mapping of the marine habitats and species of lamlash bay, arran

152
Scottish Natural Heritage Commissioned Report No. 400 Mapping of the marine habitats and species of Lamlash Bay, Arran

Upload: buihanh

Post on 25-Jan-2017

216 views

Category:

Documents


0 download

TRANSCRIPT

Page 1: Mapping of the marine habitats and species of Lamlash Bay, Arran

Scottish Natural HeritageCommissioned Report No. 400

Mapping of the marine habitats and species of Lamlash Bay, Arran

Page 2: Mapping of the marine habitats and species of Lamlash Bay, Arran

COMM ISS IONED REPORT

Commissioned Report No. 400

Mapping of the marine habitats and species of Lamlash Bay, Arran

For further information on this report please contact:

Chris Leakey

Scottish Natural Heritage

Battleby

Redgorton

PERTH

PH1 3EW

Telephone: 01738 458661

E-mail: [email protected]

This report should be quoted as:

Axelsson, M., Dewey, S., Doran, J. and Plastow, L. (2010). Mapping of the marine habitats and species of Lamlash Bay, Arran. Scottish Natural Heritage Commissioned Report No.400

This report, or any part of it, should not be reproduced without the permission of Scottish Natural Heritage.

This permission will not be withheld unreasonably. The views expressed by the author(s) of this report

should not be taken as the views and policies of Scottish Natural Heritage.

© Scottish Natural Heritage 2010.

Page 3: Mapping of the marine habitats and species of Lamlash Bay, Arran

Mapping of the marine habitats and species of Lamlash Bay, Arran

Commissioned Report No. 400 (iBids no. 1370)

Contractor: Seastar Survey Ltd.

Year of publication: 2010

Background

In September 2008 a No Take Zone (NTZ) was established in Lamlash Bay (Isle of Arran, North Ayrshire) in Scotland following a consultation for a fishery order (under the Inshore Fishing (Scotland) Act 1984). The proposed NTZ was passed by the Scottish Parliament prohibiting all sea fishing within the NTZ. Scottish Natural Heritage has a commitment to contribute to the monitoring of the No Take Zone in Lamlash Bay and the purpose of the current study is to inform the design and initiation of the monitoring of the Lamlash Bay NTZ, which will establish the future success of the management measures in delivering benefits to both the biodiversity of the bay and the scallop fishery in the wider area. The current study involved an acoustic mapping survey with ground-truthing (camera deployments and sediment sampling) to create maps of the habitats and biotopes within the site in a Geographical Information System (GIS).

Main findings

• The ground-truthing of the sidescan sonar data revealed a heterogeneous sediment environment with sand dominating the central and southern Lamlash Bay and mixed coarse sediments dominating the northern Lamlash Bay and North Channel.

• Nine biotopes were recognised during the ground-truthing (using drop-down camera) of the 2008 and 2009 sidescan sonar data: CR.LCR.BrAs.AmenCio, CR.LCR.BrAs.AmenCio.Bri, SS.SMx.CMx.ClloMx(Nem), SS.SMX.CMx.OphMx, SS.SMP.Mrl.Pcal, SS.SMu.CSaMu.(AfilMysAnit), SS.SSa.CMuSa.(AbraAirr), SS.SMP.KSwSS.LsacR.CbPb and SS.SMP.KSwSS.LsacR.Sa.

• All Lamlash Bay NTZ Day grab locations were classified as ‘sand’ according to the Folk sediment classification system (greater than 80%); with fine and very fine sand being the dominant ‘sand’ categories.

• Two further biotopes have tentatively been classified among the NTZ Day grab locations as SS.SSa.CMuSa.(AalbNuc) and SS.SMx.CMx.(MysThyMx).

• Maerl beds, dominated by Phymatolithon calcareum, were identified in the northern Lamlash Bay, North Channel and the Lamlash Outer regions as well as along the northern, eastern and southern coastline of Holy Island.

• Based on comparisons with the few previous studies the sediment environment and the faunal communities are similar overall.

• Recommendations are made regarding future monitoring including repeat drop-down camera (still photography and video) and sediment sampling surveys at 2 year intervals.

COMMISSIONED REPORT

Summary

Page 4: Mapping of the marine habitats and species of Lamlash Bay, Arran

For further information on this project contact:

Chris Leakey, Scottish Natural Heritage, Battleby, Redgorton, Perth, PH1 3EW

Tel: 01738 458661

For further information on the SNH Research & Technical Support Programme contact:

Policy & Advice Directorate Support, SNH, Great Glen House, Leachkin Rd, Inverness, IV3 8NW

Tel: 01463 725000 or [email protected]

Page 5: Mapping of the marine habitats and species of Lamlash Bay, Arran

Acknowledgements

We would thank the captain and crew of the vessel Alba na Mara for their help and positive attitude throughout the survey period in Lamlash Bay in October 2008. We would also like to thank staff at Scottish Natural Heritage for their support and feedback throughout the project.

Page 6: Mapping of the marine habitats and species of Lamlash Bay, Arran

ii

CONTENTS SUMMARY 1 INTRODUCTION 1 1.1 Project background 1 1.2 The environment 3 1.2.1 Geology and the sedimentary environment 3 1.2.2 The physical environment 4 1.2.3 Biological distributions 4

2 METHODOLOGY 5 2.1 The acoustic surveys 5 2.1.1 Horizontal control 8 2.1.2 Vertical control 8 2.1.2.1 Tides 8 2.1.2.2 The 2008 split beam echosounder 8 2.1.2.3 The 2009 dual frequency echosounder 9

2.1.3 Vessel motion reference 9 2.1.4 Sidescan sonar 9 2.1.5 Weather conditions 11 2.1.6 Processing and charting 11

2.2 Drop-down camera survey methodology 11 2.2.1 Drop-down camera system 11 2.2.2 Camera operation and deployment 11 2.2.3 Data handling 14

2.3 Video and photographic analysis 14 2.3.1 Analysis of the photographs and video records 14 2.3.1.1 Semi-quantitative analysis of the video footage 14 2.3.1.2 Quantitative analysis of the stills photographs 15

2.4 Sediment sampling 15 2.5 Data analysis 17 2.6 Survey limitations 18 2.6.1 Survey vessel 18 2.6.2 Obstructions 18 2.6.3 Equipment failure 18

3 RESULTS 18 3.1 Acoustic analysis 18 3.1.1 Target 1 - anchor sinkers 19 3.1.2 Target 2 - anchor drag marks 20 3.1.3 Target 3 - scallop dredge marks 21 3.1.4 Target 4 - exposed bedrock 21 3.1.5 Target 5 - Deacon Rock 22 3.1.6 Target 6 - sediment furrows 23 3.1.7 Target 7 - low backscatter return (sand) 23 3.1.8 Target 8 - mottled high and low backscatter 24 3.1.9 Target 9 - mottled high and low backscatter 24 3.1.10 Target 10 - area of seabed with Maerl 25 3.1.11 Target 11 – high backscatter area with maerl 26 3.1.12 Target 12 – mooring sinkers and chain 26 3.1.13 Target 13 - coarse sediment with macroalgae and maerl 27 3.1.14 Target 14 – mixed seabed environment off Holy Island 28

3.2 Photographic and video analysis – ground-truthing 28 3.2.1 Ground-truthing of the sidescan sonar imagery 28 3.2.1.1 Target 5 - Deacon Rock 30 3.2.1.2 Target 7 - low backscatter return 30 3.2.1.3 Target 8 - mottled high and low backscatter 31

Page 7: Mapping of the marine habitats and species of Lamlash Bay, Arran

iii

3.2.1.4 Target 9 - mottled high and low backscatter 31 3.2.1.5 Target 10 - area of seabed with maerl 32 3.2.1.6 Target 13 – area of seabed with maerl and seaweed 32 3.2.1.7 Target 14 – mixed sediment north-east of Holy Island 33

3.2.2 General descriptions of the biological communities in Lamlash Bay 33 3.2.2.1 Central and southern Lamlash Bay 33 3.2.2.2 Northern Lamlash Bay and North Channel 35 3.2.2.3 Outer Lamlash Bay 39 3.2.2.4 Southern entrance to Lamlash Bay 41 3.2.2.5 Scallops and other noteworthy species 42 3.2.2.6 Maerl beds 45 3.2.2.7 Brittlestar beds 46

3.3 Sediment samples 47 3.3.1 Particle Size Analysis 48 3.3.2 Macrofaunal distributions 49 3.3.2.1 Abundance 50 3.3.2.2 Diversity 51 3.3.2.3 Macrofaunal composition 51

3.3.3 Comparisons between macrofaunal trends and the sediment data 55 3.3.4 Biotope classification of the 2008 Lamlash Bay Day grab samples 56

3.4 The seabed environment in Lamlash Bay 57 4 DISCUSSION 64 4.1 Survey methodology 64 4.2 Survey limitations 64 4.2.1 Survey extent 64 4.2.2 Discrepancies between the sidescan sonar and photographic survey results 65

4.3 Confidence assessment 65 4.4 Comparisons with previous studies 66 4.4.1 Lamlash Bay rocky reefs 66 4.4.2 Lamlash Bay maerl beds 67 4.4.3 Other fauna and flora in Lamlash Bay 67 4.4.4 Biotope distribution in Lamlash Bay 68 4.4.5 Evidence of fishing activity 69

4.5 Recommendations for future monitoring at Lamlash Bay 69 4.5.1 Requirement for additional survey effort 69 4.5.2 Future monitoring in Lamlash Bay 70 4.5.2.1 Sediment character 70 4.5.2.2 Distribution of biotopes 70 4.5.2.3 Extent of sub-features 70 4.5.2.4 Species composition and population measures 70

5 CONCLUSIONS 71 6 REFERENCES 73 6.1 General references 73 6.2 References used for video and still photography identification 76

Page 8: Mapping of the marine habitats and species of Lamlash Bay, Arran

iv

APPENDICES

Appendix 01. Survey equipment specification 77

Appendix 02. Navigation checks and tidal curves 78

Appendix 03. Field survey logs (2008 and 2009) 81

Appendix 04. Particle Size Analysis (PSA) 121

Appendix 05. Modified FOLK sediment classification 122

Appendix 06. Lamlash Bay Day Grab sample species list 123

Appendix 07. Lamlash Bay still photography analysis – species list (2008 and 2009) 128

Appendix 08. Lamlash Bay video analysis – summary of results (2008 and 2009) 132

Appendix 09. Faunal densities for noteworthy species 136

LIST OF FIGURES

Figure 1. Lamlash Bay located on the east coast (Firth of Clyde) of the Isle of Arran in North Ayrshire, Scotland.

1

Figure 2. Lamlash Bay No Take Zone boundary. 2

Figure 3. Seasearch survey sites in Lamlash Bay. 4

Figure 4. The bathymetric survey coverage (lines) for the Lamlash Bay habitat mapping survey 2008.

6

Figure 5. The bathymetric survey coverage (lines) for the Lamlash Bay habitat mapping survey 2009.

7

Figure 6. Sidescan sonar coverage for the 2008 Lamlash Bay habitat mapping survey.

10

Figure 7. Sidescan sonar coverage for the 2009 Lamlash Bay habitat mapping survey.

10

Figure 8. Camera system mounted on a similar seabed frame to that used during the Lamlash Bay survey.

12

Figure 9. Drop-down camera sampling locations (tracks) for the Lamlash Bay habitat mapping survey 2008.

12

Figure 10.

Drop-down camera sampling locations (tracks) for the Lamlash Bay habitat mapping survey 2009.

13

Figure 11. Sediment sampling locations for the Lamlash Bay habitat mapping survey 2008.

16

Figure 12. Sidescan sonar image with selected targets to illustrate some of the features identified in Lamlash Bay.

19

Figure 13.

Anchor sinkers seen in the southern section of Lamlash Bay (located approximately at East (X): 204779.6 and North (Y): 629190.2).

20

Figure 14. Anchor drag marks in the southern section of Lamlash Bay (located approximately at East (X): 204804.31 and North (Y): 629718.87).

20

Figure 15. Scallop dredge marks in the southern section of Lamlash Bay (circular pattern centred on East (X): 205270.52 and North (Y): 629791.71).

21

Page 9: Mapping of the marine habitats and species of Lamlash Bay, Arran

v

Figure 16. Exposed bedrock near the shallow sill in North Channel in Lamlash Bay (located approximately at East (X): 206105.47 and North (Y): 632224.95).

22

Figure 17. Deacon Rock in Lamlash Bay. 22

Figure 18. Sediment furrows in the centre of Lamlash Bay. 23

Figure 19. An area of low backscatter return in the south of Lamlash Bay. 23

Figure 20.

An area of high mottled backscatter return in the south of Lamlash Bay (located at East (X): 205110.26 and North (Y): 628864.17).

24

Figure 21.

An area of high, low and mottled backscatter illustrating cobbles, boulders and rocks surrounded by sand (located at East (X): 204182.71 and North (Y): 630457.02).

25

Figure 22. An area of mottled backscatter to the North West of Lamlash Bay where maerl is present (located at East (X): 204581.00 and North (Y): 631678.00).

25

Figure 23. An area of high backscatter on the shallow bank at the north-eastern Lamlash Bay where maerl is present (located at East (X): 205551.00 and North (Y): 632298.00).

26

Figure 24.

Mooring sinkers and chain along the north-western edge of Lamlash Bay (located at East (X): 203236.00 and North (Y): 630883.00).

27

Figure 25.

An area of mixed coarse sediment at the southern entrance of Lamlash Bay where maerl and macroalgae are present (located at East (X): 206527.00 and North (Y): 628297.00).

27

Figure 26.

An area of mixed backscatter along the north-eastern coast off Holy Island (located at East (X): 205026.00 and North (Y): 631114.00).

28

Figure 27.

Ground-truthing of target 5 - Deacon Rock (photograph LNC07_26 and 32).

30

Figure 28. Ground-truthing of target 7 - low backscatter return (photograph LSC03_20).

30

Figure 29. Ground-truthing of target 8 (photographs LSC04_49 and LNC05_48). 31

Figure 30. Ground-truthing of target 9 (photographs LNC10_21 and LNC08_20). 31

Figure 31. Ground-truthing of target 10 (photographs LNC11_10). 32

Figure 32. Ground-truthing of target 13 (photograph E01_05 and E01_19). 32

Figure 33. Ground-truthing of target 14 (photograph NTZ05_09 and NTZ05_18). 33

Figure 34.

Photographs from the southern and central regions of Lamlash Bay (LSC01_07, LSC02_29, LSC06_26 and LSC07_39).

35

Figure 35.

Photographs from transect A in the North Channel of Lamlash Bay (A01_06 and A02_11).

37

Figure 36.

Photographs from the northern Lamlash Bay and North Channel areas of Lamlash Bay (LNC03_09, LNC05_04, LNC07_35, LNC10_02, LNC10_06, LNC11_06, LNC11_20, LNC12_03).

38

Figure 37. Photographs from the Outer Lamlash Bay area (LNC01_09, LNC02_17, LOC01_35, LNC01_56, LNC01_71, LNC03_04, D01_02, D03_13).

40

Figure 38. Photographs from the southern entrance to Lamlash Bay (E01_07, E01_11, E01_26, E02_13).

41

Page 10: Mapping of the marine habitats and species of Lamlash Bay, Arran

vi

Figure 39. Examples of scallops observed in Lamlash Bay. 45

Figure 40. Maerl beds observed in and around Lamlash Bay. 46

Figure 41. Brittlestar beds in Lamlash Bay (LNC08_19, LNC08_20, LOC3_ 08 and LSC01_55).

47

Figure 42. Cumulative percentage sediment weight at the NTZ locations in Lamlash Bay.

49

Figure 43. Cluster analysis of the 2008 Lamlash Bay macrofaunal Day grab data. 52

Figure 44. Ordination analysis of the 2008 Lamlash Bay macrofaunal Day grab data (all 10 locations).

52

Figure 45. Further ordination analysis of the 2008 Lamlash Bay macrofaunal Day grab data (the 7 locations grouped).

53

Figure 46. Bathymetry and sidescan sonar data in Lamlash Bay acquired in 2008 and 2009 (bathymetry data overlaying the sidescan sonar imagery).

58

Figure 47. Biotope complex distribution in Lamlash Bay (from the video and stills photography analysis) overlaying the sidescan sonar imagery.

59

Figure 48. Bathymetry within the NTZ in Lamlash Bay acquired in 2008 and 2009 (overlaying the sidescan sonar imagery).

60

Figure 49. Biotope complex distribution within the NTZ in Lamlash Bay (from the video and stills photography analysis) overlaying the sidescan sonar imagery.

61

Figure 50. The biotope distribution within the NTZ in Lamlash Bay from photographic ground-truthing.

62

Figure 51. The sediment distribution within the NTZ in Lamlash Bay from photographic ground-truthing.

63

LIST OF TABLES

Table 1.

Coordinates of Lamlash Bay No Take Zone boundary (Scottish Government, 2008)

2

Table 2. Lamlash Bay 2008 acoustic survey areas. 6

Table 3. Lamlash Bay 2009 acoustic survey lines. 7

Table 4. Summary of video and still photography data collected during the 2008 Lamlash Bay surveys.

29

Table 5.

Summary of video and still photography data collected during the 2009 Lamlash Bay surveys.

29

Table 6.

Summary of the video and photographic analyses with the main substrata, fauna and biotope in central and southern Lamlash Bay.

34

Table 7.

Summary of the video and photographic analyses with the main substrata, fauna and biotopes (mixed sediments are muddy sandy gravel with shell material) in northern Lamlash Bay and North Channel.

36

Table 8.

Summary of the video and photographic analyses with the main substrata, fauna and biotopes (mixed sediments are muddy sandy gravel with shell material) in Outer Lamlash Bay.

39

Page 11: Mapping of the marine habitats and species of Lamlash Bay, Arran

vii

Table 9. Summary of the video and photographic analyses with the main substrata, fauna and biotopes (mixed sediments are muddy sandy gravel with shell material) at the southern entrance to Lamlash Bay.

41

Table 10.

The approximate densities (ind/100 m2) of scallops and two other species present in the still photographs from the 2008 Lamlash Bay survey.

42

Table 11.

The approximate densities (ind/100 m2) of scallops and two other species present in the still photographs from the 2009 Lamlash Bay survey.

43

Table 12.

The approximate densities (ind/100 m2) of scallops and two other species present in the video footage from the 2008 Lamlash Bay survey.

44

Table 13.

The approximate densities (ind/100 m2) of scallops and two other species present in the video footage from the 2009 Lamlash Bay survey.

44

Table 14. Day grab sampling locations. 47

Table 15.

Summary of the Particle Size Analysis (sediment weight by percentage) from the NTZ in Lamlash Bay.

48

Table 16.

Total percentage sediment weight (brown: ‘gravel’; yellow: ‘sand’; pink: ‘silt’) of the Day grab samples at the NTZ locations in Lamlash Bay (NB. processing differential resulting in total percentage being in excess of / less than 100%).

49

Table 17.

Total number of individuals of the most abundant taxa/species present in the 2008 Lamlash Bay grab samples (rank order).

50

Table 18. Total number of individuals (N), number of species (S), Margalef’s species richness (d), Pielou’s equitability index (J) and Shannon-Wiener diversity index (H’) for all the samples in the 2008 Lamlash Bay Day grab survey.

51

Table 19. SIMPER analysis of the 2008 macrofauna in Lamlash Bay. 54

Table 20. SIMPER analysis of taxa within the ‘fine’ cluster groups in the 2008 Lamlash Bay survey.

55

Table 21. Results from the Spearman rank correlation coefficient analysis comparing the faunal data with the sediment data collected at Lamlash Bay 2008.

55

Table 22. Summary of the biotope complexes / biotopes for the 2008 Lamlash Bay Day grab samples.

56

Table 23. Lamlash Bay monitoring attributes. 70

Page 12: Mapping of the marine habitats and species of Lamlash Bay, Arran

viii

Data Storage and display All of the data collected during the survey have been stored in the following formats:

• Data logs have been created in Excel and Word documents.

• All still photographs and images included in the report have been stored as JPEG.

• Raw video data have been stored digitally using MiniDV tapes and DVD discs.

• Edited video data supplied as MPEG files on DVD.

• All bathymetry data have been stores in ASCII text format (X, Y, Z).

• Raw sidescan sonar data have been stored in HyPack and XTF format.

• Mosaic sidescan sonar data have been stored in GeoTiff format.

• The GIS data have been stored in ArcView 9.3.

These are kept by Scottish Natural Heritage.

Page 13: Mapping of the marine habitats and species of Lamlash Bay, Arran

1

1 INTRODUCTION

1.1 Project background Seastar Survey Ltd. was contracted by Scottish Natural Heritage (SNH) to conduct a habitat mapping survey of Lamlash Bay and the adjacent area, Isle of Arran (figure 1). The survey follows the establishment of a No Take Zone (NTZ) via the Inshore Fishing (Prohibition on Fishing) (Lamlash Bay) (Scotland) Order 2008. This Order was introduced by the Scottish Parliament on 20th September 2008 and prohibits all forms of fishing within the NTZ (figure 2 and table 1). The aim of the survey was to carry out a baseline survey in Lamlash Bay to describe the seabed environment and map the seabed habitats in Lamlash Bay. The survey will inform the monitoring of the area to establish the success of the management measure in delivering benefits to both the biodiversity of the bay and the scallop fishery in the area. The survey area encompasses the whole of Lamlash Bay and an area outside the bay extending approximately 1 mile beyond the northern and southern entrances of the bay and seawards to the 30 m contour (figure 1), with the NTZ (figure 2) given the highest priority. Figure 1. Lamlash Bay located on the east coast of the Isle of Arran (Firth of Clyde) in North Ayrshire, Scotland.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 14: Mapping of the marine habitats and species of Lamlash Bay, Arran

2

Figure 2. Lamlash Bay No Take Zone boundary.

