces of the relocated hominin

    bed or other Pucangan sandstones (and occur very rarely as

    loose surface finds) in the discovery district. These observa-

    tions support the conclusion based upon historical evidence

    (Huffman et al., 2005) that the hominin fossil was encased

    in Pucangan Formation sandstone and protected from surface

    exposure when discovered.

    The good condition of the skull and the large size of the an-

    cient Mojokerto Delta favor the conclusion that the hominin

    died in the deltaic environment in which it was deposited.

    The Mojokerto child therefore provides evidence for a sea-

    coast Homo erectus population in Southeast Asia, and raises

    interest in the role that maritime adaptation might have played

    in the dispersal and paleoecology of early hominins.

    Acknowledgements

    We thank Aart Berkhout, Bernhard W. Seubert and Johan

    Volker for the translating documents; Richard Buffler, Dju-

    haeni and Frank Wesselingh for contributing toFig. 4; Simon

    R. Gowen and Rony L. Swennen for evaluating the age of the

    banana plant in old site photographs; Todd Green and Christo-

    pher Huffman for producing challenging graphics; Dale

    Hudler for Total Data Station training and post-field analysis;

    Hisao Kumai, Michael Morwood and Carl Swisher III for dis-

    cussing their past work in the discovery area; Mark Leckie and

    William Mclntosh for advice on planktic foraminifera and ra-

    dioisotopic dating, respectively; and Pat Shipman for contrib-

    uting observations on the Mojokerto skull taphonomy and

    1930s documents. The manuscript was improved by commentsfrom Richard Buffler, Pat Shipman, Lucy Todd and four anon-

    ymous reviewers. Financial support from the Leakey Founda-

    tion and the National Science Foundations (BCS 0113688) to

    OFH and the German Research Foundation (HE-3593/1-1) to

    CH is gratefully acknowledged.

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