Table 1. Coordinates of Lamlash Bay No Take Zone boundary (Scottish Government, 2008)

Location Latitude Longitude

Mount Pleasant Farm 55˚ 32.603’ N 006˚ 06.512’ W Holy Isle West 55˚ 31.556’ N 005˚ 05.216’ W Holy Isle East 55˚ 31.876’ N 005˚ 04.304’ W Hamilton Rock 55˚ 33.000’ N 005˚ 04.823’ W

Clauchlands Point 55˚ 33.002’ N 005˚ 04.957’ W

Page 15: Mapping of the marine habitats and species of Lamlash Bay, Arran

3

In order to record and characterise the habitats in Lamlash Bay, Seastar Survey Ltd. undertook an acoustic and a ground-truthing survey (sediment sampling and video/still photography surveys) in 2008. Additional acoustic and camera surveys were carried out in 2009 to acquire data in the areas not surveyed in 2008. The aim of the acoustic element of the survey was to identify different backscatter returns and describe as well as delineate the extent of the various seabed habitat types occurring in and around Lamlash Bay. A digital sidescan sonar mosaic, in conjunction with single beam echo sounder derived bathymetry, provided the initial broadview to map the substrata present throughout the survey area as well as allowing the identification of any features of interest. The aim of the ground-truthing element of the survey was to provide a description of the richness and diversity of the habitats on both rocky reef and the softer sediments. The biotope distribution and species composition was developed through interpretation of drop-down video footage and digital still photography, taken after evaluation of the sidescan mosaic. Sediment samples (Day grabs) collected in 2008 were analysed to ground-truth the sidescan sonar data as well as describe and quantify the infaunal communities in the NTZ. The results of all elements of the survey were used to create a Geographical Information System (GIS) which enabled a high level of processing, interpretation and display of the sidescan sonar mosaic, bathymetry, substrata types, biotopes and the digital photography. 1.2 The environment Lamlash Bay is located on the east coast of the Isle of Arran in North Ayrshire, Scotland. Arran is the largest island in the Firth of Clyde, the most southerly fjord in the northern hemisphere (COAST, 2005). Arran is sheltered from the Atlantic by the Kintyre Peninsula and separated from mainland Scotland by the Firth of Clyde to the east and the Sound of Bute and the mouth of Loch Fyne to the north (Gazetteer for Scotland, 2008). The island is 36 km in length and 22 km wide (COAST, 2005). Lamlash Bay is approximately 5 km across, bounded by the headlands of Clauchlands Point to the north and Kingscross point to the south (COAST, 2005; Gazetteer for Scotland, 2008). Over half of the bay is occupied by Holy Island which is approximately 3 km long and 1.5 km wide (COAST, 2005; Gazetteer for Scotland, 2008).

1.2.1 Geology and the sedimentary environment

There is limited published information regarding the seabed environment in Lamlash Bay, however, the northern and southern margins of the bay are dominated by rocky coastlines with sublittoral rock being present along the northern coast and the northern channel (Duncan, 2003). The south-western coast of Holy Island consists of a steep rocky slope that descends to 20 m below chart datum (Duncan, 2003). On admiralty charts the seabed within the bay is generally dominated by mud and sand with additions of broken shell particularly in the southern area of Lamlash Bay and along the western side of Holy Island. The North Channel and the outer reaches of the survey area are dominated by mixed sediments (sand, mud, broken shell and gravel). These records seem to be in overall agreement with the observations made by Duncan (2003), reporting the sediment to be primarily composed of soft sediments with sand being the most common sediment type. Areas of mixed sediment (rock, cobble and mud) and other areas dominated by boulders have also been identified (see Duncan, 2003), the latter being a feature not identified on the admiralty chart.

Page 16: Mapping of the marine habitats and species of Lamlash Bay, Arran

4

1.2.2 The physical environment

The maximum water depth in Lamlash Bay is 43 m below chart datum (admiralty charts) with the majority of the area being > 20 m deep below chart datum. Shallower sills exist at the northern and southern entrances to the bay. The sill at the southern entrance of the bay has a maximum water depth of 25 m below chart datum, at the northern entrance to the bay water depths are recorded as 11.3 m below chart datum (Seasearch 2003, COAST 2005). Current speeds range from approximately 0.5 knots (neap tides) to 0.75 knots (spring tides) at the entrance of the North Channel and from 0.75 knots (neap tides) to 1.5 knots (springs) at the entrance of the South Channel.

1.2.3 Biological distributions

There is a limited amount of published information regarding the biological distributions in Lamlash Bay and prior to 2003 no records existed for Lamlash Bay within the Marine Nature Conservation Review (MNCR) database (now recorded as NBN Marine Recorder) (Duncan, 2003; COAST, 2005). In 2003, however, Seasearch divers surveyed 21 sites in Lamlash Bay (results given in figure 3) to create a basic sublittoral baseline for the area (Duncan, 2003). The sites are marked by flags with the small blue circles illustrating the exact position of the site location. Sites with maerl are highlighted in purple and scallops (abundance estimated as ‘common’ and/or noteworthy on the Seasearch forms) in brown (locations 6-8, 10, 12, 14 and 20). Sites with both maerl and scallops (‘common’) were recorded in blue (sites 7 and 16) and sites with both sea grass and maerl were recorded in green (2). All other sites were recorded in orange (see Duncan (2003) for further details). Figure 3. Seasearch survey sites in Lamlash Bay (from Duncan, 2003).

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 17: Mapping of the marine habitats and species of Lamlash Bay, Arran

5

Of particular note is the presence of maerl and sea grass (Zostera sp.) beds in the North Channel as well as the presence of ‘reefs’ (e.g. Deacon Rock). Although none of the species recorded in Lamlash Bay are listed as Annex II species in the EC Habitats Directive, both maerl and Zostera sp. are listed on the UK Biodiversity Action Plan (UKBAP) Habitat Action Plan as priority habitats (UKBAP, 2008) and included in the Scottish Biodiversity List (BiodiversityScotland, 2008). In addition to being listed as a priority habitat on the UKBAP’s Habitat Action Plan, maerl beds are covered by four different habitat types of Annex I of the EC Habitats Directive ‘sandbanks which are slightly covered by seawater at all times’, ‘large shallow bays and inlets’; ‘estuaries’ and the priority habitat ‘lagoons’ (UKBAP, 2008). Apart from the ecologically important species such as Phymatolithon (maerl) and Zostera sp. (eelgrass), other taxa recorded in the bay (see Duncan, 2003) include Cerianthus lloydii, Metridium senile, Echinus esculentus and species in the subphylum Pisces (fish); dogfish, conger eels (Conger conger), leopard-spotted gobies (Thorogobius ephippiatus), pipefish (Syngnathinae), gurnards (Triglidae), dragonets (Callionymus lyra) and plaice (Pleuronectes platessa) as well as commercially important species such as scallops (Pectinidae), edible crabs (Cancer pagurus) and flatfish (Pleuronectiformes).

2 METHODOLOGY

The 2008 field work (acoustic and biological surveys) was undertaken between the 13th and 21st October 2008 with all survey operations conducted from the Fisheries Research Services’ (FRS) survey vessel Alba na Mara. The survey mobilisation and de-mobilisation took place in Ardrossan Harbour, Scotland on 13th October and 21st October respectively. During the survey the vessel anchored overnight in either Lamlash Bay or Brodick Bay. The draught of Alba na Mara restricted the 2008 survey operations to areas outside the 10 m contour. The additional survey work required to acquire baseline environmental data from the shallow sections (areas inside the 10 m contour) was carried out in 2009. The 2009 field work (acoustic and drop-down camera surveys) was undertaken between 9 and 13 September 2009 with all survey operations conducted from Seastar Survey Ltd.’s vessel Otarie. The survey mobilisation and de-mobilisation took place in Ardrossan Harbour, Scotland, on 7th September and 14th September respectively. 2.1 The acoustic surveys The acoustic surveys in 2008 and 2009 were carried out using a Leica 1200 RTK Rover GPS system and an EdgeTech FS 4200 dual frequency sidescan sonar system (see appendix 1 for detailed equipment specifications) but the echosounder used in 2008 was a Simrad EK60 split beam echosounder whilst the 2009 echosounder system comprised a Marimatech E-Sea Sound 206 dual frequency echosounder (33/200 kHz). The 2008 acoustic survey area was divided into four main areas of planned acoustic lines (figure 4 and table 2) to simplify survey operations. SNH defined the area within the NTZ as the priority survey area. The survey effort was therefore structured to ensure that the acoustic and ground-truthing was completed in its entirety within the NTZ in the event that the survey work was delayed preventing the completion of the survey work in the remainder of the area.

Page 18: Mapping of the marine habitats and species of Lamlash Bay, Arran

6

Table 2. Lamlash Bay 2008 acoustic survey areas.

Survey area Number of lines Line numbers Total length (km)

Lamlash North 15 101-115 48,882 Lamlash South 19 201-219 59,994 Lamlash Outer 10 401-410 58,496 Deacon Rock 11 301-311 11,807

Total length (km) 179,179

Figure 4. The bathymetric survey coverage (lines) for the Lamlash Bay habitat mapping survey 2008 (with the four main areas illustrated by 1) Lamlash North, 2) Lamlash South, 3) Lamlash Outer and 4) Deacon Rock).

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

The 2009 acoustic survey was divided into nine different areas (A – I) and the locations of these lines were selected in the planning stage of the project (see figure 5 and table 3). As previously, SNH defined the area within the NTZ as the priority survey area. The survey effort was again structured to ensure that the acoustic and ground-truthing was completed in its entirety within the NTZ in the event that the survey work was delayed preventing the completion of the survey work in the remainder of the study area.

Page 19: Mapping of the marine habitats and species of Lamlash Bay, Arran

7

Table 3. Lamlash Bay 2009 acoustic survey lines (line C03 not surveyed).

Survey area Number of lines Line numbers Total length (km)

Area A 9 A01 – A09 8,638 Area B 6 B01 – B06 17,363 Area C 10 C01 – C11 15,428 Area D 18 D01 – D18 36,698 Area E 10 E01 – E10 18,059 Area F 3 F01 – F03 3,204 Area G 6 G01 – G06 4,218 Area H 6 H01 – H06 6,497 Area I 6 I01- I06 9,491

Total length (km) 119,596 km

Figure 5. The bathymetric survey coverage (lines) for the Lamlash Bay habitat mapping survey 2009 (with the nine main areas illustrated). .

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 20: Mapping of the marine habitats and species of Lamlash Bay, Arran

8

2.1.1 Horizontal control

Horizontal control was achieved using a Leica Real Time Kinematic (RTK) GPS (GX1230 Real Time Rover System). In 2008 the GPS antenna was mounted on the handrail on the upper deck above the bridge (on Alba na Mara) and in 2009 it was pole-mounted at the stern of Mariner to ensure a signal could be received from the maximum number of satellites available. The GPS obtained a satellite derived position in WGS84 latitude and longitude which was recorded using the Hypack 2008 survey management software. A data transformation was undertaken within the Hypack software using the Ordnance Survey model OSTN02 to convert from WGS84 latitude and longitude to OSGB36 easting and northing. A known position within Ardrossan Harbour was obtained from the Ordnance Survey in OSGB36 grid coordinates. A navigation check was then carried out against that known position at the start and the end of the survey to check the accuracy of the RTK GPS signal (see navigation checks in appendix 02).

2.1.2 Vertical control

2.1.2.1 Tides

Tide corrections were achieved via the vertical component of the RTK GPS. The raw bathymetric soundings produced by the echosounder were reduced relative to Ordnance Datum Newlyn (ODN) using the Ordnance Survey OSGM02 model within Hypack. This allowed for the tidal component to be removed from the raw soundings in real-time (see example of tidal curve in appendix 02).

2.1.2.2 The 2008 split beam echosounder

Vertical control during the 2008 survey was achieved using the Alba na Mara’s Simrad EK60 scientific split beam echosounder (38/200 kHz) with the coverage of the survey illustrated in figures 4 and 5. The vessels echosounder transducers are pole mounted in a pod fitted to a shaft which enabled the pod to be deployed to 1 m below the keel to improve acoustic performance. When the echosounder pole was down the transducers were approximately 5.25 m below the sea surface. It was intended that the high frequency (200 kHz) channel would be used for the survey. However, the data provided by the high frequency channel was found to experience signal loss when the vessel was moving; therefore the low frequency (38 kHz) data was recorded for the duration of the survey. The speed of sound through the water column was measured each day using a Marimatech HMS 1820-P CTD. Additional sound velocity profiles were taken when the survey vessel moved to a different survey area. The sound velocity profiles have been applied to the bathymetric data during processing within the Hypack survey management software. The bathymetric soundings were reduced relative to ODN to account for the effect of tide in real-time within Hypack, via the vertical component of the RTK GPS. Hypack applied the real time corrections using the Ordnance Survey OSGM02 model.

Page 21: Mapping of the marine habitats and species of Lamlash Bay, Arran

9

2.1.2.3 The 2009 dual frequency echosounder

Vertical control during the 2009 survey was achieved using a Marimatech E-Sea Sound dual-frequency (33 and 200 kHz) digital echosounder. Data from both frequencies were recorded on a paper trace and digitally using the Hypack survey management software. The echosounder transducer was pole mounted over the port side of the vessel and secured using fore and aft stays to prevent any movement of the transducer head. Bathymetric survey coverage from the 2008 and 2009 surveys are illustrated in figure 4 and 5. The bathymetric soundings were reduced relative to ODN to account for the effect of tide in real-time within Hypack, via the vertical component of the RTK GPS. Hypack applied the real time corrections using the Ordnance Survey OSGM02 model. The speed of sound through the water column was measured each day using a Marimatech HMS 1820-P CTD. Additional sound velocity profiles were taken when the survey vessel moved to a different survey area. The sound velocity will be applied to the bathymetric data during processing within the Hypack survey management software.

2.1.3 Vessel motion reference

Possible errors in the bathymetric data relating to vessel motion were minimised by the use of a TSS CMS 25 Compact Motion Sensor, to account for any vessel motion to an accuracy of ±5 cm. Corrections due to vessel motion have been applied during processing using the Hypack software.

2.1.4 Sidescan sonar

Seabed backscatter data was collected using an EdgeTech 4200-FS dual frequency (120/410 kHz) sidescan sonar. During the survey the unit was operated using the high frequency at 410 kHz. The system comprised a towfish, a slip-ring winch carrying 700 m of coax cable and transceiver. The sidescan data was viewed in real-time through Hypack. Heading and layback information were also provided through Hypack. Data was recorded in HS2 format (Hypack sidescan data file). In 2008 the line spacing was set at 50 m with a range of 50 m (covering 50 m either side of the fish – 100m swath). The line spacing allowed for 100 % overlap of the sidescan data with adjacent lines. The towfish was deployed from the Alba na Mara’s CTD winch. The towfish was lowered into the water by the ships crew. Once the towfish was at the sea surface control of the winch was transferred to winch controls in the survey lab, which were operated by the surveyor. The sidescan was towed at speeds of 4-5 knots. The line spacing in 2009 was set at 50 m with a range of 100 m (covering 100 m either side of the fish creating a 200 m swath). The line spacing allowed for a 200% overlap of the sidescan data with adjacent lines. The towfish was deployed from a capstan winch on a rope and connected to the surface using a 200 m soft tow cable which was deployed at the same rate as the rope. As in 2008 the sidescan was towed at speeds of 4-5 knots. The sidescan sonar survey resulted in a coverage illustrated in figures 6 and 7 (see survey limitations below).

Page 22: Mapping of the marine habitats and species of Lamlash Bay, Arran

10

Figure 6. Sidescan sonar coverage for the 2008 Lamlash Bay habitat mapping survey.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Figure 7. Sidescan sonar coverage for the 2009 Lamlash Bay habitat mapping survey.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 23: Mapping of the marine habitats and species of Lamlash Bay, Arran

11

2.1.5 Weather conditions

The weather conditions in both 2008 and 2009 were overall suitable for survey operations to take place throughout the survey periods. The only exceptions were on 19 October 2008 when weather downtime was incurred as a result of strong wind (Force 7 - 8), rough seas (moderate – rough), heavy rain and poor visibility; and on 8 September 2009 when strong winds (Force 6 – 8), a rough sea state (rough or very rough) and moderate to poor visibility prevented survey operations from taking place. In these conditions it was deemed unsafe for the vessel to continue with camera work or to carry out grabbing and survey operations were ceased.

2.1.6 Processing and charting

Processing of the bathymetric data was undertaken using the post processing tools within the survey management software. This was undertaken with reference to the echo sounder paper trace records as part of our quality control procedures. The processing of the data involved editing out the water column, the removal of spikes and other erroneous points from the data and the reduction of the depths using corrected data acquired from the RTK. The data were then imported into an ArcView GIS for display and interpretation. The sidescan sonar data were initially processed and mosaiced within the survey management software. This involved trimming the edges wherever possible and appropriate as well as mosaicing the sidescan traces together to obtain a map as complete as possible with a resolution of 2 m bins (i.e. 1 pixel equals 2 m of seafloor). Paper print-outs and a GeoTiff were delivered. This data was then also imported into the ArcView GIS, and again the GIS was used as the medium of display and ongoing data analysis and interpretation. 2.2 Drop-down camera survey methodology

2.2.1 Drop-down camera system

The camera equipment used during the survey was a Kongsberg Simrad OE14-208 Digital Stills Camera System mounted obliquely on a seabed camera frame (figure 8). A flash unit and two sub-sea lights were also mounted on the frame. These units were linked to the surface with a soft umbilical cable. All controls of the camera system were kept in the wheelhouse of the survey vessel.

2.2.2 Camera operation and deployment

The two drop-down camera surveys were carried out between 16th and 20th October 2008 and 9th and 13th September 2009 respectively. The camera system was only deployed in suitable weather conditions by a team of experienced personnel. Before each deployment a ‘clapper board’ containing site name, date and weather conditions was videoed and photographed as a quality assurance (QA) record. All camera deployments (figures 9 and 10) were established as transects across sites of interest, which were selected after scrutinising the acoustic survey data (figures 6 and 7). The vessel was positioned at one end of each transect using DGPS. The camera was then deployed and lowered to the seabed. Once the camera system was at the seabed and had time to settle, the onboard surveyors started to log navigation and the skipper was given approval to start drifting along the transect at about 0.5 knots.

Page 24: Mapping of the marine habitats and species of Lamlash Bay, Arran

12

Figure 8. Camera system mounted on a similar seabed frame to that used during the Lamlash Bay surveys (red rectangle: camera; yellow rectangle: flashgun and green rectangle: sub-sea light).

Figure 9. Drop-down camera sampling locations (tracks) for the Lamlash Bay habitat mapping survey 2008.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 25: Mapping of the marine habitats and species of Lamlash Bay, Arran

13

Figure 10. Drop-down camera sampling locations (tracks) for the Lamlash Bay habitat mapping survey 2009.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

During the 2008 deployments, the control of the camera and winch were solely with the camera operator. The winch was controlled using a remote control, whilst photographs were taken using a surface trigger in the wheel house. The 2009 deployments were carried out manually (using a manual winch fitted on a davit) with verbal instructions given from the camera operator to the winch controller to direct the camera operations. The camera system sent a continuous, real-time, analogue video feed to the surface, where the deployment was monitored and the camera / winch were controlled. The analogue video feed was recorded digitally. In 2008 the signal was recorded using Mini Digital Video (miniDV) tapes and backed-up using super VHS tapes whilst in 2009 the signal was recorded on DVDs with a hard-disc recorder being used as a back-up. In 2008 the video was recorded to tape rather than DVD as the quality and life expectancy of tape is much greater but in 2009 this facility was not available. Individual still photographs were taken using a surface controlled trigger. Photographs were taken regularly at random along the transects to capture the general seabed environment and the fauna present but also at the discretion of the camera operator to capture particular taxa (to allow identification) and ensure that any changes in the seabed environment were recorded.

Page 26: Mapping of the marine habitats and species of Lamlash Bay, Arran

14

Throughout each camera deployment, navigation data was recorded. All camera deployment log keeping was synchronised to the navigation data using the time from the RTK GPS. The log keeper recorded the time from the GPS at the start and end of each deployment and the time each photograph was taken. After each deployment the camera was removed from the frame and secured in the wheel house. The digital photographs were then uploaded from the camera to a laptop computer via a USB lead (the software used for this was Canon Zoom Browser EX). During the upload process each photograph was named with the site-name and photograph number.

2.2.3 Data handling

The photographs and video footage were then used for analysis in the GIS. When the camera surveys were completed the miniDV tapes and DVDs were taken back to Seastar Survey Ltd. offices by a member of staff and the backup VHS tapes and hard drives were transported to an offsite data storage facility. After the survey the digital miniDV tapes were up-loaded to a computer, edited, titled and burnt to DVD as mpeg files. All DVDs, photographs and logs were checked for errors as part of Seastar Survey Ltd’s standard quality control procedures and all data supplied to the client. Finally the client was supplied with DVDs with mpeg video files, sets of all seabed photographs and the seabed photographs were also incorporated into the ArcView GIS. 2.3 Video and photographic analysis

2.3.1 Analysis of the photographs and video records

The analysis of the photographs and video records was carried out ‘blind’, without any prior knowledge of the sites, using a personal computer and a SONY digital videocassette recorder and a television monitor respectively. The latter system allows slow-motion, freeze-frame and standard play analysis. An initial assessment of a site was carried out by first looking through the photographs and film from the particular site quickly to get a brief understanding of the substratum, flora and fauna.

2.3.1.1 Semi-quantitative analysis of the video footage

The video analysis included an initial assessment of a deployment to get a broad understanding of the substratum, flora and fauna as well as the identification of the different biotopes/habitats on the seabed. The boundary positions of the different biotopes/habitats were determined using the time codes and related back to the navigation data. The detailed analysis consisted of a general seabed description and the fauna were identified to the lowest practical level. General descriptions of the faunal life-forms were made and any other features of interest such as trawl marks were also recorded. The abundance data were recorded using the SACFOR scale (general fauna) and actual abundance (commercial taxa) with the analysis concentrating on the commercially important species present together with species readily identified in the video material. A list of the encountered fauna was produced for each site using species reference numbers as cited in the Marine Conservation Society Species Directory (Howson and Picton 1997) to avoid problems in species nomenclature. The sections or deployments were subsequently classified into designated biotopes according to Connor et al. (2004). Faunal densities were calculated based on the distance covered for each habitat, biotope or transect, basing the calculations on the estimated assumption that the camera covers 1 m in width (previous

Page 27: Mapping of the marine habitats and species of Lamlash Bay, Arran

15

camera calibration test gives a seafloor coverage of approximately 0.8 m by 0.8 m when the camera frame is in contact with the seabed). The distances for each habitat were calculated from the position of the start of each habitat / biotope to the position of the end of each habitat / biotope rather than attempting to calculate the exact distance along the transect resulting in approximate density values.

2.3.1.2 Quantitative analysis of the stills photographs

The still photography analysis was carried out using a personal computer. The methodology is in large parts similar to the video analysis methodology described for the video analysis and included a general seabed description where the substrata were described according to the Wentworth scale (see Leeder, 1982) with boulders and cobbles being described within ‘gravel’ and rock referring to bedrock. The fauna was identified to the lowest practical level with the nomenclature according to the Marine Conservation Society Species Directory (Howson and Picton 1997) and actual abundance of the fauna and flora was recorded whenever possible (faunal densities are based on the approximation that each photograph covers 1 m2; see camera calibration notes above). The presence of faunal life-forms and any other features of interest such as trawl marks were also recorded. Each photograph was subsequently classified into designated biotopes according to Connor et al. (2004), wherever possible. These results were subsequently incorporated into the ArcView GIS. 2.4 Sediment sampling Sediment samples were taken in 2008 using a 0.1 m2 Day grab. Grab samples were taken at 10 locations across the NTZ (figure 11). The grab sites were selected to represent the different sediment types indicated by the sidescan sonar data and seabed video footage and to allow additional data for the biotope classification process. The number of sites was restricted to the NTZ as a result of limited funding but also as the photographic material at the shallow sites was considered to be of sufficient quality to allow biotope classification. A further consideration was the potential of damaging the maerl beds had the grab sampling survey been extended to include the shallow sections. Prior to deployment the grab was cleaned out using the vessels seawater deck wash to remove any remaining sediment or contamination. The Day grab was then deployed by the vessels crew from the crane on the port side of the vessel. The grab was lowered to the seabed until the cable went slack indicating that the grab had reached the seabed. The grab was then recovered to deck where the sample was inspected by the environmental scientist. Grab samples were rejected on the grounds of poor quality for the following reasons:

• Uneven surface indicative of striking the seabed at an angle

• Washed out sample

• Disturbed surface sediment

• Contamination of the sediment (e.g. hagfish, paint chips, oil)

• Sample touching the top of the grab

• Sample <40% of the grabs capacity If the grab sample was acceptable a photograph was taken with a sample label in the image indicating the sampling area and drop number. A small sub sample (approximately 100g) was taken from each of the grab samples for particle size analysis (PSA). The contents of the grab were then washed into a collecting tray under the grab stand. The sediment was gently washed over a 1 mm sieve. The material retained on the sieve was then transferred into a 5 dm3 (litres) plastic bucket and fixed using a 4 % buffered formaldehyde solution to prevent sample degradation.

Page 28: Mapping of the marine habitats and species of Lamlash Bay, Arran

16

Figure 11. Sediment sampling locations for the Lamlash Bay habitat mapping survey 2008.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

The processing of the macrofaunal sediment samples took place at Seastar Survey’s laboratory in Southampton. The sediment samples were gently re-sieved on 1 mm sieves and sorted (using trays and low-magnification microscopes). The fauna was subsequently identified to the lowest practical level (species in most cases) with reference to the species directory (Howson and Picton, 1997) to avoid problems with nomenclature. A full list of taxa encountered and abundances per sample are recorded on a standard species / sample matrix (appendix 06) A reference collection was created and a Quality Control (QC) exercise was also carried out by Artoo Marine Ecological Consultants to check the identification results. The invertebrate specimens collected were separated by species and by station, preserved in alcohol, and stored in glass sample vials with polyethylene closures to facilitate their incorporation into the collections of The National Museum of Scotland. The particle size analysis also took place in Seastar Survey’s laboratory following standard protocols using a combination of wet and dry sieving techniques at 1 phi intervals. The PSA followed the sediment grades used by the MNCR (Marine Nature Conservation Review) as described below:

Page 29: Mapping of the marine habitats and species of Lamlash Bay, Arran

17

Pebble – medium (> 8 mm) Pebble – small (4-8 mm) Granule (2-4 mm)

Sand very coarse (1000-2000 µm)

Sand coarse (500-1000 µm)

Sand medium (250-500 µm)

Sand fine (125-250 µm)

Sand very fine (63-125 µm)

Silt and clay (mud) (<63 µm) The dry weight of the whole sediment sample was determined and any muddy samples were disaggregated using a suitable method (e.g. sodium hexametaphosphate). The sample was

wet sieved on a 63 µm mesh then dried and re-weighed to establish the weight percentage of

the sub 63 µm fraction. The remainder of the sample was dry sieved with an appropriate sequence of mesh sizes to yield weight percentage data for particle size fractions at half phi

intervals. This was done from a minimum sieve size of 63 µm to a maximum sieve size of 16 mm. 2.5 Data analysis Ecological interpretation of the results was carried out by reference to the JNCC Marine Nature Conservation Review Biotope Classification (Connor et al., 2004), and the presence of rare or unusual species identified against the list in Sanderson (1996). All the identified and enumerated benthic macrofauna were used in the analysis and the abundance data are given in the SACFOR scale (see Connor et al., 2004) for the video records and actual abundance, given as number of individuals/m2, for the still photographs. The macrofaunal sediment data are given in number of individuals/0.1 m2. The data analyses comprised both univariate and multivariate analyses all of which were calculated using Primer (Plymouth Routines in Multivariate Ecological Research) v 5.2.0 (Clarke and Warwick, 1994). The univariate analysis included the total number of individuals (N), total number of species (S), species diversity where the Shannon-Wiener (H’), Pielou’s (J) and Margalef’s (d) diversity indices (see e.g. Gage and Tyler, 1991; Fowler and Cohen, 1992; Clarke and Warwick, 1994) were used with the natural log (loge) being the chosen parameter in the case of the Shannon-Wiener diversity index. The multivariate analysis was carried out using cluster analysis and ordination (non-metric multi-dimensional scaling, MDS). These data were then transformed to square root to down-weigh the importance of common types of macrofauna in relation to rarer types. The transformed data were then analysed using the Bray-Curtis similarity coefficient (using Primer v.5) followed by a cluster analysis where the sites were group averaged and the resultant dendrogram plotted. Non-metric multi-dimensional scaling (MDS) was then carried out to further assess the presence of any similarities between sites (Clarke and Warwick, 1994). Spearman’s rank correlation coefficient (see e.g. Fowler and Cohen, 1992) was then used to assess any correlations with the particle size analysis variables. The BIOENV routine in PRIMER was used to carry out this task on untransformed abiotic data. The SIMPER routine in PRIMER was subsequently used to assess the difference in characteristic species/ taxa in the samples.

Page 30: Mapping of the marine habitats and species of Lamlash Bay, Arran

18

2.6 Survey limitations

2.6.1 Survey vessel

The main factor limiting the Lamlash Bay survey in 2008 was the use of the vessel Alba na Mara as the draught of the vessel (5.25 m with the transducer pole down) prevented any survey operations shallower than the 10 m contour. This excluded a large section of Lamlash Bay area from the survey including some areas in the north of the No Take Zone believed to be dominated by rocky ‘reef’ habitats (as defined in the European Habitats Directive 92/43/EEC). This issue was raised during the survey operations but after consultation SNH decided to complete the current stage of the survey using Alba na Mara. The second survey in 2009 using a different vessel was therefore required to cover the entire Lamlash Bay study area.

2.6.2 Obstructions

In 2008 a line of fishing pots prevented a large section in the centre of the Lamlash Outer (the survey area outside Lamlash Bay) from being surveyed. In addition, a section in the south-west was also excluded from the survey as a result of the presence of fish farm cages. In 2009 the planned lines C01 to C04 were not completed because of obstructions in the water associated with the mussel farm. Line B03 was cut short at the southern end as a result of the presence of mooring buoys. The presence of fishing pots obstructed lines D08, D09, D10 and D11, although sidescan sonar data were acquired from this area.

2.6.3 Equipment failure

During the 2009 survey the portside high frequency transmitter on the towfish failed (9 September 2009) and it was not possible to repair the towfish in the field. Drop-down camera operations were therefore conducted in the NTZ whilst waiting for a new sidescan sonar towfish to be delivered.

3 RESULTS

3.1 Acoustic analysis The processing and analysis of the acoustic data (bathymetric data and sidescan sonar mosaic) from the acoustic survey (410 kHz sidescan sonar and 38/200 kHz single beam echo sounder) revealed areas of different levels of backscatter, suggesting clear changes in bathymetry, sediment type and or sediment composition within the survey area (see figures 6 and 7). These patterns and features were used to direct the selection of deployment locations for the underwater video (see figure 9 and 10) and still photography system as well as the sediment grab sampling (see figure 11). The analysis of the sidescan sonar mosaic reveals a number of areas with a low backscatter return, particularly in the inner, deeper parts of Lamlash Bay (central and southern sections) whilst high backscatter returns are found in shallower water, particularly around the northern and southern sections of the North Channel, a number of small areas northwest of Holy Island and along the coast around Holy Island and in the Outer Lamlash area (see figures 6 and 7). To illustrate some of these features, both natural and anthropogenic, a number of targets (sections of the sidescan sonar mosaic) have been selected (figures 13 - 26) and the positions of the features are given in figure 12 (NB. the sidescan sonar images are displayed using inverted grey scale, therefore shadows appear black).

Page 31: Mapping of the marine habitats and species of Lamlash Bay, Arran

19

Figure 12. Sidescan sonar image with selected targets to illustrate some of the features identified in Lamlash Bay.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

3.1.1 Target 1 - anchor sinkers

Two anchor sinker marks (approximately 1 m x 2 m in dimension with multiple chains) were noted at Target 1 in the southern part of Lamlash Bay (figure 13). The distance between the two sinkers is approximately 14 m. This is a good example of the detail and clarity of the data obtained by the sidescan sonar and illustrates one of the human activities / impacts in the bay.

Page 32: Mapping of the marine habitats and species of Lamlash Bay, Arran

20

Figure 13. Anchor sinkers seen in the southern section of Lamlash Bay (located approximately at East (X): 204779.6 and North (Y): 629190.2).

3.1.2 Target 2 - anchor drag marks

Anchor drag marks with sediment build up are present in the central and southern sections of Lamlash Bay (figure 14). This image is approximately 50 m wide with the anchor drag marks being approximately 58 m long. There are several of these drag marks across this section of the bay, some of which are over 200 m long. Figure 14. Anchor drag marks in the southern section of Lamlash Bay (located approximately at East (X): 204804.31 and North (Y): 629718.87).

Page 33: Mapping of the marine habitats and species of Lamlash Bay, Arran

21

3.1.3 Target 3 - scallop dredge marks

There is some evidence of fishing activity along the west coast of Holy Island, in this case scallop dredging activity, with the dredge marks (individual tracks are approximately 5 m wide) creating a circular pattern centred around a small mound (figure 15) just outside the survey area. The conical shapes of the positive targets also visible on the sidescan image are possibly a result of deposition of sediment material from the dredges. These features were detected at the outer limit of the sidescan sonar trace (as a result of the relatively large draught of the survey vessel) but the additional sidescan sonar coverage in the shallow water along the Holy Island west coast (in 2009) did not reveal any further evidence of this activity. Figure 15. Scallop dredge marks in the southern section of Lamlash Bay (circular pattern centred on East (X): 205270.52 and North (Y): 629791.71).

3.1.4 Target 4 - exposed bedrock

Target 4 is from North Channel and it consists of an area of high backscatter return with relief, surrounded by a low backscatter. This target has been interpreted as exposed bedrock and boulders surrounded by coarse (boulder, cobble and gravel debris) and soft sediment (sand) types (figure 16). The bedrock target is approximately 24 m long, 12 m wide and 1.8 m high.

Page 34: Mapping of the marine habitats and species of Lamlash Bay, Arran

22

Figure 16. Exposed bedrock near the shallow sill in North Channel in Lamlash Bay (located approximately at East (X): 206105.47 and North (Y): 632224.95).

3.1.5 Target 5 - Deacon Rock

Target 5 (figure 17) is a section (image width is approximately 100 m) of Deacon Rock with a mottled appearance of high and low backscatter with some relief discernable in the high backscatter returns. This seabed environment has been interpreted to consist of boulders (largest being approximately 0.8 m in height), cobbles and other coarse sediment types surrounded by sediments returning low backscatter, suggesting the presence of sand.

Figure 17. Deacon Rock in Lamlash Bay (located at East (X): 204509.00 and North (Y): 630898.00).

Page 35: Mapping of the marine habitats and species of Lamlash Bay, Arran

23

3.1.6 Target 6 - sediment furrows

Target 6 is from an area of sediment giving low backscatter returns, interpreting the sediment as sand. Figure 18 illustrates several sediment furrows (the approximate image dimensions are: 80 m in length and 60 m in width). The origin of these furrows is unknown. Figure 18. Sediment furrows in the centre of Lamlash Bay (located at East (X): 204081.00 and North (Y): 630704.00).

3.1.7 Target 7 - low backscatter return (sand)

The most predominant seabed sediment feature across Lamlash Bay is illustrated in figure 19 (the image is approximately 50 m wide). This low backscatter return is interpreted as muddy sand and a feature present throughout the central and southern sections of Lamlash Bay. These areas are interspersed with areas of coarser sediment (including gravel) and shell debris. Figure 19. An area of low backscatter return in the south of Lamlash Bay (located at East (X): 205989.00 and North (Y): 628454.00).

Page 36: Mapping of the marine habitats and species of Lamlash Bay, Arran

24

3.1.8 Target 8 - mottled high and low backscatter

Areas of mottled high and low backscatter are interpreted as gravel interspersed with sand. It is a seabed environment found across Lamlash Bay and in central and northern sections in particular; appearing as mottled high and low backscatter returns (figure 20). The image is approximately 50 m wide and the largest boulder is about 1 m in height (1.5 m wide and 2.5 m long). Figure 20. An area of high mottled backscatter return in the south of Lamlash Bay (located at East (X): 205110.26 and North (Y): 628864.17).

3.1.9 Target 9 - mottled high and low backscatter In some of the shallower areas of Lamlash Bay, such as the central region at Deacon Rock and southern and northern boundaries of North Channel, the seabed comprised areas of coarse sediment. This mottled high back scatter return with relief is interpreted as cobbles and small and large boulders, surrounded by sediments returning low backscatter, suggesting the presence of sand (figure 21). The images are approximately 100 m wide.

Page 37: Mapping of the marine habitats and species of Lamlash Bay, Arran

25

Figure 21. An area of high, low and mottled backscatter illustrating cobbles, boulders and rocks surrounded by sand (located at East (X): 204182.71 and North (Y): 630457.02).

3.1.10 Target 10 - area of seabed with Maerl The strong returns and mottled appearance are indicative of very coarse sediment (figure 22). Maerl was found to be present at shallower regions, in particular to the north of Lamlash Bay (North Channel) where coarse sediment is present. The image (figure 22) is approximately 50 m across. Figure 22. An area of mottled backscatter to the North West of Lamlash Bay where maerl is present (located at East (X): 204581.00 and North (Y): 631678.00).

Page 38: Mapping of the marine habitats and species of Lamlash Bay, Arran

26

3.1.11 Target 11 – high backscatter area with maerl

Target 11 has been selected from the shallow bank in the north-eastern corner of Lamlash Bay. The image (figure 23) illustrates an area of mottled and high backscatter returns in a section of the seabed where maerl is present together with macroalgae (brown and red seaweed). The image is approximately 20 m across. Figure 23. An area of high backscatter on the shallow bank in the north-eastern Lamlash Bay where maerl is present (located at East (X): 205551.00 and North (Y): 632298.00).

3.1.12 Target 12 – mooring sinkers and chain

There were a number of obstructions during the field work period one of which was in the north-western corner of Lamlash Bay where a number of moorings were present. The presence of these moorings is illustrated in target 12 (figure 24) where mooring sinkers and chain can be seen. The image is approximately 20 m across.

Page 39: Mapping of the marine habitats and species of Lamlash Bay, Arran

27

Figure 24. Mooring sinkers and chain along the north-western edge of Lamlash Bay (located at East (X): 203236.00 and North (Y): 630883.00).

3.1.13 Target 13 - coarse sediment with macroalgae and maerl

There is a shallow bank extending down from the southern tip of Holy Island. This shallow bank is dominated by coarse sediment (large and small boulders, cobbles and gravel) with macroalgae and maerl gravel (figure 25). The larger algae cover the seabed and prevent the identification of maerl in some areas but maerl is most likely present on the majority of this bank. The image is approximately 20 m wide. Figure 25. An area of mixed coarse sediment at the southern entrance of Lamlash Bay where maerl and macroalgae are present (located at East (X): 206527.00 and North (Y): 628297.00).

Page 40: Mapping of the marine habitats and species of Lamlash Bay, Arran

28

3.1.14 Target 14 – mixed seabed environment off Holy Island

The seabed around target 14 (figure 26) from the area to the north-east of Holy Island consists of mixed sediments. Some areas are dominated by a truly mixed sediment environment with muddy sandy gravel with some shell material but there are also sections of muddy sand with a different faunal community. The image is approximately 20 m wide. Figure 26. An area of mixed backscatter along the north-eastern coast off Holy Island (located at East (X): 205026.00 and North (Y): 631114.00).

3.2 Photographic and video analysis – ground-truthing

3.2.1 Ground-truthing of the sidescan sonar imagery

In 2008 a total of 19 miniDV tapes with approximately 11 hours of video footage were recorded during the drop-down camera survey and a total of 848 seabed still photographs (869 with clapper boards to identify each deployment) were taken at 22 different locations (figure 9 and table 4). The 2009 ground-truthing survey resulted in a total of just over 4 hours of video footage (recorded on DVD) and 278 still photographs (294 with clapper boards to identify each deployment) from 16 different locations (see figure 10 and table 5).

Page 41: Mapping of the marine habitats and species of Lamlash Bay, Arran

29

Table 4. Summary of video and still photography data collected during the 2008 Lamlash Bay surveys (navigation failed on LOC03).

Location Depth (m) Video duration (miniDV) recorded

Number of still photographs

LNC01 30.0 00:22:05 18

LNC02 15.3 00:24:28 27

LNC03 11.3 00:31:18 35

LNC04 24.1 00:30:03 50

LNC05 25.4 00:48:38 54

LNC07 20.9 00:49:02 37

LNC08 27.6 00:31:09 69

LNC09 16.1 00:24:56 21

LNC10 29.4 00:37:22 26

LNC11 14.2 00:40:51 23

LNC12 25.6 00:31:46 58

LSC01 40.1 00:35:05 61

LSC02 32.3 00:39:17 48

LSC03 33.2 00:33:47 37

LSC04 39.4 00:44:18 51

LSC05 38.9 00:20:35 19

LSC06 37.6 00:14:28 31

LSC07 34.1 00:31:05 47

LOC01 30.3 00:45:43 79

LOC02 22.8 00:14:16 19

LOC03 - 00:15:26 34

LOC04 20.2 00:09:13 25

TOTAL 11:14:51 869

Table 5. Summary of video and still photography data collected during the 2009 Lamlash Bay surveys (depth from the first still photography in each transect).

Location Depth (m) Video duration (DVD) recorded

Number of still photographs

NTZ01 3.8 0:11:22 17

NTZ02 5.3 0:12:24 15

NTZ03 9.6 0:18:44 22

NTZ04 17.5 0:15:22 22

NTZ05 5.7 0:17:28 21

A01 8.0 0:25:01 27

A02 17.7 0:14:57 15

A03 11.3 0:08:50 9

B01 23.0 0:12:46 11

D01 45.6 0:11:53 11

D02 47.6 0:11:06 9

D03 12.0 0:13:53 15

E01 10.8 0:21:10 27

E02 13.1 0:10:42 15

F01 2.7 0:15:20 18

H01 36.3 0:38:33 40

TOTAL 04:19:31 294

Page 42: Mapping of the marine habitats and species of Lamlash Bay, Arran

30

The selection of sidescan sonar targets (section 3.1) for illustration purposes was carried out subsequent to processing and analysis of the sidescan sonar data. The selection of ground-truthing using the camera was, however, carried out in the field prior to any detailed analysis of the sidescan sonar data. The selected targets in section 3.1 did therefore not always correspond directly to these targets as some features were not obvious on initial interpretation or there simply were too many desired targets to allow ground-truthing of all of them within the allocated survey period. However, the quality of the sidescan sonar images at most of these targets is sufficient to verify the identity of the features seen.

3.2.1.1 Target 5 - Deacon Rock

The photographic transect (LNC07) near Deacon Rock (figure 27) allows ground-truthing of target 5 (see figure 17) confirming the initial interpretation of a seabed environment with boulders and cobbles surrounded by sand. The species found on Deacon Rock include Ascidiacea, Antedon bifida, Ophiura sp., Munida rugosa, hydroids and various polychaetes. Figure 27. Ground-truthing of target 5 - Deacon Rock (photographs LNC07_26 and 32).

3.2.1.2 Target 7 - low backscatter return

Target 7 has been interpreted as sand and it is a widespread feature across the entire Lamlash Bay survey area. The ground-truthing photographs (example given in figure 28) verify the initial interpretation and illustrate sandy mud and fauna including Ophiura sp, Turritella communis, gobies and burrowing polychaetes. Figure 28. Ground-truthing of target 7 - low backscatter return (photograph LSC03_0020).

Page 43: Mapping of the marine habitats and species of Lamlash Bay, Arran

31

3.2.1.3 Target 8 - mottled high and low backscatter

The mottled high and low backscatter seen in target 8 has been interpreted as gravel, an interpretation confirmed in the photographs, exemplified by the photographs in figure 29. The fauna found in these areas include C. lloydii, various burrowing and tubiculous polychaetes, T. communis, Ascidiacea and the occasional E. esculentus and Liocarcinus puber. Figure 29. Ground-truthing of target 8 (photographs LNC05_48 and LSC04_49).

3.2.1.4 Target 9 - mottled high and low backscatter

The interpretation of target 9 (mottled high back scatter return with relief surrounded by low backscatter) was interpreted as cobbles and small and large boulders, surrounded by sand, features confirmed in the ground-truthing as given in figure 30. Fauna such as brittlestar and maerl beds are found on similar substrata together with hydroids, polychaetes, scallops, Liocarcinus spp. and T. communis as well as some red algae. Figure 30. Ground-truthing of target 9 (photographs LNC10_21 and LNC08_20).

Page 44: Mapping of the marine habitats and species of Lamlash Bay, Arran

32

3.2.1.5 Target 10 - area of seabed with maerl

The ground-truthing of target 10 confirms the initial interpretation of a seabed with strong backscatter returns and mottled appearance to be very coarse sediment (figure 31). Maerl was found at this location, a feature not visible on the sidescan sonar, but this type of seabed environment was shown to be relatively common in the northern and Outer Lamlash regions. Fauna associated with this habitat included hydroids, C. lloydii and polychaetes as well as some Liocarcinus spp. and scallops. Figure 31. Ground-truthing of target 10 (photograph LNC11_10).

3.2.1.6 Target 13 – area of seabed with maerl and seaweed

The shallow bank extending from the southern tip of Holy Island is most likely dominated by maerl and seaweed communities throughout although the ground-truthing using the drop-down camera only acquired footage along two transects (see figures 10 and 12). The video footage and still photographs confirm the presence of coarse sediment (small boulders, cobbles and gravel in particular) but also a seabed dominated by maerl, some of which is hard to identify as a result of the dense cover of large seaweed taxa in places (see figure 32). There is some live maerl present in this region (approximately 1 – 8%) but also patches of brittlestars (Ophiocomina nigra).

Figure 32. Ground-truthing of target 13 (photograph E01_05 and E01_19).

Page 45: Mapping of the marine habitats and species of Lamlash Bay, Arran

33

3.2.1.7 Target 14 – mixed sediment north-east of Holy Island

The sedimentary environment north-west of Holy Island is mixed, as predicted from the sidescan sonar imagery and confirmed by the photographic material. Some areas are dominated by a truly mixed seabed (muddy sandy gravel and shell) with fauna such as C. lloydii, T. communis, hydroids and polychaetes (see NTZ05_09 in figure 33). The other main sediment type (muddy sand) present is dominated by T. communis and Amphiura sp., a community similar to that seen in the centre of the bay (see NTZ05_18 in figure 33). Figure 33. Ground-truthing of target 14 (photograph NTZ05_09 and NTZ05_18).

3.2.2 General descriptions of the biological communities in Lamlash Bay

The analysis of the sidescan sonar mosaic, together with the ground-truthing using the photography (see summary of species in appendix 07) and video (see summary of biotopes in appendix 08) records, allowed for a detailed assessment of the habitats in Lamlash Bay. The interpretation of the mosaic and the ground-truthing revealed a number of features: 1) soft sediments dominated by muddy sand in central and southern regions of Lamlash Bay; 2) a mixture of sediment environments (muddy sand and mixed sediments) in the northern Lamlash Bay and Outer Lamlash Bay area with sections of rocks and brittlestar beds; 3) relatively small rocky reefs (including Deacon Rock); and 4) shallower areas / banks in northern Lamlash Bay, North Channel, Outer Lamlash Bay and the southern entrance to Lamlash Bay with maerl beds. The majority of the photographic / video transects cross a substratum boundary in an attempt to ground-truth more than one type of backscatter return in each deployment in order to increase sampling efficiency. This resulted in more than one habitat as well as different taxa present within each transect, making the descriptive tasks more complicated (see tables 6, 7 and 8).

3.2.2.1 Central and southern Lamlash Bay

The central and southern regions of Lamlash Bay are characterised by soft sediment, mainly muddy sand, and the dominant fauna include Amphiura sp., Ophiura sp., T. communis and polychaetes (figure 34 and table 6). However, the seabed is not uniformly muddy sand and there are several photographic transects that cross several (minor) boundaries both in terms of the substratum and fauna present. Transect LSC02 illustrate this point perfectly with the first section being dominated by Amphiura sp.; the second section being dominated by Ophiura sp. and T. communis; and the third section being dominated by T. communis. The

Page 46: Mapping of the marine habitats and species of Lamlash Bay, Arran

34

sediment appears to be the same in the first two sections but changes to a mixed type (muddy sand and gravel) in the third section. A total number of 5 biotopes have been recognised for this region of the survey area. The dominant types are SS.SSa.CMuSa.(AbraAirr), exemplified by photograph LSC01 (in figure 34) and SS.SMu.CSaMu.(AfilMysAnit), exemplified by photograph LSC02 (in figure 34). Both of these biotopes have been somewhat tentatively selected (hence the brackets) in the photographic analysis. The latter of the two has been selected rather than SS.SSa.CMuSa.AalbNuc (see Day grab samples below) as T. communis is common on the photographs at AfilMysAnit locations but it is not a taxa recorded in AalbNuc. These two biotopes appear to be very closely linked and grade into each other (Connor et al., 2004). The biotopes resulting from the Day grab and photographic survey may therefore be: 1) a combination of the two biotopes; or 2) different patches of the two biotopes in the same region. The difference between AbraAirr and AalbNuc is subtle but obvious on the photographs in the large number of Amphiura sp. present in the former. Furthermore, the sediment is slightly sandier in AbraAirr compared to AalbNuc. Table 6. Summary of the video and photographic analyses with the main substrata, fauna and biotope in central and southern Lamlash Bay.

Location Main substrata Main fauna Biotope(s)

LSC01 Muddy sand with shell

Amphiura sp. Polychaetes

SS.SSa.CMuSa.(AbraAirr) SS.SMx.CMx

CR.LCR.BrAs.AmenCio.Bri

LSC02 Muddy sand with shell

Amphiura sp. Ophiura sp.

Turritella communis Polychaetes

SS.SSa.CMuSa.(AbraAirr) SS.SMu.CSaMu.(AfilMysAnit)

SS.SMx.CMx

LSC03 Muddy sand Ophiura sp. Turritella communis

Polychaetes

SS.SSa.CMuSa.(AbraAirr) SS.SMu.CSaMu.(AfilMysAnit)

LSC04 Muddy sand with shell

Amphiura sp. Polychaetes

SS.SSa.CMuSa.(AbraAirr) SS.SMx.CMx

CR.LCR.BrAs.AmenCio

LSC05 Muddy sand Amphiura sp. Polychaetes

SS.SSa.CMuSa.(AbraAirr)

LSC06 Muddy sand Amphiura sp. Polychaetes

SS.SSa.CMuSa.(AbraAirr)

LSC07 Muddy sand Ophiura sp. Turritella communis

Polychaetes

SS.SMu.CSaMu.(AfilMysAnit) CR.LCR.BrAs.AmenCio

B01 Muddy sand Ophiura sp. Turritella communis

Polychaetes

SS.SSa.CMuSa.(AbraAirr)

F01 Muddy sandy gravel with some shell material

Brown / red algae Cerianthus lloydii

Polychaetes

SS.SMP.KSwSS.LsacR SS.SMx.CMx

H01 Mixed sediments Polychaetes Hydroidea spp. Antedon bifida

SS.SMx.CMx CR.LCR.BrAs.AmenCio

Figure 34. Photographs from the southern and central regions of Lamlash Bay (LSC01_07, LSC02_29, LSC06_26 and LSC07_39).

Page 47: Mapping of the marine habitats and species of Lamlash Bay, Arran

35

3.2.2.2 Northern Lamlash Bay and North Channel

The northern Lamlash Bay and North Channel area consists of a very mixed seabed environment with substrata ranging from muddy sand to bedrock (figures 35 and 36 and table 7). The fauna is also different in different part of the area including T. communis, M. rugosa, C. lloydii, polychaetes, ascidians and maerl (figures 36 and table 7). A total of 6 biotopes have been classified for this area. The dominant biotopes are SS.SMu.CSaMu.(AfilMysAnit) and SS.SMx.CMx.ClloMx(Nem) but the maerl biotope (SS.SMP.Mrl.Pcal) is common in the shallower regions of the survey area and extends further into the infralittoral region (see figures 10 and 35). The maerl is dominant on the main (deeper) part of the banks but there is a mixture of maerl and macroalgae (brown and red seaweeds) in the shallowest areas. The circalittoral rock biotope CR.LCR.BrAs.AmenCio (exemplified by LNC10_06 and LNC11_20 in figure 36) has been selected rather than CR.LCR.BrAs.AntAsH as the latter is found in the most sheltered positions in sea lochs, in weak to very weak tidal currents and the fauna is slightly different to the AmenCio biotope. SS.SMu.CSaMu.(AfilMysAnit), exemplified by LNC05, LNC07 and LNC12 in figure 36, has been selected in the photographic analysis rather than SS.SSa.CMuSa.AalbNuc (see Day grab samples below) as T. communis is common on the photographs but it is not a taxa recorded in AalbNuc. These two biotopes appear to be very closely linked and grade into each other (Connor et al., 2004). The biotopes found here may therefore be: 1) a combination of the two; or 2) different patches of the two biotopes in the same region. The

Page 48: Mapping of the marine habitats and species of Lamlash Bay, Arran

36

classification of biotopes during the video analysis allowed classification only in lower levels in the hierarchy as less detail is visible in the videos compared to the still photographs (allowing zooming in). Table 7. Summary of the video and photographic analyses with the main substrata, fauna and biotopes (mixed sediments are muddy sandy gravel with shell material) in northern Lamlash Bay and North Channel.

Location Main substrata Main fauna Biotope(s)

A01 Mixed sediments Brown / red algae Maerl

Polychaetes Cerianthus lloydii

SS.SMP.Mrl.Pcal

A02 Mixed sediments Brown / red algae Maerl

Polychaetes Cerianthus lloydii

SS.SMP.Mrl.Pcal

A03 Mixed sediments Brown / red algae Maerl

Polychaetes Cerianthus lloydii

SS.SMP.Mrl.Pcal SS.SMP.KSwSS.LsacR

LNC02 Muddy sand and gravel with shell

material

Maerl Polychaetes

Cerianthus lloydii Ascidiacea

SS.SMP.Mrl.Pcal

LNC03 Muddy sand and gravel with shell

material

Maerl Polychaetes

Cerianthus lloydii Ascidiacea

SS.SMP.Mrl.Pcal SS.SMu.CSaMu.(AfilMysAnit)

LNC04 Muddy sand and mixed sediments

Turritella communis Polychaetes

SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem)

LNC05 Muddy sand and mixed sediments

Turritella communis Polychaetes

SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem) CR.LCR.BrAs.AmenCio

LNC07 Muddy sand and mixed sediments

with rocks

Turritella communis Polychaetes Ophiura sp.

SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem) CR.LCR.BrAs.AmenCio

LNC08 Muddy sand and mixed sediments

with rocks

Ophiura sp. Ophiocomina nigra

Ascidiacea Turritella communis

Polychaetes

SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem) CR.LCR.BrAs.AmenCio

CR.LCR.BrAs.AmenCio.Bri

LNC09 Muddy sandy gravel with shell

material

Maerl Polychaetes

Cerianthus lloydii Turritella communis

SS.SMP.Mrl.Pcal SS.SMu.CSaMu.(AfilMysAnit)

CR.LCR.BrAs.AmenCio

LNC10 Mixed sediments Turritella communis Munida rugosa Polychaetes

SS.SMx.CMx.ClloMx(Nem) SS.SMu.CSaMu.(AfilMysAnit)

CR.LCR.BrAs.AmenCio CR.LCR.BrAs.AmenCio.Bri

LNC11 Mixed sediments Maerl Polychaetes

SS.SMP.Mrl.Pcal SS.SMu.CSaMu.(AfilMysAnit)

Page 49: Mapping of the marine habitats and species of Lamlash Bay, Arran

37

Turritella communis Ascidiacea

c CR.LCR.BrAs.AmenCio

LNC12 Muddy sand and mixed sediment

Polychaetes Turritella communis

Ophiura sp.

SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem)

NTZ01 Sandy gravel Brown / red algae Maerl

SS.SMP.KSwSS.LsacR SS.SMP.Mrl.Pcal

NTZ02 Mixed sediments Maerl Polychaetes

Cerianthus lloydii

SS.SMP.Mrl.Pcal

NTZ03 Sandy gravel and mixed sediments

Brown / red algae Maerl

Polychaetes

SS.SMP.KSwSS.LsacR SS.SMP.Mrl.Pcal

NTZ04 Sandy gravel and mixed sediments

Brown / red algae Maerl

Polychaetes

SS.SMP.KSwSS.LsacR SS.SMP.Mrl.Pcal

NTZ05 Mixed sediments Polychaetes Cerianthus lloydii Turritella communis

Amphiura sp.

SS.SMx.CMx.ClloMx(Nem) SS.SSa.CMuSa.(AbraAirr)

Figure 35. Photographs from transect A in the northern part of the North Channel area of Lamlash Bay (A01_06, and A02_11).

Page 50: Mapping of the marine habitats and species of Lamlash Bay, Arran

38

Figure 36. Photographs from the northern Lamlash Bay and North Channel areas of Lamlash Bay (LNC03_09, LNC05_04, LNC07_35, LNC10_02, LNC10_06, LNC11_06, LNC11_20, LNC12_03).

Page 51: Mapping of the marine habitats and species of Lamlash Bay, Arran

39

3.2.2.3 Outer Lamlash Bay

The Outer Lamlash Bay area consists of a very mixed seabed environment with sections of muddy sand and other sections of mixed and coarse (with gravel, cobbles and boulders) substrata (see figure 37 and table 8). Two transects (LNC01* and LOC02) are dominated by muddy sand and taxa dominated by T. communis, M. rugosa and polychaetes. The other three transects are characterised by a mixed sediment (muddy sand and gravel with rocks in some areas) and with a varied faunal composition (maerl, featherstars, T. communis, O. nigra, C. lloydii and polychaetes). A total of 6 different biotopes have been classified in the Outer Lamlash Bay region (see comments about the selection of the various biotopes above). Photographic examples of the biotopes are given as follows: LOC03 exemplifies the SS.SMx.CMx biotope complex, photograph LNC01_56 exemplifies the CR.LCR.BrAs.AmenCio.Bri (see further examples below) sub-biotope and photograph LNC03_04 exemplifies the SS.SMx.CMx.ClloMx(Nem) biotope (all given in figure 37). SS.SSa.CMuSa.(AbraAirr) has been identified in the deeper areas to the east of Holy Island (D01 and D02). The sediment and the fauna are very similar but the abundance of Amphiura sp. is slightly lower than in the central parts of Lamlash Bay. The seabed environment appears to be very mixed in this region and none of the biotopes seems to be particularly dominant relative to others. Maerl is present, particularly on the shallower banks, but there is a big difference in the quality of the maerl with the maerl at LOC01 being relatively healthy whilst the maerl at LOC04 is difficult to detect at all. Table 8. Summary of the video and photographic analyses with the main substrata, fauna and biotopes (mixed sediments are muddy sandy gravel with shell material) in Outer Lamlash Bay (* LNC01 has been placed with Outer Lamlash Bay in the general description stage).

Location Main substrata Main fauna Biotope(s)

*LNC01 Muddy sand Turritella communis Munida rugosa Polychaetes

SS.SMu.CSaMu.(AfilMysAnit)

LOC01 Mixed sediments Maerl Turritella communis Antedon bifida

SS.SMP.Mrl.Pcal SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem) CR.LCR.BrAs.AmenCio

LOC02 Muddy sand Turritella communis Polychaetes

SS.SMu.CSaMu.(AfilMysAnit)

LOC03 Muddy sand and gravel

Maerl Antedon bifida

Ophiocomina nigra Ascidiacea Polychaetes

SS.SMP.Mrl.Pcal SS.SMu.CSaMu.(AfilMysAnit)

SS.SMx.CMx CR.LCR.BrAs.AmenCio

CR.LCR.BrAs.AmenCio.Bri

LOC04 Muddy sand and gravel

Maerl Polychaetes

Cerianthus lloydii

SS.SMP.Mrl.Pcal SS.SMu.CSaMu.(AfilMysAnit) SS.SMx.CMx.ClloMx(Nem)

D01 Muddy sand Amphiura sp. Turritella communis

Polychaetes

SS.SSa.CMuSa.(AbraAirr)

D02 Muddy sand Amphiura sp. Turritella communis

Polychaetes

SS.SSa.CMuSa.(AbraAirr)

D03 Mixed sediments Maerl Polychaetes

Cerianthus lloydii

SS.SMP.Mrl.Pcal

Page 52: Mapping of the marine habitats and species of Lamlash Bay, Arran

40

Figure 37. Photographs from the Outer Lamlash Bay area (LNC01_09, LNC02_17, LOC01_35, LNC01_56, LNC01_71, LNC03_04, D01_02, D03_13).

Page 53: Mapping of the marine habitats and species of Lamlash Bay, Arran

41

3.2.2.4 Southern entrance to Lamlash Bay

A shallow bank extends from the southern tip of Holy Island. This bank is dominated by maerl (P. calcareum) but brown and red seaweeds (macroalgae) are present in the shallowest sections obscuring the seabed making identification and quantification of maerl difficult (figure 38 and table 9) in some sections. In addition to maerl there are also patches of brittlestar beds. These patches are dominated by O. nigra have been classified as SS.SMx.CMx.OphMx. Some sections contain both maerl and brittlestars but as maerl beds are recognised as important habitats these areas have been classified as SS.SMP.Mrl.Pcal. The deeper area immediately west of the bank appears to be characterised by mixed sediment but potentially also muddy sands. However, as there are no ground-truthing data available from this section further survey work is required to assess this fully. Table 9. Summary of the video and photographic analyses with the main substrata, fauna and biotopes (mixed sediments are muddy sandy gravel with shell material) at the southern entrance to Lamlash Bay.

Location Main substrata Main fauna Biotope(s)

E01 Mixed sediments Brown / red algae Maerl

Polychaetes Cerianthus lloydii Ophiocomina nigra

SS.SMP.KSwSS.LsacR SS.SMP.Mrl.Pcal

SS.SMX.CMx.OphMx CR.LCR.BrAs.AmenCio.Bri

E02 Mixed sediments Maerl Polychaetes

Cerianthus lloydii Ophiocomina nigra

SS.SMP.Mrl.Pcal SS.SMx.CMx.ClloMx(Nem)

SS.SMX.CMx.OphMx

Figure 38. Photographs from the southern entrance to Lamlash Bay (E01_07, E01_11, E01_26, E02_13).

Page 54: Mapping of the marine habitats and species of Lamlash Bay, Arran

42

3.2.2.5 Scallops and other noteworthy species

The analysis of the photographic material resulted in identification of scallops (Pecten maximus and Aequipecten opercularis) and a few other noteworthy species (see appendix 09 with a summary in tables 10 and 11; and figure 39). M. rugosa is the most abundant taxon in the examples given in table 10 with the highest densities within the CR.LCR.BrAs.AmenCio biotope in 2008 and SS.SSa.CMuSa.(AbraAirr) in 2009. Only a few individuals of C. pagurus and Nephrops norvegicus were identified in the still photographs over the two years whilst Homarus gammarus was not identified at all. In 2008 scallops were found in all of the survey areas in Lamlash Bay (table 10 and appendix 09) in 2008 with P. maximus being, overall, the most common scallop taxon (in the still photography analysis), with the highest densities found in the SS.SMx.CMx.ClloMx(Nem), SS.SMP.Mrl.Pcal and CR.LCR.BrAs.AmenCio biotopes. The highest densities of A. opercularis in 2008 were found in the SS.SMx.CMx.ClloMx(Nem) biotope. The densities of P. maximus and A. opercularis appear to be very similar within the NTZ, with densities of both species being around 3 individuals/100 m2. Table 10. The approximate densities (ind/100 m2) of scallops and two other species present in the still photographs from the 2008 Lamlash Bay survey (total: total number of individuals/100m2; total (N): total number of individuals; NTZ: individuals/100m2 in the NTZ area; LNC: total number of individuals/100m2 in northern Lamlash Bay; LOC: total number of individuals/100m2 in Outer Lamlash Bay; LSC: total number of individuals/100m2 in southern and central Lamlash Bay; Pectinidae sp. are unidentified individuals i.e. not the total of all scallops).

Still photography analysis

Munida rugosa

Cancer pagurus

Pectinidae sp.

Pecten maximus

Aequipecten opercularis

Total 9.4 0.5 1.5 4.4 2.0

Total (N) 80 4 13 37 17

LNC 8.8 0.5 1.2 4.7 2.0

LOC 16.3 0.0 2.0 2.6 0.7

All

LSC 6.6 0.7 1.7 4.9 2.8

NTZ 6.6 0.7 1.8 3.3 2.6 NTZ

NTZ (N) 18 2 5 9 7

SaMu Total 8.3 0.4 0.4 1.5 1.5

Total (N) 22 1 1 4 4

CMx Total 10.6 0.5 1.0 7.0 3.0

Total (N) 21 1 2 14 6

Mrl.Pcal Total 1.7 0.8 5.0 7.6 0.0

Total (N) 2 1 6 9 0

AmenCio Total 27.5 0.0 1.4 10.1 0.0

AmenCio Total (N) 19 0 1 7 0

MuSa Total 0.0 8.9 0.6 1.7 1.7

Total (N) 0 15 0 3 3

SaMu SS.SMu.CSaMu.(AfilMysAnit) CMx SS.SMx.CMx.ClloMx(Nem) Mrl.Pcal SS.SMP.Mrl.Pcal AmenCio CR.LCR.BrAs.AmenCio AmenCio CR.LCR.BrAs.AmenCio.Bri MuSa SS.SSa.CMuSa.(AbraAirr)

Page 55: Mapping of the marine habitats and species of Lamlash Bay, Arran

43

In 2009 scallops were identified in the No Take Zone as well as within the SS.SMx.CMx.ClloMx(Nem) and SS.SMPMrl.Pcal biotopes (table 11). The 2009 densities of P. maximus are higher than those of A. opercularis within the NTZ; with densities of P. maximus of approximately 1.5 individuals/100 m2 whilst A. opercularis was not identified at all within this area (N.B. observing and identifying some species on camera footage is notoriously difficult (see below)). Table 11. The approximate densities (ind/100 m2) of scallops and two other species present in the still photographs from the 2009 Lamlash Bay survey (total: total number of individuals/100m2; total (N): total number of individuals; Pectinidae sp. are unidentified individuals i.e. not the total of all scallops).

Still photography analysis

Munida rugosa

Cancer pagurus

Pectinidae sp.

Pecten maximus

Aequipecten opercularis

Total 3.3 0.5 1.1 2.7 0.5 All Total (N) 6 1 2 5 1

NTZ 0.0 0.0 0.7 1.5 0.0 NTZ NTZ (N) 0 0 1 2 0

CMx Total 5.8 0.0 1.9 3.8 1.9

Total (N) 3 0 1 2 1

Mrl.Pcal Total 0.0 0.8 0.8 2.3 0.0

Total (N) 0 1 1 3 0

AmenCio Total 0.0 0.0 0.0 0.0 0.0

AmenCio Total (N) 0 0 0 0 0

MuSa Total 8.1 0.0 0.0 0.0 0.0

Total (N) 3 0 0 0 0

OphMx Total 0.0 0.0 0.0 0.0 0.0

Total (N) 0 0 0 0 0

LsacR Total 0.0 0.0 0.0 0.0 0.0

LsacR Total (N) 0 0 0 0 0

AmenCio CR.LCR.BrAs.AmenCio

AmenCio CR.LCR.BrAs.AmenCio.Bri

CMx SS.SMx.CMx.ClloMx(Nem)

OphMx SS.SMX.CMx.OphMx

Mrl.Pcal SS.SMPMrl.Pcal

MuSa SS.SSa.CMuSa.(AbraAirr)

LsacR SS.SMP.KSwSS.LsacR.CbPb

LsacR SS.SMP.KSwSS.LsacR.Sa

The 2008 video analysis rarely allowed identification of the scallops to species level. This group was therefore recorded within the family Pectinidae (table 12). In 2008 the highest densities of scallops were found in the CR.LCR.BrAs.AmenCio and SS.SMP.Mrl.Pcal biotopes with overall densities of scallops being approximately 2 individuals per 100 m2. In 2009 the density values are slightly lower overall (c. 1 individual per 100 m2) with the highest densities found in SS.SMx.CMx and CR.LCR.BrAs.AmenCio biotopes (table 13 and appendix 9). The video density values are lower than the values seen in the still photography analysis, a result most likely linked to the fact that the identification (observe and count) of fauna is harder on the video compared to the photographs (some scallops were difficult to see even on still photographs; see figure 39). It should be noted that the biotope / habitat classification of individual photographs involves a subjective selection process and it is possible that some of the photographs or areas are classified incorrectly but the densities given in tables 10 to 13 give estimates of the densities of these taxa.

Page 56: Mapping of the marine habitats and species of Lamlash Bay, Arran

44

Table 12. The approximate densities (ind/100 m2) of scallops and two other species present in the video footage from the 2008 Lamlash Bay survey (total: total number of individuals/100m2; total (N): total number of individuals; LNC: total number of individuals/100m2 in northern Lamlash Bay; LOC: total number of individuals/100m2 in Outer Lamlash Bay; LSC: total number of individuals/100m2 in southern and central Lamlash Bay; Pectinidae sp. are the total number of scallop individuals as identification of scallops species is difficult on video footage. * MuSa not distinguished during the video analysis hence no data in these fields).

Video footage analysis 2008

Pectinidae sp.

Cancer pagurus

Nephrops norvegicus

Pleuronectidae sp.

Total 2.3 0.4 0.1 0.0

Total (N) 342 61 9 2

LNC 2.7 0.3 0.0 0.0

LOC 2.0 0.7 0.2 0.0

All

LSC 1.9 0.2 0.0 0.0

SaMu Total 2.3 0.5 0.1 0.0

CMx Total 1.8 0.5 0.0 0.0

Mrl.Pcal Total 2.5 0.1 0.0 0.0

AmenCio

AmenCio Total 4.4 0.1 0.0 0

MuSa Total * * * *

SaMu SS.SMu.CSaMu.(AfilMysAnit)

CMx SS.SMx.CMx.ClloMx(Nem)

Mrl.Pcal SS.SMP.Mrl.Pcal

AmenCio CR.LCR.BrAs.AmenCio

AmenCio CR.LCR.BrAs.AmenCio.Bri

MuSa SS.SSa.CMuSa.(AbraAirr)

Table 13. The approximate densities (ind/100 m2) of scallops and two other species present in the video footage from the 2009 Lamlash Bay survey (with the approximate densities in the different survey areas and biotope; total: total number of individuals/100m2; total (N): total number of individuals; Pectinidae sp. are the total number of scallop individuals as identification of scallops species is difficult on video footage).

Video footage analysis 2009 Cancer pagurus

Nephrops norvegicus

Munida rugosa

Pectinidae sp. Pecten maximus

Total 0.3 0.3 3.0 0.1 1.1 All

Total (N) 7 9 82 2 31

NTZ Total 0.2 0.0 0.0 0.1 0.5

CMx Total 0.0 0.0 4.4 0.2 2.4

Mrl.Pcal Total 0.6 0.0 0.0 0.0 0.6

AmenCio

AmenCio Total 1.3 0.0 1.3 0.0 6.3

MuSa Total 0.0 2.8 18.2 0.3 0.9

OphMx Total 0.0 0.0 0.0 0.0 2.6

LsacR

LsacR Total 0.0 0.0 0.0 0.0 0.0

AmenCio CR.LCR.BrAs.AmenCio AmenCio CR.LCR.BrAs.AmenCio.Bri

CMx SS.SMx.CMx.ClloMx(Nem) OphMx SS.SMX.CMx.OphMx

Mrl.Pcal SS.SMPMrl.Pcal MuSa SS.SSa.CMuSa.(AbraAirr)

LsacR SS.SMP.KSwSS.LsacR.CbPb LsacR SS.SMP.KSwSS.LsacR.Sa

Page 57: Mapping of the marine habitats and species of Lamlash Bay, Arran

45

Figure 39. Examples of scallops observed in Lamlash Bay (LNC08_37, LNC03_31, LNC08_22 and LNC03_18).

3.2.2.6 Maerl beds

As noted above, maerl beds (SS.SMP.Mrl.Pcal) were identified in the northern Lamlash Bay, in North Channel and in the Outer Lamlash Bay as well as around several regions of Holy Island (figure 40). These beds are typically found in the shallower areas but extend down to approximately ≤26 m. The maerl beds are found at NTZ01 - NTZ04, A01 - A03, D03, E01-E02, LNC02, LNC03, LNC09, LNC11, LOC01, LOC03 and LOC04 (see figure 40) but the vast majority of these beds are in a poor condition. The maerl beds at A01, A02 and LOC01 are the healthiest populations with the largest proportion (5-10 %) of live maerl. LOC04 is an area with some maerl in particularly poor health with very few scattered individuals present (all dead). In fact, in many cases it was difficult to identify the maerl without zooming in on the images and maerl was not detected during the video analysis, illustrating the need for both still photography and video footage to ensure recording large-scale and small-scale features. LOC04 can therefore not be classified as a maerl bed and this location is best described as an area of scattered maerl. Other regions with maerl are mixed with macroalgae (covering the seabed) making identification of the maerl challenging. The assessments of the condition of these beds are therefore more difficult and uncertain to some extent.

Page 58: Mapping of the marine habitats and species of Lamlash Bay, Arran

46

Figure 40. Maerl beds observed in and around Lamlash Bay (LNC02_04, LNC03_02, LNC0902, LNC11_02, LOC01_03, LOC03_19).

3.2.2.7 Brittlestar beds

Patches of seabed with brittlestars (CR.LCR.BrAs.AmenCio.Bri) were observed at several locations (E01, E02, LNC08, LNC10, LOC03, and LSC01) of the Lamlash Bay survey (figures 38 and 41). The dominant taxon was O. nigra. The other brittlestar taxon Ophiothrix fragilis, which was often associated with O. nigra elsewhere, was not seen in these samples.

Page 59: Mapping of the marine habitats and species of Lamlash Bay, Arran

47

Figure 41. Brittlestar beds in Lamlash Bay (LNC08_19, LNC08_20, LOC3_ 08 and LSC01_55).

3.3 Sediment samples A total of 10 Day grab samples were collected within the NTZ in Lamlash Bay (table 14 and appendix 03). A sub-sample was collected for Particle Size Analysis and the remaining sample was retained for further macrofaunal processing and analysis. Table 14. Day grab sampling locations (navigation data in ddmmss.sss).

Location Date Latitude (N) Longitude (W) Depth (m)

NTZ01#01 20/10/2008 553239.274 -50547.237 27.5

NTZ02#02 20/10/2008 553228.927 -50567.708 28.6

NTZ03#01 20/10/2008 553218.679 -50591.838 30.0

NTZ04#01 20/10/2008 553224.860 -50608.735 26.9

NTZ05#01 20/10/2008 553208.646 -50581.213 22.1

NTZ06#03 20/10/2008 553198.133 -50567.949 15.6

NTZ07#01 20/10/2008 553225.128 -50538.758 20.4

NTZ08#02 20/10/2008 553241.085 -50507.473 19.7

NTZ09#01 20/10/2008 553254.641 -50562.384 20.3

NTZ10#01 20/10/2008 553251.697 -50467.579 22.5

Page 60: Mapping of the marine habitats and species of Lamlash Bay, Arran

48

3.3.1 Particle Size Analysis

The results from the particle size analysis from the Day grab samples are given in appendix 04 with a summary in table 15 (mud, sand and gravel refer to all size fractions within each category). The results illustrate that all the sites are dominated by “sand” with at least a proportion of mud being present at all the locations. The highest percentage of mud (>10 %) is found at NTZ02, NTZ03 and NTZ04. Gravel is also present but not at all the locations. NTZ08 and NTZ10 have the highest percentage of gravel but smaller percentage is also present at NTZ07 and NTZ09 (table 15). Overall, however, and the most important aspect of these results is that sand represents more than 80 % of the sediment at all but one of the 10 sites (see table 15). These 9 locations can therefore classified as “Sand” (greater than 80% sand; see appendix 05) according to the modified Folk triangle (Folk, 1954) sediment classification system as adapted by the Joint Nature Conservation Committee (JNCC; see Connor et al., 2004) with NTZ08 also dominated by sand (77 %). In terms of the sediment present, 9 sites could therefore also classify as SS.SSA (sublittoral sediment, sublittoral sands and muddy sands) but as both the sediment type and fauna have to be considered when classifying a site as a particular biotope according to the JNCC Marine Nature Conservation Review Biotope Classification (Connor et al., 2004) system, some of the sites are classified as different biotopes (see below). Table 15. Total number of individuals of the most abundant taxa/species present in the 2008 Lamlash Bay grab samples.

Location Depth (m)

Gravel (%)

Sand (%)

Mud (%)

Classification (Wentworth scale)

NTZ01#01 27.5 0.04 94.76 5.44 muddy sand

NTZ02#02 28.6 0.03 90.24 10.00 muddy sand NTZ03#01 30.0 0.04 88.85 11.29 muddy sand NTZ04#01 26.9 0.06 87.61 12.10 muddy sand NTZ05#01 22.1 0.27 90.52 9.35 muddy sand NTZ06#03 15.6 0.14 93.78 6.17 muddy sand NTZ07#01 20.4 5.33 88.45 6.36 gravely muddy Sand NTZ08#02 19.7 15.77 77.20 7.09 muddy gravely Sand

NTZ09#01 20.3 1.49 89.68 8.86 gravely muddy sand

NTZ10#01 22.5 11.93 81.86 6.32 muddy gravely Sand

The “sand” size class can be further divided according to the sediment size classification based on the Wentworth scale (see e.g. Leeder, 1982). Further analysis of the data shows that within the ‘sand’ category, all the sites are dominated by fine or very fine sand (figure 42 and table 16). Relatively higher proportions of medium and coarse sand, as well as the gravel fractions noted above, are found at three locations (NTZ07, NTZ08 and NTZ10).

Page 61: Mapping of the marine habitats and species of Lamlash Bay, Arran

49

Table 16. Total percentage sediment weight (brown: ‘gravel’; yellow: ‘sand’; light yellow: ‘silt’) of the Day grab samples at the NTZ locations in Lamlash Bay (NB. processing differential resulting in total percentage being in excess of / less than 100%). Sieve size

NTZ01 NTZ02 NTZ03 NTZ04 NTZ05 NTZ06 NTZ07 NTZ08 NTZ09 NTZ10

16.000 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

8.000 0.00 0.00 0.00 0.00 0.20 0.00 0.29 5.13 1.49 4.39

4.000 0.00 0.00 0.02 0.00 0.00 0.00 1.07 3.45 0.00 3.92

2.000 0.04 0.03 0.02 0.06 0.07 0.14 3.97 7.20 0.00 3.63

1.000 0.38 0.18 0.22 0.19 0.57 0.72 9.15 7.36 0.09 4.96

0.500 1.55 0.65 0.75 0.95 1.32 2.31 6.77 5.84 1.28 6.19

0.250 7.08 3.49 3.55 4.28 5.77 9.72 19.06 13.03 7.28 17.79

0.125 53.04 37.14 26.68 16.85 41.50 50.42 42.63 38.11 38.91 39.31

0.063 32.71 48.78 57.66 65.34 41.36 30.62 10.84 12.86 42.12 13.60

< 0.063 5.44 10.00 11.29 12.10 9.35 6.17 6.36 7.09 8.86 6.32

Figure 42. Cumulative percentage sediment weight at the NTZ locations in Lamlash Bay.

3.3.2 Macrofaunal distributions

The macrofaunal analysis revealed a total of 2726 individuals and 160 taxa (appendix 06) in the Lamlash Bay Day grab samples (including meiofauna and epi-fauna). Overall the macrofauna is dominated by Mollusca (45.0%) and Annelida (30.7%), representing 75.7% of the identified fauna. The Echinodermata and Crustacea comprise 16.0% and 6.3% of the macrofauna respectively while the Nemertea represent 1.0% and the remaining groups (Cnidaria, Sipuncula and Ascidiacea) represent 0.08% of the fauna. One individual of Gobiidae sp. was also found in one of the Day grab samples.

Page 62: Mapping of the marine habitats and species of Lamlash Bay, Arran

50

3.3.2.1 Abundance

The abundance of the identified macrofauna collected in the NTZ in Lamlash Bay Day grab survey is given in appendix 06 with some examples of the most abundant species / taxa given in table 17. As with the overall data, Mollusca and Annelida dominated the macrofaunal community. The most abundant taxon is Mysella bidentata, followed by Amphiura filiformis, Prionospio fallax and Thyasira flexuosa. M. bidentata is typically found in muddy sand and fine gravel (sometimes in burrows of sipunculids or associated with ophiuroids) while the suspension feeder A. filiformis is found in muddy sand (Hayward and Ryland, 1991). T. flexuosa is typically found in sandy muds (Tebble, 1966). The ecology of the dominant fauna therefore seems to correlate with the dominant sediment types in Lamlash Bay (silty sand) and although little information is available for the ecology of P. fallax, the current data suggest that P. fallax is also found in muddy sand (most abundant in locations with the highest percentage of silty sand: see appendix 06). Table 17. Total number of individuals of the most abundant taxa/species present in the 2008 Lamlash Bay grab samples (rank order).

MCS code

Taxon Species Abundance

W1906 Mysella bidentata 940

ZB154 Amphiura filiformis 370

P765 Prionospio fallax 172

W1837 Thyasira flexuosa 111

P94 Pholoe synophthalmica 66

P1098 Owenia fusiformis 66

W2128 Dosinia lupinus 42

P494 Nephtys spp. 35

P499 Nephtys hombergii 35

P579 Lumbrineris gracilis 30

P919 Mediomastus fragilis 30

ZB121 Ophiuroidea sp. 29

P502 Nephtys kersivalensis 28

P1178 Trichobranchus roseus 28

S440 Ampelisca tenuicornis 28

W1569 Nucula nitidosa 28

P1124 Melinna palmata 25

S440 Tanaopsis graciloides 24

G1 NEMERTEA spp. 23

S248 Urothoe elegans 23

P1139 Ampharete lindstroemi 22

ZB223 Echinocardium cordatum 22

Page 63: Mapping of the marine habitats and species of Lamlash Bay, Arran

51

3.3.2.2 Diversity

The results from the species diversity analysis are given in table 18. The total number of individuals present at the 10 Lamlash Bay Day grab samples range from 133 individuals per sample to 474 individuals per sample whilst the total number of taxa range from 37 to 64 among the samples, indicating that there are some differences between the different locations. The species diversity (Shannon-Wiener diversity index) overall is low (to medium) with diversity being highest at locations NTZ 05 to NTZ 10. The equitability (J) results suggest a similar pattern where locations NTZ 01 to NTZ 04 have low equitability, indicating higher dominance by a small number of different species, while the results from the other locations (NTZ 05 to NTZ 10) indicate a more equal distribution between species. Table 18. Total number of individuals (N), number of species (S), Margalef’s species richness (d), Pielou’s equitability index (J) and Shannon-Wiener diversity index (H’) for all the samples in the 2008 Lamlash Bay Day grab survey.

Location S N d J H' (loge)

NTZ 01#01 49 474 7.79 0.51 1.98

NTZ 02#02 53 369 8.80 0.54 2.14

NTZ 03#01 37 425 5.95 0.52 1.87

NTZ 04#01 40 304 6.82 0.63 2.33

NTZ 05#01 63 240 11.31 0.80 3.31

NTZ 06#03 47 183 8.83 0.81 3.12

NTZ 07#01 55 162 10.61 0.91 3.64

NTZ 08#02 64 159 12.43 0.94 3.91

NTZ 09#01 43 267 7.52 0.73 2.73

NTZ 10#01 46 133 9.20 0.87 3.33

3.3.2.3 Macrofaunal composition

The results from the cluster analysis and ordination of all the Lamlash Bay infaunal data are given in figures 43 and 44. The cluster analysis group locations NTZ07, NTZ08 and NTZ10 in one cluster whilst the remaining locations are grouped together in a separate cluster (figure 43), results that appear to closely follow the sediment distributions described above (section 3.3.1) where locations NTZ07, NTZ08 and NTZ10 are found in the slightly coarser sediment while the remaining samples are found in ‘fine’ sediments.

Page 64: Mapping of the marine habitats and species of Lamlash Bay, Arran

52

Figure 43. Cluster analysis of the 2008 Lamlash Bay macrofaunal Day grab data.

The ordination analysis support the cluster analysis results (figure 44) grouping NTZ07, NTZ08 and NTZ10 together (coarser sediment group) whilst the remaining seven locations are found in the ‘fine’ sediments. The ordination furthermore reveals a very low level of stress (0.01) giving “an excellent representation with no prospect of misinterpretation” (Clarke and Warwick, 1994). The data were analysed using a number of different variables but the results were unchanged further strengthening the results given here. Figure 44. Ordination analysis of the 2008 Lamlash Bay macrofaunal Day grab data (all 10 locations).

Page 65: Mapping of the marine habitats and species of Lamlash Bay, Arran

53

The ordination results given in figure 44 illustrate the seven ‘fine’ sediment locations grouped very closely together. To assess any potential patterns further a second MDS plot of the remaining seven locations was produced (figure 45). This second ordination analysis largely supports the results in the cluster analysis, grouping NTZ05, NTZ06 and NTZ09 in one group and NTZ01 to NTZ04 together in the other, suggesting some small differences between the two faunal communities. The level of stress of the ordination is slightly higher (0.06) in this analysis but still “corresponds to a good ordination with no real prospect of a misleading interpretation” (Clarke and Warwick, 1994). Figure 45. Further ordination analysis of the 2008 Lamlash Bay macrofaunal Day grab data (the 7 locations grouped).

To assess these groups further, a SIMPER analysis of the 2008 macrofaunal Day grab samples in Lamlash Bay was carried out to assess the most dominant species in the ‘coarse’ (NTZ07, NTZ08 and NTZ10) and ‘fine’ sediment groups as identified in the cluster and ordination analyses (table 19).

Many of the dominant fauna in the ‘fine’ sediment group are typically found in muddy sands (and sandy muds) typified by M. bidentata, A. filiformis and T. flexuosa (see above; Tebble, 1966 and Hayward and Ryland, 1995). The relative contribution of M. bidentata to this community is very high compared to any of the other species. The most characterising fauna in the ‘coarse’ sediment cluster are typically found on substrata of a mixed character, preferences supporting the overall clustering. Dosinia lupinus is a suspension feeder found on sandy mud, sand and shell gravel. Mediomastus fragilis is a sub-surface deposit feeder found on muddy gravel, sandy mud, muddy sand and mud. Lumbrineris gracilis is a predator / omnivore found on gravel / shingle, muddy sand and mud.

Page 66: Mapping of the marine habitats and species of Lamlash Bay, Arran

54

Table 19. SIMPER analysis of the 2008 macrofauna in Lamlash Bay.

% contribution of characterising species Site

Contribution (%)

MCS code

Taxa/species

‘Coarse’ 11.09

8.23

7.92

6.88

5.33

4.81

4.07

3.71

3.66

3.07

3.07

P579

P919

W

P765

P1098

G1

S440

P94

S1142

W1837

P1139

Lumbrineris gracilis

Mediomastus fragilis

Dosinia lupinus

Prionospio fallax

Owenia fusiformis

NEMERTEA spp.

Ampelisca tenuicornis

Pholoe synophthalmica

Tanaopsis graciloides

Thyasira flexuosa

Ampharete lindstroemi

‘Fine’ 40.13

17.81

9.94

7.22

2.44

2.04

1.76

1.72

1.54

1.23

W1906

ZB154

P765

W1837

P94

P499

P1098

P1124

P494

W1569

Mysella bidentata

Amphiura filiformis

Prionospio fallax

Thyasira flexuosa

Pholoe synophthalmica

Nephtys hombergii

Owenia fusiformis

Melinna palmata

Nephtys spp.

Nucula nitidosa

A second SIMPER analysis was carried out assessing the potential differences between the two groups in the ‘fine’ sediment clusters (‘fine 1’ consists of NTZ01 to NTZ04 while ‘fine 2’ consists of NTZ05, NTZ06 and NTZ09) and although there are differences in the communities (table 20), the main difference seem to be the relative contribution of the taxa present as the four most dominant taxa are present in both groups. The results suggest that the relative importance of the number of individuals of both M. bidentata and A. filiformis are the most important differences between the two groups, suggesting that there are subtle differences in the habitats forming these two groups.

Page 67: Mapping of the marine habitats and species of Lamlash Bay, Arran

55

Table 20. SIMPER analysis of taxa within the ‘fine’ cluster groups in the 2008 Lamlash Bay survey (‘fine 1’ consists of NTZ01 – NTZ04 while ‘fine 2’ consists of NTZ05, NTZ06 and NTZ09).

% contribution of characterising species Site

Contribution (%)

MCS code

Taxa/species

‘Fine 1’ 56.24

21.15

3.74

3.71

1.62

W1906

ZB154

W1837

P765

P94

Mysella bidentata

Amphiura filiformis

Thyasira flexuosa

Prionospio fallax

Pholoe synophthalmica

‘Fine 2’ 19.47

18.83

13.56

11.19

6.71

P765

W1906

ZB154

W1837

P1098

Prionospio fallax

Mysella bidentata

Amphiura filiformis

Thyasira flexuosa

Owenia fusiformis

3.3.3 Comparisons between macrofaunal trends and the sediment data

The sediment variables (sediment retained on the different sieve sizes used) were used to assess any correlations between these data and the cluster analysis results (figure 43) using the Spearman rank correlation coefficient (r

s) and the results are given in table 21.

The results show that the highest correlation is found between the faunal distributions and sediment retained on sieve size 1.0 (very coarse sand) with a positive correlation of 0.813 but other correlations are also high with sieve sizes 0.063 (0.777), 0.5 (0.751) and 0.25 (0.612) of particular note (these sieve sizes refer to very fine sand, coarse sand and fine sand respectively). Various combinations of sediment sizes were also tested and the highest correlation was found with a combination of 8.0, 4.0, 1.0 and 0.063 sieve sizes which resulted in a Spearman rank correlation coefficient (r

s) of 0.85. Overall therefore the results

indicate that the sediment size has a strong positive correlation with the faunal distributions in the Lamlash Bay Day grab samples. Table 21. Results from the Spearman rank correlation coefficient analysis comparing the faunal data with the sediment data collected at Lamlash Bay 2008.

Variable

(sieve size, mm) Spearman Rank correlation

16.00

8.0

4.0

2.0

1.0

0.5

0.25

0.125

0.063

<0.063

0

-0.117

-0.115

0.413

0.813

0.751

0.612

-0.181

0.777

0.102

Page 68: Mapping of the marine habitats and species of Lamlash Bay, Arran

56

3.3.4 Biotope classification of the 2008 Lamlash Bay Day grab samples

The combination of the sediment distribution, depth and the macrofauna present has resulted in a biotope classification for the Lamlash Bay Day grab samples (table 22). As noted previously, the dominant sediment type throughout the 10 samples was sand (>80 % in all but one of the samples) suggesting that all the locations should be classified within the ‘main habitat’ SS.SSa (sublittoral sands and muddy sands; see above and Connor et al., 2004). The sediment samples clustered as ‘fine’ in the cluster and ordination analyses (NTZ01 to NTZ06 and NTZ09) could be (based on the depth and sediment size data) classified as biotope complex SS.SSa.CMuSa (circalittoral muddy sand) as the mud content is >5% at all seven locations (see Connor et al., 2004). The dominant fauna in these seven samples (described above) also support this classification but neither of the two biotopes at level 5 in the hierarchy (SS.SSa.CMuSa.AalbNuc and SS.SSa.CMuSa.AbraAirr) seems to fit the fauna identified in Lamlash Bay perfectly. The strongest agreement between many of the taxa present and those described by Connor et al. (2004) are those in the SS.SSa.CMuSa.AalbNuc biotope. This biotope is therefore tentatively selected as the biotope for the locations in the ‘fine’ sediment cluster group (NTZ01 to NTZ06 and NTZ09). The samples clustered as ‘coarse’ (NTZ07, NTZ08 and NTZ10) were classified as muddy gravely sand in the Particle Size Analysis (see PSA section above) and the fauna was dominated by different taxa compared to those seen in the ‘fine’ cluster group. Considering the sediment content and the fauna present, these locations have been classified as biotope complex SS.SMx.CMx. A more detailed assessment of the fauna in the ‘coarse’ cluster suggests that these locations could be classified as SS.SMx.CMx.MysThyMx (level 5 of the biotope classification hierarchy). This particular biotope is found in moderately exposed or sheltered areas with common infaunal polychaetes species including P. fallax and Lumbrineris gracilis as well as amphipods such as Ampelisca spp. together with the brittlestar Amphiura filiformis. However, this biotope is characterised by the bivalves Thyasira spp. (often T. flexuosa) and M. bidentata, and although these taxa are present in the samples, they are not very common. The classification of the locations NTZ07, NTZ08 and NTZ10 can therefore only tentatively be classified as SS.SMx.CMx.MysThyMx at this stage. It should be noted that maerl (P. calcareum), has been recorded (in shallow water) as part of this biotope but at low abundances (Connor et al., 2004), an observation that seems to be valid in the current study (see other sections of this report). Table 22. Summary of the biotope complexes / biotopes for the 2008 Lamlash Bay Day grab samples.

Locations Biotope complex / biotope Sediment type Dominant fauna

NTZ 01#01 NTZ 02#02 NTZ 03#01 NTZ 04#01 NTZ 05#01 NTZ 06#03 NTZ 09#01

SS.SSa.CMuSa.(AalbNuc) muddy sand

Mysella bidentata

Amphiura filiformis

Prionospio fallax

Thyasira flexuosa

Pholoe synophthalmica

Nephtys hombergii

NTZ 07#01 NTZ 08#02 NTZ 10#01

SS.SMx.CMx.(MysThyMx) muddy gravely sand

Lumbrineris gracilis

Mediomastus fragilis

Dosinia lupinus

Prionospio fallax

Owenia fusiformis

NEMERTEA spp.

Page 69: Mapping of the marine habitats and species of Lamlash Bay, Arran

57

3.4 The seabed environment in Lamlash Bay The analysis of the sidescan sonar mosaic and bathymetry together with the ground-truthing using the photography and video records as well as the incorporation of these data into the ArcView GIS allowed for a more detailed assessment of the habitats in Lamlash Bay. The interpretation of the sidescan sonar mosaic (figures 6 and 7) together with the ground-truthing (see examples of photographs above) reveals a complex, patchy and heterogeneous seabed environment difficult to delineate and summarise in a single image. The results are therefore displayed in six images (figures 46 to 51) with the underlying aim not to overly extrapolate the data sets available. Figure 46 illustrates the sidescan sonar imagery together with the bathymetry across the study area while figure 48 focuses on the NTZ. The results not only show a very good match between the features seen but also positions of the shallow banks and regions not surveyed in 2008. Figures 47, 49 and 50 illustrate the distribution of biotope complexes and biotopes identified in the Lamlash Bay study. The maerl biotopes (SS.SMP.Mrl.Pcal) are illustrated in red and these are found in Outer Lamlash Bay, in the North Channel, at the southern entrance to Lamlash Bay and in northern Lamlash Bay with extensive beds within the NTZ. Note, however, that one location (LOC04) north of the entrance to North Channel, is described as scattered maerl rather than a maerl bed and it is currently unclear whether the maerl found here is more extensive or not. The purple polygons can be exemplified by Deacon Rock (figure 47) in the centre of the bay. Some coarse material is found around this outcrop but the substrata change gradually in several directions. The direction of the sidescan sonar survey lines seems to affect the quality of the imagery in this region suggesting a need for careful planning for future sidescan sonar monitoring surveys. Other rocky outcrops (CR.LCR.BsAs.AmenCio) are found in the study area, some of which are found within the brown polygons (in figures 47 and 49), which consist of mixed coarse sediments (SS.SMx.CMx) of muddy, sandy gravel. Most of the rocky outcrops are found in northern Lamlash Bay, but also in the shallow waters across the Bay. The surrounding areas in figures 47 and 50 consist of muddy sand and sandy mud but some of these features need additional ground-truthing (e.g. sediment sampling) to increase the levels of confidence in the data. Some of the data have been extrapolated to create these biotope maps using the confidence model described above. Figures 49 to 51 illustrate the still photography camera transects across the sidescan sonar imagery with each coloured circle signifying one photograph. The colour of the circles signify a biotope (figures 49 and 50) or a substrata type (figure 51) and the changes can be seen along each photographic transect. Overall the correlations between the boundaries of sediment / substrata type and biotope / habitat type are very good throughout the survey area but some of the subtle changes in the faunal distributions are not always apparent in the patterns seen in the substrata. This is best exemplified in the sector in the south-southwest where there is no apparent evidence of faunal change across an area that changes from muddy sand to sandy mud (figures 47 and 50). Additional sediment sampling is required to assess the current infaunal compositions within the bay and allow monitoring of any future potential changes in the infaunal composition in these areas. Furthermore, areas classified as muddy sand biotopes have in many places been classified as sandy mud in the sediment classification and the other way around. The difference in sandy mud and muddy sand is at times very difficult to discern on seabed photographs and video footage but with the presence of burrows and other features the current classification using the photographs is deemed appropriate. Therefore these discrepancies are more likely linked to the biotope classification system which is based on both epi- and infaunal data potentially explaining some of the discrepancies between the two results.

Page 70: Mapping of the marine habitats and species of Lamlash Bay, Arran

58

Figure 46. Bathymetry and sidescan sonar data in Lamlash Bay acquired in 2008 and 2009 (bathymetry data overlaying the sidescan sonar imagery).

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 71: Mapping of the marine habitats and species of Lamlash Bay, Arran

59

Figure 47. Biotope complex distribution in Lamlash Bay (from the video and stills photography analysis) overlaying the sidescan sonar imagery.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 72: Mapping of the marine habitats and species of Lamlash Bay, Arran

60

Figure 48. Bathymetry within the NTZ in Lamlash Bay acquired in 2008 and 2009 (overlaying the sidescan sonar imagery).

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 73: Mapping of the marine habitats and species of Lamlash Bay, Arran

61

Figure 49. Biotope complex distribution within the NTZ in Lamlash Bay (from the video and stills photography analysis) overlaying the sidescan sonar imagery.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 74: Mapping of the marine habitats and species of Lamlash Bay, Arran

62

Figure 50. The biotope distribution within the NTZ in Lamlash Bay from photographic ground-truthing.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 75: Mapping of the marine habitats and species of Lamlash Bay, Arran

63

Figure 51. The sediment distribution within the NTZ in Lamlash Bay from photographic ground-truthing.

Based upon Ordnance Survey material with the permission of the Controller of HMSO © Crown Copyright (2009) Licence no. 100017908

Page 76: Mapping of the marine habitats and species of Lamlash Bay, Arran

64

4 DISCUSSION

4.1 Survey methodology An integrated approach using a range of survey techniques to study an area of seabed has been shown to be successful in many studies. The use of sidescan sonar and photography together with traditional survey techniques (e.g. grab sampling and trawling) have become more commonly used in recent years (e.g. Bett and Masson, 1998; Brown et al., 2002; Axelsson, 2003; Masson et al., 2003; Brown et al., 2004a, b; Stevens and Connolly, 2005; Axelsson et al., 2006). The use of video and still photography in surveys has been shown to be cost-effective with large areas being covered in a relatively short time (Brown et al., 2004a; Stevens and Connolly, 2005) with a number of studies concluding that video and still photography are appropriate for the assessment of the presence and extent of biotopes (Sanderson and Holt, 2001; Service and Golding, 2001) as well as ground-truthing of acoustic images (Brown et al., 2002; Brown et al., 2004a, b). However, it is acknowledged that there is some loss in the taxonomic resolution when using photography rather than biological sampling techniques (e.g. Stevens and Connolly, 2005) and some video records are not of a sufficient quality to allow biotope classifications to be carried out. Still photography should be carried out simultaneously to supply meaningful images (Hiscock and Seeley, 2006). Even though these suggestions may reflect some of the difficulties associated with the biotope classification system, combining digital video and still photography with sediment sampling in surveys appears to have advantages over a single system and allows an overall higher quality assessment to be carried out. Most recently new techniques have been developed to allow seabed discrimination using the acoustic signals from sidescan sonar and multi-beam systems with the potential advantages of being relatively low cost, rapid and allowing coverage of large areas (Preston, 2006a, b). As some techniques and methods used in habitat mapping and seabed discrimination are relatively new, problems are still likely to occur during at least some part of a project involving these methods. The analysis of the sidescan sonar together with the ground-truthing using the drop-down camera system and sediment sampling has proven to be a very successful method for studying the large-scale features in Lamlash Bay but also to investigate the finer biological and sedimentary features within the study area. Overall the sediment distributions and the positions of most of the rocky areas are broadly consistent with those previously reported (see Duncan, 2003) although the current study is more extensive than previous work. Undoubtedly, there are some aspects that could be improved with further work, particularly with more extensive sediment sampling to obtain additional survey coverage and understanding of the infaunal habitats and species in the area. However, the current survey has provided a broad overview of the habitats within the area and can be considered successful in achieving the original aims.

4.2 Survey limitations

4.2.1 Survey extent

The extent of the survey coverage in 2008 was limited as a result of the draught of the survey vessel used, excluding the seabed environment shallower than 10 m. The additional survey work in 2009 resulted in a more extensive coverage, particularly with regards to the acoustic aspect of the work. However, the 2009 ground-truthing coverage had to be limited according to available funds, which therefore restricted the time available to complete survey work in all areas of interest. It would be useful to acquire additional sediment samples as well as further drop-down camera ground-truthing data to allow a more complete description of the biotopes present, assess the densities of commercial and other noteworthy species as well as enable future monitoring of any spill-over effects of the NTZ. In particular, it would be

Page 77: Mapping of the marine habitats and species of Lamlash Bay, Arran

65

very useful to acquire additional drop-down camera data to allow a higher confidence in (and better coverage of) the biotope classification in, for example, the southern region of the NTZ and at the southern entrance to Lamlash Bay. The current ground-truth coverage limits the interpretation and extrapolation of the data (see section 4.3 (limitations) and figures 47, 49 and 50), largely as a result of the complexity of the seabed environment.

4.2.2 Discrepancies between the sidescan sonar and photographic survey results

The interpretation of the sidescan sonar and the subsequent ground-truthing resulted in some discrepancies in the boundaries between different habitats / biotopes. This discrepancy has a number of possible explanations including: 1) positioning of camera frame relative to navigation data; 2) boundaries of biotope / habitat classifications; and 3) depth of acoustic signal penetration. 1) Positioning of camera frame relative to navigation data During the survey operations, a lay-back error was calculated for the camera frame position in relation to the vessel; however, as this was based on the position of the A-frame from which the camera frame was deployed and did not allow for movements of the camera on seabed, slight inaccuracies in positions of the camera deployments are possible (maximum inaccuracies are estimated to be 5 m). 2) Biotope / habitat classifications The exact positions of the boundaries between different biotopes were often difficult to determine as many boundaries were transitional in nature. The change in sediment tended to be gradual along transects and the determination of the boundary between two types became difficult. Some of the boundaries identified from the camera deployments did therefore not match exactly with those seen on the sidescan sonar imagery but overall the results were good. 3) The depth of the acoustic signal It has been shown that some sidescan sonar frequencies (e.g. 30 kHz) penetrate into the sediment and therefore return volume backscatter rather than a backscatter signal from the uppermost sediment surface (Blondel and Murton, 1997; Axelsson, 2003; Masson et al., 2003). The lower frequencies penetrate the most (Blondel and Murton, 1997) and at 30 kHz the signal could penetrate tens of centimetres (Masson et al., 2003). It is therefore possible that in certain sediments, acoustic signals at 100 kHz could penetrate the sediment to some degree and therefore generate results different to those seen on seabed photographs. However, as 410 kHz was used in the Lamlash Bay survey, signal penetration is unlikely to explain any of the discrepancies seen in this study. 4.3 Confidence assessment In this study attempts have been made to minimise interpolation of the data as much as possible but as with many similar studies ground-truth coverage is not as extensive as perhaps desired. In order to illustrate the quality and interpretation and interpolation of the data confidence ratings have been given to the figures. The confidence in the level of certainty of the interpretation of the seabed environment and biotope maps (e.g. figures 47 and 50) at point source (i.e. a photograph) is estimated at 90-100 % accurate (level 1), while confidence level 2 (areas with the same backscatter surrounding a photograph) is estimated to be 80 % accurate and confidence level 3 (areas with the same levels of backscatter but some distance away (c. 50 - 100 m) from the photographs) is estimated to be approximately 50-60 % accurate (e.g. southern NTZ). The reasoning behind the relatively low confidence rating for level 3 is best illustrated by the ground-truthing of the southern and central Lamlash Bay (see figure 47 and 50), where some areas of the seabed appear similar on the sidescan sonar records but are shown to be different by the photographic ground-truthing data.

Page 78: Mapping of the marine habitats and species of Lamlash Bay, Arran

66

Video and still photography have been shown to be appropriate for the assessment of the presence and extent of biotopes, but the quality of the data interpretation and biotope records from video tapes relies on the identification skills of the person scoring the tapes (Holt et al., 2001). The classification of biotopes is also somewhat subjective and not all seabed environments ‘fit’ the biotope classification scheme resulting in some biotopes being classified to ‘best fit’ the communities present (Holt et al., 2001). Some studies therefore develop a specific habitat or biotope classification scheme (see for example Brown et al., 2004a) for that survey but long-term perhaps the biotope classification system by Connor et al. (2004) will be developed further and therefore more widely used. However, obtaining high level of confidence in the data still requires an experienced marine biologist, ideally with local knowledge, although a quality assurance programme including re-assessments of the tapes can recover any lost data or mistakes in the classifications (Holt et al., 2001).

4.4 Comparisons with previous studies As mentioned above, there are no records for Lamlash Bay within the Marine Nature Conservation review (MNCR) database (now Marine Recorder) prior to 2003 (Duncan, 2003; COAST, 2005). Furthermore, only a couple of studies appear to have been carried out since then (Duncan, 2003; Kamenos et al., 2004), both of which describe Lamlash Bay as primarily composed of soft substrata apart from a number of rocky reefs (e.g. Deacon Rock and areas along the North Channel). Although direct comparisons are difficult as a result of the lack of exact positions of many features in previous studies, the ground-truthing of the seabed environment in 2008 and 2009 largely agrees with the previous studies as well as the records on the admiralty charts, with mixed sediments (muddy, sand and gravel) in the North Channel and muddy sand in the central and southern regions of Lamlash Bay. The admiralty charts furthermore illustrate increased mud content along the south-western section of Lamlash Bay, a feature also seen in the 2008 survey. In 2003 the main biological features of note in Lamlash Bay were maerl and sea grass (eelgrass) beds as well as sublittoral rocky reefs with kelp (Duncan, 2003). In 2008 and 2009, maerl was found to be extensive in Lamlash Bay but the sea grass beds were not identified at all. This could be explained by the fact that survey work in the north-western region, where the seagrass has previously been found, was obstructed by moorings. Additional survey work may therefore be required to assess this feature further. Rocky reefs were also identified in both the 2008 and 2009 Lamlash Bay surveys.

4.4.1 Lamlash Bay rocky reefs

A number of rocky reefs (mainly boulder, cobble debris areas) were found in the 2008 survey, including the rocky outcrop known as Deacon Rock. Additional reef features were found during the 2009 survey work in the shallow waters along the northern coastline, including Gurnard Bay (Holy Isle) and the northern coast near Clauchlands Point, and around Holy Island.

Deacon Rock has been described as a submerged reef rising from 17 m up a steep slope of sea squirt covered boulders to a sugar kelp (Laminaria saccharina) and mixed red algae crowned apex at 10 m. The 2008 study did collect acoustic data across Deacon Rock and camera transects did record the sea squirt covered boulders around it but the kelp covered apex was not surveyed using the drop-down camera (too shallow for the vessel used), although kelp debris was found on the seabed in the surrounding areas. Other flora and fauna found on and around the boulders included red algae, short and long hydroid turf, C. lloydii, T. communis, Ophiura sp. and sponges (Suberites sp.) as well as O. nigra brittlestar beds on some sections of the outcrop.

Page 79: Mapping of the marine habitats and species of Lamlash Bay, Arran

67

A number of bedrock ledges near the fish farm cages are also apparently present (Duncan, 2003; Seasearch, 2003). These rocks are reportedly covered in black silt with relatively sparse fauna present but because of the presence of the fish cages it was not possible to cover these areas in the 2008 or 2009 surveys.

4.4.2 Lamlash Bay maerl beds

Maerl is the collective name for several species of calcified red seaweed with three species of maerl occurring in the UK, P. calcareum, Lithothamnion glaciale and Lithothamnion corallioides. Maerl grows as unattached nodules on the seabed and under favourable conditions can form extensive beds (UKBAP, 2008). Maerl is slow growing but over long periods its dead calcareous skeleton can accumulate into deep deposits (an important habitat in its own right), overlain by a thin layer of pink living maerl. Maerl beds typically develop on coarse sediments and in clean water where there is some tidal flow, such as in the narrows and rapids of sea lochs, or in open sea where wave action is sufficient to remove fine sediments (Hall-Spencer and Moore, 2000; Kamenos et al., 2004; UKBAP, 2008). Although maerl beds cover a very small area of UK waters, all of the beds studied to date have been found to harbour disproportionately high diversity and abundance of associated organisms in comparison with surrounding biotopes; some of these species are confined to maerl habitat or rarely found elsewhere (Nunn, 1992; Hall-Spencer and Moore, 2000). In the Lamlash Bay surveys only P. calcareum was identified with certainty, with occurrences in the northern and Outer Lamlash Bay areas as well at the southern entrance to Lamlash Bay (area E). Maerl was not identified in the central regions of Lamlash Bay. Duncan (2003) reported the remains of a deep maerl bed (Phymatolithon sp.) in the North Channel entrance. This maerl bed is still evident in 2008 but as with the majority of the maerl beds found in 2008 and 2009, these beds are in relatively poor health. The healthiest maerl bed was found in the northern region of the Outer Lamlash Bay area (camera transects LOC01, A01 and A02) with up to 5-10% live maerl whilst only scattered maerl in poor condition was found at the northern edge of the North Channel entrance (LOC04). The maerl beds along the northern border of Lamlash Bay and along the North Channel were generally found to have approximately 1 % of live maerl, at least in sections, but the healthy beds described by Kamenos et al. (2004) south of Clauchlands Point (up to 85-95 % live maerl) were not identified in either of the 2008 or 2009 surveys despite the increased survey coverage in 2009 (including the shallower depths on the banks surrounding North Channel in particular). The deepest record of maerl in the Lamlash Bay 2008 survey was at approximately 26 m. These results are in line with those reported in a study describing maerl being found down to about 30 m depth in European waters (Hall-Spencer and Moore, 2000).

4.4.3 Other fauna and flora in Lamlash Bay

Apart from the maerl beds mentioned above, no other rare species (see Sanderson, 1996; Vincent et al., 2004) were identified in the Lamlash Bay surveys; although it is possible that some remain undetected as a result of the taxonomic limitations of using camera equipment (see Stevens and Connolly, 2005). As a general rule sponges were rarely recorded in the Lamlash Bay study but comparisons with Duncan (2003) and Seasearch (2003) are difficult as sponges were equally rare in these studies. The most common sponges group in the Lamlash Bay study were the encrusting sponges but individuals of Amphilectus fucorum and Suberites sp. were also recorded, all of which were found in the rocky areas in the northern and Outer Lamlash Bay areas.

Page 80: Mapping of the marine habitats and species of Lamlash Bay, Arran

68

As in the previous studies (Duncan, 2003; Seasearch, 2003) the Cnidaria was dominated by small and tall hydroid turf (including Nemertesia spp.) in rocky areas, as well as C. lloydii in soft sediment areas. As in these earlier studies M. senile and Alcyonium digitatum were also recorded. In Crustacea the squat lobster M. rugosa was by far the most common taxon in 2008 with juvenile Galathea sp. being relatively abundant in the grab samples. Hermit crabs were recorded but some were most likely undetected and thus might be more common than recorded in this study. Squat lobsters were recorded in earlier studies but not to species level and hermit crabs were found to be widespread (see Duncan, 2003). Edible crab (C. pagurus) and lobsters (H. gammarus) were reportedly present at a number of sites in 2003 (7 and 5 locations respectively) but in 2008 and 2009 edible crabs were rare and lobsters were not recorded at all. The dive surveys in 2003 are more likely to detect and record these crustaceans compared to drop-down camera deployments which may at least partly explain this difference. T. communis was by far the most common Mollusca taxon in the 2008 and 2009 camera footage, a taxon only mentioned in passing in the 2003 surveys. Scallops, on the other hand, were widely distributed in the 2003 surveys. Of particular note in 2003 was a large number of juveniles ‘in the middle of the north channel’, the return of scallops to Deacon Rock as well as the complete absence of scallops near the fish cages (south-southwest) (Duncan, 2003; Seasearch, 2003). Scallops are also widely distributed in the 2008 and 2009 Lamlash Bay surveys but only at densities of approximately 3 individuals/100m2 (there are no density values for the 2003 surveys). The dominant taxa in the Day grab samples are given in table 17 but as there are no other available data, comparisons are not possible. Among the Echinodermata brittlestars were the most abundant taxa in both 2008 and 2009 with Ophiura sp. and Amphiura sp. being the most abundant taxa overall. Other recorded echinoderms were O. nigra, A. bifida and E. esculentus. References were made to brittlestars and featherstars in the 2003 surveys (Duncan, 2003; Seasearch, 2003) but these taxa were not identified to species level. Among the infaunal samples (the 2008 Day grab samples) other echinoderms were present but no comparable data are available from previous studies. A large number of fish species (19 in total) were recorded in the 2003 surveys with flatfish being the most common (Duncan, 2003). Pollock was also present together with several different species of gobies and wrasse. In 2008 and 2009 only a few fish species were recorded, presumably as a result of this group being highly mobile and avoiding the camera during deployments. A few commercial species including Haddock (Melanogrammus aeglefinus) and some flatfish were identified but the most common fish were gobies and dragonets, species also identified in 2003.

4.4.4 Biotope distribution in Lamlash Bay

There are no biotope classification records for Lamlash Bay prior to the 2008 Lamlash Bay survey. A general overview of the number of biotope complexes present within the Irish Sea region has been produced as part of the Irish Sea Pilot (Vincent et al., 2004). These results suggest approximately four biotope complexes in the Lamlash Bay area (results based on data collation and some ground-truthing) but there is a general lack of detailed information for the Clyde region outside the sea lochs. A habitat mapping study was initiated by the Scottish Sustainable Marine Environment Initiative Firth of Clyde Pilot Project but the final report has not been completed as a result of a number of complications together with the lack of ground-truthing of acoustic data (SSMEI,

Page 81: Mapping of the marine habitats and species of Lamlash Bay, Arran

69

2007). However, a predictive model of the seabed habitats present within the Clyde Sea region was produced indicating a total of five seabed habitat types within Lamlash Bay and six seabed habitat types in the entire Lamlash Bay region (Lamlash Bay and the Outer Lamlash Bay areas). These results agree with those seen in the 2008 Lamlash Bay survey (see figure 47), however, three additional biotopes were identified in the shallow regions (<10 m) in the bay in the 2009 survey and additional biotopes may be present in regions not covered by the 2008 and 2009 ground-truthing surveys.

4.4.5 Evidence of fishing activity

Indications of fishing activity were evident in October 2008 when fishing vessels were observed moving into Lamlash Bay immediately after the completion of the 2008 survey operations. In addition, scallop dredging marks were identified in the photographic and sidescan sonar imagery close to Rocky Bank in central Lamlash Bay (see target 3 above). However, there was no evidence of any dredging marks in the 2009 data. Continued monitoring by sidescan sonar could potentially be used to assess any future dredging, although bearing in mind that such marks will gradually be removed by physical processes and the likely duration of persistence is not known at this location. 4.5 Recommendations for future monitoring at Lamlash Bay The primary issue fundamental to any future monitoring will be the enforcement of the fishing ban to ensure there is no fishing activity within the NTZ. Without enforcement there might be a need for additional monitoring survey work to assess any changes to the seabed environment following such activities. Assuming there is no fishing activity within the NTZ the recommendations below are based on Special Areas of Conservation (SAC) monitoring attributes (Davies et al., 2001). Lamlash Bay is not an SAC but the attributes and targets proposed for monitoring are relevant here (table 23). Table 23. Lamlash Bay monitoring attributes.

Attribute Target Sediment character Monitor any changes in composition of sediment types

across the bay. Distribution of biotopes Monitor any changes in the distribution of biotopes.

Extent of sub-feature Monitor any changes in extent of biotopes. Monitor the extent of maerl and sea grass beds Monitor the quality of maerl and sea grass beds Monitor any changes in other characteristic species

Species composition Monitor any changes in biotope quality as a result of addition or removal of notable species

Species population measures - population structure of a species - presence or abundance of specified species

Monitor species abundance and composition of biotopes Monitor age/size class structure Monitor presence / abundance of characteristic species

4.5.1 Requirement for additional survey effort

Page 82: Mapping of the marine habitats and species of Lamlash Bay, Arran

70

There are several habitats and species within Lamlash Bay requiring future monitoring measures. The 2008 and 2009 Lamlash Bay surveys collected considerable amounts of baseline data (as outlined above) but additional baseline data are required to allow future monitoring of all the habitats and species present in Lamlash Bay. The Day grab sediment sampling programme should be extended to the areas surrounding the NTZ to allow monitoring of the habitats and species present (e.g. changes in species composition and animal sizes as well as potential spill-over effects). Scuba-diving surveys are also recommended to collect additional qualitative (transect surveys) and quantitative data (using quadrats) on the sea grass beds (e.g. shoot count and sea grass length measurements) to allow future recovery assessment studies.

4.5.2 Future monitoring in Lamlash Bay

4.5.2.1 Sediment character

The character of the seabed environment might change as a result of the classification of the NTZ. The sediment composition could be assessed using the same methodology as in this survey to allow continuity of analysis and monitoring. Camera deployments and sediment sampling using a 0.1 m2 stainless steel Day grab should be carried out at 2 year intervals.

4.5.2.2 Distribution of biotopes

This attribute could be assessed by monitoring the spatial extent of the different biotopes within Lamlash Bay simply by monitoring the identity of the biotopes at the existing sampling stations. Camera deployments and sampling of infauna and epifauna using a 0.1 m2 stainless steel Day grab should be carried out every 2 years. Monitoring of the presence and abundance of characteristic species for each biotope should be carried out and it would be useful to extend the grabbing survey into the areas surrounding the NTZ. It would also be useful for future monitoring (every 6 years) to continue to identify biotope boundaries using broad scale techniques (e.g. sidescan sonar) so that at least very conspicuous changes will become apparent.

4.5.2.3 Extent of sub-features

The extent of biologically important biotopes such as the sea grass beds is unknown. Investigations into the extent and distribution of this feature should be carried out. The use of drop-down video together with scuba-diving transects and quadrat sampling (for repeatability) during favourable weather conditions is recommended. Continued monitoring of the extent of the important sub-features (maerl in particular) should be carried out using drop-down camera and Day grab sediment sampling at 2 year intervals with acoustic (sidescan sonar) monitoring carried out at 6 year intervals.

4.5.2.4 Species composition and population measures

Repeat drop-down camera and sediment sampling surveys should be carried out at 2 year intervals to assess any changes in the main habitats and species present in Lamlash Bay. The quality and extent of the maerl beds as well as the size, quality and extent of other habitats (e.g. size and extent of brittlestar beds and hydroid turf, the latter potentially associated with scallop recovery) could be assessed using the current methodology. Additional scuba-diving surveys of the sea grass beds could also be carried out at 2 year intervals to assess any changes to the sea grass beds and the associated species.

Page 83: Mapping of the marine habitats and species of Lamlash Bay, Arran

71

Changes to scallop populations and other noteworthy taxa (e.g. commercial fish and shellfish species as well as invertebrates forming part of the diet of commercial species) could also be monitored using the methodology described in this study. Both P. maximus and A. opercularis spawn twice a year (environmental conditions allowing) with P. maximus being sexually mature after two years and A. opercularis after one year (Barnes, 2009a; Marshall and Wilson, 2009). A three year old P. maximus can produce 15-21 million oocytes (no data for A. opercularis) with a high dispersal potential (10 km or more), allowing any changes in population sizes to occur within one or two years. Other commercial species noted above, such as C. pagurus, N. norvegicus and H. gammarus, take 6-10, 3-4 and 6-10 years respectively to reach sexual maturity (Tuck et al., 2000; Barnes, 2009b; Neal and Wilson, 2009). Consequently, as well as the absolute abundance of organisms, the reproductive capacity of these populations is important in considering the recovery times of such species. Monitoring by means of camera deployments and sediment sampling using a 0.1 m2 stainless steel Day grab should therefore be carried out at 2 year intervals to allow continuous and regular monitoring of any changes to the faunal populations and species composition. With the sediment sampling effort extended to include areas outside the NTZ, direct comparisons (e.g. abundance and spill-over effects) between the changes of faunal populations within the NTZ with those outside the area will be possible.

5 CONCLUSIONS

The habitat mapping surveys in Lamlash Bay in 2008 and 2009 were a success in terms of collecting high quality data including sidescan sonar and echosounder data together with high quality ground-truthing still photographs and video footage. The sediment sampling programme was also deemed a success but additional sampling outside the NTZ is required to allow direct comparisons (e.g. changes in species composition, changes in abundance and spill-over effects) between the two areas in future monitoring. The ground-truthing of the sidescan sonar data revealed a heterogeneous and complicated seabed environment dominated by sand but areas of mixed sediment, bedrock as well as areas of boulder / cobble debris are also present. The analysis of the photographic material (stills photographs and video) revealed a total of nine biotopes within Lamlash Bay to date: SS.SMx.CMx.ClloMx(Nem), SS.SMX.CMx.OphMx, SS.SMu.CSaMu.(AfilMysAnit), SS.SSa.CMuSa.(AbraAirr), SS.SMP.KSwSS.LsacR.CbPb, SS.SMP.KSwSS.LsacR.Sa, CR.LCR.BrAs.AmenCio, CR.LCR.BrAs.AmenCio.Bri and SS.SMP.Mrl.Pcal. The classification of some of these biotopes is tentative (part of biotope in brackets) as the faunal composition does not fit the biotope descriptions perfectly. An additional two biotopes have tentatively (for the same reasons as with the camera footage) been classified within the sediment samples: SS.SSa.CMuSa.(AalbNuc) and SS.SMx.CMx.(MysThyMx). The fauna include C. lloydii, M. rugosa, T. communis, P. maximus, A. opercularis, featherstars and various polychaetes as well as brittlestars including Amphiura sp., Ophiura sp and O. nigra. Of particular note is the presence of maerl beds in the northern and outer Lamlash Bay regions, an important biological feature shown to harbour disproportionately high diversity and abundance of associated organisms in comparison with surrounding habitats.

Page 84: Mapping of the marine habitats and species of Lamlash Bay, Arran

72

It has been recommended that future monitoring using photographic (stills photography and video) and sediment sampling surveys should be carried out at 2 year intervals to assess any changes and benefits of the establishment of the NTZ in Lamlash Bay.

Page 85: Mapping of the marine habitats and species of Lamlash Bay, Arran

73

6 REFERENCES

6.1 General references Axelsson, M.B. 2003. The deep seabed environment of the UK continental margin – integration and interpretation of geological and biological data. Ph.D. thesis, University of Southampton, 266 pages. Axelsson, M., Dewey, S., Chaddock, S. and Duke, S. 2006. Survey of the reef habitat around Eddystone Rocks, Plymouth. Report to English Nature, 55 pages. Ballantine, B. 1994. Networks of ‘no-take’ marine reserves are practical and necessary. International Conference on Science and Management of Protected Areas, in Halifax, Canada, 8 pages. Barnes, M. 2009a. BIOTIC Species Information for Aequipecten opercularis. Available online from: www.marlin.ac.uk/biotic Barnes, M. 2009b. Biotic species information for Homarus gammarus. Available online from: www.marlin.ac.uk/biotic Bett, B.J. and Masson, D.G. 1998. Atlantic Margin Environmental Surveys. In, In depth, Southampton Oceanography Centre, issue 3, pages 4-5. BiodiversityScotland, 2008. The Scottish Biodiversity List. Biodiversity Implementation Team Scottish Natural Heritage. Available (online): www.biodiversityscotland.gov.uk/pageType2 .php?id=35&type=2&navID=92 (accessed January 2010). Blondel, P. and Murton, J.M. 1997. Handbook of seafloor sonar imagery. John Wiley & Sons Ltd., 314 pages. Brown, C.J., Cooper, K.M., Meadows, W.J., Limpenny, D.S. and Rees, H.L. 2002. Small-scale mapping of seabed assemblages in the Eastern English Channel using sidescan sonar and remote sampling techniques. Estuarine, Coastal and Shelf Science, 54, 263-278. Brown, C.J., Hewer, A.J., Limpenny, D.S., Cooper, K.M., Rees, H.L. and Meadows, W.J. 2004a. Mapping seabed biotopes using sidescan sonar in regions of heterogeneous substrata: case study east of the Isle of Wight, English Channel. International Journal of the Society for Underwater Technology, 26, 27-36. Brown, C.J., Hewer, A.J., Meadows, W.J., Limpenny, D.S., Cooper, K.M. and Rees, H.L. 2004b. Mapping seabed biotopes at Hastings Shingle Bank, eastern English Channel. Part 1. Assessment using sidescan sonar. Journal of the Marine Biological Association of the United Kingdom, 84, 481-488. Clark, B. 1999. MCS campaign for No Take Zones in UK waters. Marine Conservation, 4 (4), 10-11. Clarke, K.R. and Warwick, R.M. 1994. Change in marine communities: an approach to statistical analysis and interpretation. Plymouth Marine Laboratory, 144 pages. COAST 2005. Isle of Arran, Community of Arran Seabed Trust (COAST) 2005. [online]. Available from: www.arrancoast.co.uk

Page 86: Mapping of the marine habitats and species of Lamlash Bay, Arran

74

Connor, D.W., Allen, J.H., Golding, N., Howell, K.L., Lieberknecht, L.M., Northen, K.O. and Reker, J.B. 2004. The marine habitat classification for Britain and Ireland (version 04.05). Joint Nature Conservation Committee, 49 pages. Davies, J., Baxter, J., Bradley, M., Connor, C., Khan, J., Murray, E., Sanderson, W., Turnball, C. and Vincent, M. 2001. Marine Monitoring Handbook. Joint Nature Conservation Committee, 405 pages. Duncan, C. 2003. Lamlash Bay Seasearch 2003. Interim report by the Marine Conservation Society to Scottish Natural Heritage. 25 pp. Folk. R.L. 1954. The distinction between grain size and mineral composition in sedimentary-rock nomenclature. Journal of Geology, 62, pp 344–359. Fowler, J. and Cohen, J. 1992. Practical Statistics for Field Biology. John Wiley & Sons, Chichester, England, 227 pages. Gage, J.D. and Tyler, P.A. 1991. Deep-Sea Biology: A natural history of organisms at the deep-sea floor. Cambridge University Press, 504 pages. Gazetteer for Scotland 2008. Gazetteer for Scotland. 2008. [online]. Available from: http://www.geo.ed.ac.uk/scotgaz/gaztitle.html Hall-Spencer, J. M. and Moore, P. G. 2000. Scallop dredging has profound, long-term impacts on maerl habitats. ICES Journal of Marine Science. 57. 1407-1415. Hiscock, K. and Seeley, B. 2006. Quality assurance of biotope classification from drop-down video footage. Report to Countryside Council for Wales from the Marine Life Information Network (MarLin). Plymouth: Marine Biological Association of the UK, 20 pages. Holt, R.H.F., Sanderson, W.G., Kay, L., & Williams, R. 2001. Section 2.1.2 Evaluation of drop-down video for measuring biotope richness off north Pen Llŷn. In The establishment of a programme of surveillance and monitoring for judging the condition of the features of Pen Llŷn ar Sarnau cSAC:1. Progress to March 2001. W.G. Sanderson, R.H.F. Holt, L. Kay, G. Wyn & A.J. McMath eds. Bangor, CCW contract Science Report No: 380 (UK Marine SAC’s Project), 400pp. Howson, C.M. and Picton, B.E. 1997. The species directory of the marine fauna and flora of the British Isles and surrounding seas. Ulster Museum Publication, 276. The Ulster Museum: Belfast, UK. JNCC, 2008. No Take Zones. Joint Nature Conservation Committee (Online). Available from: www.jncc.gov.uk/page-1572. Kamenos, N. A., Moore, P. G., and Stevenson, T. D. I. 2004. Marine survey to identify maerl beds: Lamlash Bay Long Sea Outfall. University Marine Biological Station Millport. Scottish Water Solution Reference Number: SWS/FW/2004-0427-01, 10 pages. Leeder, M.R. 1982. Sedimentology: process and product. Chapman and Hall, 344 pages. Marshall, C. and Wilson, E. 2009. BIOTIC Species Information for Pecten maximus. Available online from: www.marlin.ac.uk/biotic.

Page 87: Mapping of the marine habitats and species of Lamlash Bay, Arran

75

Masson, D.G., Bett, B.J., Billett, D.S.M., Jacobs, C.L., Wheeler, A.J. and Wynn, R.B. 2003. The origin of deep-water, coral-topped mounds in the northern Rockall Trough, Northeast Atlantic. Marine Geology, 3288, 1-22. MCS, 2007. Marine Conservation Society (MCS) joins Natural England Lundy NTZ scientific monitoring work (online). Available from: www.mcsuk.org/downloads/biodiversity/ MCSjoinsLundy.pdf. Neal, K. and Wilson, E. 2009. Biotic species information for Cancer pagurus. Available online from: www.marlin.ac.uk/biotic Nunn, J. 1992. The molluscan fauna associated with maerl. Conchologist’s Newsletter. 125. 161-167. Pearce, J. 2002. The future of fisheries – marine protected areas – a new way forward or another management glitch? Marine Pollution Bulletin, 44, 89-91. Preston, J. 2006a. Sidescan seabed classification processing of GeoAcoustic sidescan XTF data. A report to Seastar Survey, March 2006, 24 pages. Preston, J. 2006b. Acoustic seabed classification with multibeam and sidescan images. Acoustic seabed classification tutorial notes by the Quester Tangent Corporation, 53 pages. Roberts, C.M., Bohnsack, J.A., Gell, F., Hawkins, J.P. and Goodridge, R. 2001. Effects of Marine Reserves Naso vlamingii Apo Island, Central Philippines. Marine Ecology Progress Series, 264, 15-20. Russ, G.R. and Alcala, A.C. 1996. Do marine reserves export adult fish biomass? Evidence from Apo Island, Central Philippines. Marine Ecology Progress Series, 132, 1-9. Russ, G.R., Alcala, A.C. and Maypa, A.P. 2003. Spill-over from marine reserves: the case of biotopes using video recordings. In, Marine Monitoring Handbook, (eds) Jon Davies, John Baxter, Martin Bradley, David Connor, Janet Khan, Eleanor Murray, William Sanderson, Caroline Turnball, Malcolm Vincent, JNCC, Peterborough. Sanderson, W.G. 1996. Rare marine benthic flora and fauna in Great Britain: the development of criteria for assessment. JNCC Report Number 240. Sanderson, W.G. and Holt, R.H.F 2001. Update to: Procedural guideline No. 3-5 identifying on adjacent fisheries. Science, 294, 1920-1923. SSMEI, 2007. Scottish Sustainable Marine Environment Initiative Firth of Clyde Pilot Project. Project Update (6th

Quarterly Report), November 2007, 3 pages. Scottish Government 2008. Scottish Statutory Instrument 2008 No. 317. Sea Fisheries. The Inshore Fishing (Prohibition on Fishing) (Lamlash Bay) (Scotland) Order 2008. [online]. Available from: http://www.opsi.gov.uk/legislation/scotland/ssi2008/pdf/ssi_20080317_en.pdf Seasearch, 2003. Arran Lamlash Bay Survey: summer 2003 summary report (online). Available from: www.seasearch.org.uk. Service, M. and Golding, N. 2001. Procedural guideline No. 3-14 In-situ survey of sublittoral epibiota using towed sledge video and still photography. In, Marine Monitoring Handbook, (eds) Jon Davies, John Baxter, Martin Bradley, David Connor, Janet Khan, Eleanor Murray, William Sanderson, Caroline Turnball, Malcolm Vincent, JNCC, Peterborough.

Page 88: Mapping of the marine habitats and species of Lamlash Bay, Arran

76

Solandt, J-L., Beukers-Stewart, B., Davis, C., Hoskin, M., Wood, H. and McNeish, D. 2007. Lessons learned from setting up small MPAs in the UK and Isle of Man. Poster produced by the Marine Conservation Society, found at www.mcsuk.org. Stevens, T. and Connolly, R.M. 2005. Local-scale mapping of benthic habitats to assess representation in a marine protected area. Marine and Freshwater Research, 56, 111-123. Tuck, I.D., Chapman, C.J. and Atkinson, R.J.A. 2000. Population biology of the Norway lobster Nephrops norvegicus (L.) in the Firth of Clyde, Scotland. I. Growth and density. ICES Journal of Marine Science, 54, 125-135. UKBAP, 2008. UK Biodiversity Action Plan: List of Priority Habitats. [online]. Available from: www.ukbap.org.uk/PriorityHabitats.aspx Vincent, M.A., Atkins, S.M., Lumb, C.M., Golding, N., Lieberknecht, L.M. and Webster, M. 2004. Marine nature conservation and sustainable development - the Irish Sea Pilot. Report to Defra by the Joint Nature Conservation Committee, Peterborough. 6.2 References used for video and still photography identification Barrett, J. and Yonge, C.M. 1958. Collins pocket guide to the sea shore. Collins Clear-Type Press, London, 272 pages. British Isles Seaweed Images (online). Available from: www.weedseen.co.uk Campbell, A. 1994. Seashores and shallow seas of Britain and Europe. The Hamlyn Publishing Group Ltd., 320 pages. Encyclopedia of Marine Life of Britain and Ireland (online). Available from: www.habitas.org.uk/marinelife Gibson, R., Hextall, B. and Rogers, A. 2001. Photographic guide to the sea & shore life in Britain and North-West Europe. Oxford University Press, 436 pages. Hayward, P.J. and Ryland, J.S. 1995. Handbook of the marine fauna of north-west Europe. Oxford University Press, 800 pages. Howson, C.M. and Picton, B.E. 1997. The species directory of the marine fauna and flora of the British Isles and surrounding seas. Ulster Museum and the Marine Conservation Society. Ulster Museum Publication number 276, 509 pages. Naylor, P. 2005. Great British marine animals (2nd edition). Sound Diving Publications, 272 pages. Tebble, N. 1966. British bivalve seashells: a handbook for identification. Published (electronic edition) by Pisces Conservation Ltd, 2006 by kind permission of the British Museum (Natural History). The Marine Fauna gallery of Norway (online). Available from: www.seawater.no The Marine Life Information Network for Britain and Ireland (online). Available from: www.marlin.ac.uk

Page 89: Mapping of the marine habitats and species of Lamlash Bay, Arran

77

Appendix 01. Survey equipment specification

Positioning Leica RTK GPS (GX1230 Real Time Rover System) Accuracy: Horizontal: 10 mm + 1 ppm, kinematic Vertical: 20 mm + 1 ppm kinematic Survey Management Software Hypack 2008: Navigation, sidescan sonar, echosounder Motion Sensor TSS CMS 25 Compact Motion Sensor Heave: Range ± 100 m. Resolution 1 cm. Accuracy 5 cm or 5 % of range Roll, Pitch: Range ± 30˚. Resolution 0.01˚ - accuracy dynamic/static 0.25˚ RMS Echosounder 2008 Simrad EK60 Scientific Echosounder Recording and processing at 38 kHz 2009 Marimatech E-Sea Sound 206C (33/200 kHz) Recording and processing at 200 kHz Sidescan Sonar EdgeTech 4200-FS dual frequency (100/400 kHz) sidescan sonar Survey recording and processing of 410 kHz Sound Velocity Probe Marimatech HMS 1820-P CTD Range: 0-250 m. Temperature -55˚C - +55˚C. Salinity 1-60 Resolution: Depth 1 cm, Temperature 0.01˚C, Salinity 0.001 Accuracy: Depth 0.1% of range, temperature ± 0.1˚C. Salinity ±0.001 Sediment Sampling Equipment 0.1 m2 Day grab

Page 90: Mapping of the marine habitats and species of Lamlash Bay, Arran

78

Appendix 02. Navigation checks and tidal curves

Navigation checks in 2008 An example of RTK GPS derived tide data, which was applied to the bathymetry data in real time during the Lamlash Bay survey. Tide heights relative to Ordnance Datum Newlyn (ODN) ODN is 1.5m above Admiralty Chart Datum (ACD) at Lamlash

Page 91: Mapping of the marine habitats and species of Lamlash Bay, Arran

79

Start and end of survey navigation checks. The known position was measured from a 1:500 scale Ordnance Survey Super Plan. The RTK GPS derived positions were logged over a period of 5 minutes and compared against the position measured from the super plan.

Page 92: Mapping of the marine habitats and species of Lamlash Bay, Arran

80

Navigation checks in 2009 Start of survey navigation check – Clyde Marina, Ardrossan 10/09/2009

End of survey navigation check – Clyde Marina, Ardrossan 14/09/2009

Page 93: Mapping of the marine habitats and species of Lamlash Bay, Arran

81

Appendix 03. Field survey logs

Appendix 03: Lamlash Bay Sidescan sonar and Bathymetry log - 2008SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) and WGS84 Latitude and Longitude (ddmmss.sss)

Client SNH Job Number J/08/151

Location Lamlash Bay Vessel SV Alma Na Mara

Survey Lamlash Bay Habitat Mapping Survey Date October 2008DATE Site Line SOL Time EOL Time Direction Comments

Name Name (GMT) (GMT) East (m) North (m) East (m) North (m)

15/10/2008 Lamlash North 114 06:58:03 07:25:43 SW - NE 203653.91 630307.41 206696.51 632561.97

15/10/2008 Lamlash North 111 07:31:18 08:00:41 NE - SW 206570.69 632662.04 203462.97 630364.62

15/10/2008 Lamlash North 112 08:04:58 08:35:14 SW - NE 203335.89 630201.99 206644.18 632643.96

15/10/2008 Lamlash North 115 08:40:26 09:07:59 NE - SW 206703.79 632505.29 203676.54 630273.75

15/10/2008 Lamlash North 110 09:12:58 09:42:26 SW - NE 203291.05 630296.48 206567.04 632716.11

15/10/2008 Lamlash North 113 09:46:17 10:16:38 NE - SW 206727.39 632645.69 203509.45 630274.42

15/10/2008 Lamlash North 108 11:00:55 11:18:18 NE - SW 203273.70 630394.37 205405.56 631986.20 anchor scares on sidescan

15/10/2008 Lamlash North 107 11:22:22 11:41:01 NE - SW 205417.41 632080.06 203290.94 630488.37

15/10/2008 Lamlash North 104 11:46:23 12:04:16 SW - NE 203121.56 630564.80 205301.07 632151.95

15/10/2008 Lamlash North 101 12:06:40 12:26:03 NE - SW 205240.53 632277.57 203074.35 630695.93

15/10/2008 Lamlash North 102 12:28:34 12:45:39 SW - NE 203128.55 630655.80 205193.57 632202.98 target at 12:41:00

15/10/2008 Lamlash North 103 12:48:28 13:06:30 NE - SW 205232.11 632179.42 203150.83 630629.45 target at 12:58:40

15/10/2008 Lamlash North 106 13:09:23 13:25:45 SW - NE 203189.83 630478.75 205331.05 632046.20

15/10/2008 Lamlash North 109 13:27:49 13:45:52 NE - SW 205422.80 631962.59 203295.22 630362.89

15/10/2008 Lamlash North 105 14:08:11 14:26:46 NE - SW 205308.38 632104.77 203179.84 630515.30

15/10/2008 Deacon Rock 311 14:39:41 14:47:05 S - N 204477.94 630264.01 204428.27 631295.81

15/10/2008 Deacon Rock 310 14:49:18 14:57:17 N - S 204470.92 631325.99 204497.75 630120.39

15/10/2008 Deacon Rock 309 15:01:55 15:09:59 S - N 204546.26 630189.46 204527.97 631304.40 Reef at start of line

15/10/2008 Deacon Rock 308 15:13:24 15:21:56 N - S 204582.81 631245.76 204603.54 630048.36 Reef at end of line

15/10/2008 Deacon Rock 307 15:25:46 15:34:55 S - N 204663.36 630099.99 204638.03 631302.73 Depression at 15:28:00

15/10/2008 Deacon Rock 306 16:10:18 16:18:25 N - S 204681.52 631258.25 204694.78 630060.84

15/10/2008 Deacon Rock 305 16:22:31 16:31:19 S - N 204765.53 630022.84 204716.63 631315.63

15/10/2008 Deacon Rock 304 16:34:18 16:42:24 N - S 204763.54 631244.00 204796.50 630049.33

15/10/2008 Deacon Rock 303 16:47:41 16:55:38 S - N 204863.75 630162.92 204829.68 631311.54

15/10/2008 Deacon Rock 302 16:57:56 17:06:59 N - S 204870.83 631379.22 204894.00 630066.38

15/10/2008 Deacon Rock 301 17:11:21 17:19:07 S - N 204962.79 630144.87 204933.30 631325.42

Start of Line Position End of Line Position

Page 94: Mapping of the marine habitats and species of Lamlash Bay, Arran

82

16/10/2008 Lamlash South 208 06:46:22 07:16:25 NW - SE 203532.10 630785.77 206495.16 627851.42 target at 07:10:00

16/10/2008 Lamlash South 207 07:43:55 08:06:12 SE - NW 205533.31 628790.76 203500.21 630790.27 line shortened due to buoys

16/10/2008 Lamlash South 210 08:10:37 08:40:17 NW - SE 203564.01 630940.22 206327.55 628161.52

16/10/2008 Lamlash South 209 08:43:56 09:14:39 SE - NW 206406.40 628012.92 203613.47 630818.35

16/10/2008 Lamlash South 212 09:17:34 09:41:24 NW - SE 203732.85 630910.86 206006.49 628618.46

16/10/2008 Lamlash South 211 09:46:40 10:13:27 SE - NW 206038.35 628528.52 203653.84 630933.50

16/10/2008 Lamlash South 214 11:03:35 11:22:09 NW - SE 203985.56 630781.74 205859.74 628898.61

16/10/2008 Lamlash South 213 11:26:09 11:47:06 SE - NW 205855.63 628867.56 203812.36 630895.97

16/10/2008 Lamlash South 216 11:50:54 12:08:16 NW - SE 203910.15 631010.18 205672.62 629244.01

16/10/2008 Lamlash South 215 12:13:21 12:31:55 SE - NW 205594.35 629263.33 203815.08 631058.16

16/10/2008 Lamlash South 218 12:35:26 12:50:19 NW - SE 203932.13 631120.50 205524.98 629574.39

16/10/2008 Lamlash South 217 12:56:03 13:11:41 SE - NW 205362.85 629646.74 203903.28 631088.89

16/10/2008 Lamlash South 206 13:18:26 13:36:53 NW - SE 203474.48 630738.38 205552.18 628647.27

16/10/2008 Lamlash South 205 13:42:41 14:06:52 SE - NW 205514.02 628663.19 203458.14 630693.24

16/10/2008 Lamlash South 204 14:13:03 14:27:04 NW - SE 203642.48 630421.45 205249.66 628799.69 targets at 14:14 and 14:25:27

16/10/2008 Lamlash South 203 14:34:02 14:54:45 SE - NW 205243.19 628776.96 203480.76 630515.40

16/10/2008 Lamlash South 202 14:59:35 15:06:14 NW - SE 203491.30 630430.06 204220.54 629724.57 line cut short due to creel buoys

16/10/2008 Lamlash South 201 15:13:54 15:20:40 SE - NW 204130.62 629731.45 203519.18 630349.46 NW half only due to creel buoys

17/10/2008 Lamlash Outer 401 07:14:54 08:11:05 N - S 204919.88 635157.33 207195.53 627903.72 Off line at 07:43 due to creel buoys

17/10/2008 Lamlash Outer 402 08:17:23 09:12:23 S - N 207155.86 628009.40 204798.41 635258.53 off line at 08:30:30 due to creel buoys

17/10/2008 Lamlash Outer 403 09:30:00 09:54:09 N - S 204954.99 634910.54 206529.91 631926.13 southern area not run due to buoys

17/10/2008 Lamlash Outer 404 09:59:28 10:25:31 S - N 206486.28 631941.20 204721.98 635193.06 southern area not run due to buoys

17/10/2008 Lamlash Outer 405 11:09:31 11:35:18 N - S 204805.88 634973.89 206429.44 631884.35 southern area not run due to buoys

17/10/2008 Lamlash Outer 406 11:39:32 12:05:23 S - N 206384.55 631927.22 204637.78 635175.96 southern area not run due to buoys

17/10/2008 Lamlash Outer 407 12:28:35 12:50:11 N - S 204639.12 635039.19 206323.31 631899.78 target at 12:48:30

17/10/2008 Lamlash Outer 408 12:55:14 13:18:25 S - N 206185.77 632112.09 204537.68 635140.03 bedrock at 12:56:15 - sand waves 13:13:18

17/10/2008 Lamlash Outer 409 13:24:43 13:49:24 N - S 204511.94 635097.26 206228.70 631913.93

17/10/2008 Lamlash Outer 410 14:01:39 14:09:48 S - N 206168.76 631893.12 205852.92 632901.16

17/10/2008 Lamlash Outer 403a 16:01:30 16:12:14 N - S 206926.67 629647.41 207104.59 627911.37 southern end of line

Page 95: Mapping of the marine habitats and species of Lamlash Bay, Arran

83

Appendix 03: Lamlash Bay Day Grab log - 2008SeaStar Survey - Project J/08/151

No Take ZonePositions are OSGB36 East and North (m) and WGS84 Latitude and Longitude (ddmmss.sss)

Client SNH Job Number

Location Lamlash North Vessel

Survey Lamlash Bay Habitat Mapping Survey Date

Site Day Grab Time Date East North Latitude (N) Longitude (W) Depth Samples Taken

Name Number (GMT) (m) (m) ddmmss.sss ddmmss.sss (m)

NTZ NTZ03#01 16:20:50 20/10/2008 204567.54 631330.75 553218.679 -50591.838 30.0 1mm Biology & PSA

NTZ NTZ02#02 16:39:47 20/10/2008 204829.12 631505.80 553228.927 -50567.708 28.6 1mm Biology & PSA

NTZ NTZ01#01 16:51:10 20/10/2008 205054.22 631690.37 553239.274 -50547.237 27.5 1mm Biology & PSA

NTZ NTZ09#01 17:02:37 20/10/2008 204906.90 631987.57 553254.641 -50562.384 20.3 1mm Biology & PSA

NTZ NTZ04#01 17:17:46 20/10/2008 204389.00 631447.99 553224.860 -50608.735 26.9 1mm Biology & PSA

NTZ NTZ05#01 17:30:10 20/10/2008 204671.81 631138.08 553208.646 -50581.213 22.1 1mm Biology & PSA

NTZ NTZ06#03 17:43:47 20/10/2008 204801.07 630935.10 553198.133 -50567.949 15.6 1mm Biology & PSA

NTZ NTZ07#01 17:52:10 20/10/2008 205123.23 631421.11 553225.128 -50538.758 20.4 1mm Biology & PSA

NTZ NTZ08#02 18:03:30 20/10/2008 205466.08 631712.50 553241.085 -50507.473 19.7 1mm Biology & PSA

NTZ NTZ10#01 18:12:15 20/10/2008 205896.46 631879.62 553251.697 -50467.579 22.5 1mm Biology & PSA

J/08/151

SV Alma Na Mara

October 2008

Page 96: Mapping of the marine habitats and species of Lamlash Bay, Arran

84

Appendix 03: Lamlash Bay Video log - 2008SeaStar Survey - Project J/08/151

Positions are OSGB36 East and North (m) and WGS84 Latitude and Longitude (ddmmss.sss)

Client SNH Job Number

Location Lamlash Bay Vessel

Survey Lamlash Bay Habitat Mapping Survey Date

DATE Site Location SOL Time Video VHS miniDV Number Depth DVD

Name (GMT) Duration Tape No. Tape No. of Photos East (m) North (m) Latitude (N) Longitude (W) (m) Number

17/10/2008 Lamlash North LNC01 16:53:35 00:22:05 1 2 18 206292.53 632385.68 553279.693 -50432.552 30.0

18/10/2008 Lamlash North LNC02 09:55:46 00:24:28 2 5 27 205617.42 631833.26 553247.948 -50493.647 15.3

18/10/2008 Lamlash North LNC03 12:13:51 00:31:18 2 6 35 205727.72 631909.97 553252.317 -50483.775 11.3

18/10/2008 Lamlash North LNC04 08:00:30 00:30:03 1 3 50 204373.78 631566.61 553230.772 -50611.268 24.1

18/10/2008 Lamlash North LNC05 08:48:22 00:48:38 2 4 54 204418.83 631491.12 553226.884 -50606.708 25.4

18/10/2008 Lamlash North LNC07 16:30:24 00:49:02 4 10 37 204161.29 631210.44 553211.550 -50629.876 20.9

19/10/2008 Lamlash North LNC08 07:53:58 00:31:09 4 11 69 204334.51 630572.38 553177.566 -50610.155 27.6

18/10/2008 Lamlash North LNC09 13:04:00 00:24:56 3 7 21 205772.49 632083.22 553262.103 -50480.703 16.1

18/10/2008 Lamlash North LNC10 14:03:10 00:37:22 3 8 26 204565.25 631279.72 553215.609 -50591.322 29.4

18/10/2008 Lamlash North LNC11 14:58:24 00:40:51 3 9 23 204395.72 631726.73 553239.429 -50609.868 14.2

19/10/2008 Lamlash North LNC12 08:45:02 00:31:46 4 12 58 204487.88 630294.00 553162.384 -50594.774 25.6

16/10/2008 Lamlash South LSC01 17:21:03 00:35:05 1 1 61 204880.67 629699.26 553131.739 -50555.207 40.1

19/10/2008 Lamlash South LSC02 09:36:59 00:39:17 5 13 48 204015.72 629890.10 553139.489 -50637.824 32.3

20/10/2008 Lamlash South LSC03 10:07:42 00:33:47 6 16 37 204826.09 629033.76 553095.474 -50556.669 33.2

20/10/2008 Lamlash South LSC04 12:25:46 00:44:18 6 17 51 204670.41 629376.52 553113.860 -50572.877 39.4

20/10/2008 Lamlash South LSC05 13:23:45 00:20:35 7 18 19 204325.71 629665.97 553128.432 -50606.757 38.9

20/10/2008 Lamlash South LSC06 13:58:48 00:14:28 7 18 31 204487.90 630075.81 553151.069 -50594.083 37.6

20/10/2008 Lamlash South LSC07 14:36:37 00:31:05 7 19 47 203777.05 630388.15 553165.920 -50662.918 34.1

20/10/2008 Lamlash Outer LOC01 07:17:56 00:45:43 5 14 79 204815.50 634916.60 553412.574 -50583.216 30.3

20/10/2008 Lamlash Outer LOC02 08:18:30 00:14:16 5 15 19 205562.61 633477.79 553336.701 -50506.507 22.8

20/10/2008 Lamlash Outer LOC03 08:45:25 00:15:26 5 15 34 No Navigation Data Recorded

20/10/2008 Lamlash Outer LOC04 09:17:16 00:09:13 6 16 25 205943.17 632319.34 553274.919 -50465.389 20.2

J/08/151

SV Alma Na Mara

October 2008

Start Of Line Position Start Of Line Position

Page 97: Mapping of the marine habitats and species of Lamlash Bay, Arran

85

Appendix 03: Lamlash Bay Still Photographs log - 2008SeaStar Survey - Project J/08/151

Lamlash NorthPositions are OSGB36 East and North (m) and WGS84 Latitude and Longitude (ddmmss.sss)

Client SNH Job NumberLocation Lamlash North VesselSurvey Lamlash Bay Habitat Mapping Survey Date

Site Photograph Time Date East North Latitude (N) Longitude (W) Depth CommentsName Number (GMT) (m) (m) ddmmss.sss ddmmss.sss (m)

LNC01 2 16:54:45 17/10/2008 206305.77 632380.92 553279.482 -50431.263 36.3 Depth correct to below water surfaceLNC01 3 16:55:25 17/10/2008 206312.59 632380.51 553279.373 -50430.661 37.1

LNC01 4 16:56:14 17/10/2008 206325.82 632375.75 553279.208 -50429.357 38.2

LNC01 5 16:57:54 17/10/2008 206351.14 632367.22 553278.830 -50426.916 40.0

LNC01 6 16:58:29 17/10/2008 206360.18 632364.42 553278.702 -50426.045 40.8

LNC01 7 16:59:10 17/10/2008 206370.74 632361.95 553278.568 -50425.046 41.5

LNC01 8 17:00:30 17/10/2008 206390.79 632361.04 553278.470 -50423.160 42.8

LNC01 9 17:01:00 17/10/2008 206398.06 632359.42 553278.459 -50422.460 43.2

LNC01 10 17:01:25 17/10/2008 206403.62 632358.14 553278.441 -50421.917 43.6

LNC01 11 17:03:01 17/10/2008 206424.92 632359.90 553278.506 -50419.918 44.7

LNC01 12 17:07:24 17/10/2008 206487.80 632365.72 553278.979 -50413.976 46.0

LNC01 13 17:10:19 17/10/2008 206526.35 632366.11 553279.097 -50410.314 45.0

LNC01 14 17:10:46 17/10/2008 206531.69 632366.55 553279.122 -50409.802 44.8

LNC01 15 17:13:05 17/10/2008 206561.23 632371.33 553279.398 -50407.011 43.2

LNC01 16 17:13:40 17/10/2008 206568.57 632372.38 553279.486 -50406.334 42.7

LNC01 17 17:15:13 17/10/2008 206586.65 632376.43 553279.723 -50404.609 41.5

LNC01 18 17:15:22 17/10/2008 206588.56 632376.98 553279.753 -50404.427 41.5

LNC02 2 09:56:17 18/10/2008 205627.54 631829.26 553248.016 -50493.397 19.7

LNC02 3 09:56:49 18/10/2008 205636.11 631830.37 553248.103 -50492.591 19.0

LNC02 4 09:57:02 18/10/2008 205639.60 631831.14 553248.139 -50492.262 18.7

LNC02 5 09:57:42 18/10/2008 205650.05 631832.53 553248.280 -50491.276 17.9

LNC02 6 09:58:20 18/10/2008 205659.04 631836.30 553248.415 -50490.424 17.2

LNC02 7 09:58:46 18/10/2008 205665.29 631837.37 553248.566 -50489.850 16.6

LNC02 8 09:58:53 18/10/2008 205666.66 631838.93 553248.598 -50489.717 16.4

LNC02 9 09:59:14 18/10/2008 205671.12 631840.72 553248.643 -50489.273 16.2

LNC02 10 09:59:37 18/10/2008 205675.61 631842.24 553248.757 -50488.863 15.9

LNC02 11 09:59:58 18/10/2008 205680.28 631845.86 553248.928 -50488.409 14.9

LNC02 12 10:00:23 18/10/2008 205685.52 631849.22 553249.109 -50487.917 15.5

LNC02 13 10:00:43 18/10/2008 205689.75 631852.23 553249.272 -50487.521 15.3

LNC02 14 10:01:47 18/10/2008 205702.44 631862.72 553249.845 -50486.339 15.1

LNC02 15 10:02:50 18/10/2008 205714.67 631872.52 553250.404 -50485.223 14.9

LNC02 16 10:02:59 18/10/2008 205716.40 631874.12 553250.490 -50485.054 14.8

LNC02 17 10:03:09 18/10/2008 205718.80 631875.54 553250.584 -50484.847 14.9

LNC02 18 10:04:06 18/10/2008 205729.83 631883.73 553251.067 -50483.856 15.2

LNC02 19 10:05:00 18/10/2008 205739.81 631889.78 553251.426 -50482.941 15.7

LNC02 20 10:05:10 18/10/2008 205741.69 631890.75 553251.488 -50482.769 15.8

J/08/151SV Alma Na MaraOctober 2008

Page 98: Mapping of the marine habitats and species of Lamlash Bay, Arran

86

LNC02 21 10:06:41 18/10/2008 205758.41 631901.38 553252.093 -50481.222 17.3

LNC02 22 10:08:17 18/10/2008 205779.44 631910.61 553252.685 -50479.294 19.4

LNC02 23 10:09:24 18/10/2008 205795.88 631913.70 553252.948 -50477.770 21.0

LNC02 24 10:11:56 18/10/2008 205836.40 631911.66 553253.046 -50473.904 23.8

LNC02 25 10:13:20 18/10/2008 205858.99 631908.21 553252.907 -50471.746 24.7

LNC02 26 10:15:15 18/10/2008 205898.90 631902.23 553252.682 -50467.933 26.6

LNC02 27 10:18:21 18/10/2008 205970.53 631892.07 553252.302 -50461.081 31.2

LNC03 2 12:14:46 18/10/2008 205607.62 631948.96 553254.382 -50495.789 20.8

LNC03 3 12:15:09 18/10/2008 205613.43 631950.13 553254.470 -50495.247 20.4

LNC03 4 12:15:44 18/10/2008 205621.62 631952.12 553254.591 -50494.476 19.7

LNC03 5 12:16:29 18/10/2008 205631.50 631954.75 553254.751 -50493.544 18.7

LNC03 6 12:17:13 18/10/2008 205642.51 631957.01 553254.910 -50492.514 18.1

LNC03 7 12:17:51 18/10/2008 205653.36 631958.41 553255.032 -50491.495 17.3

LNC03 8 12:18:57 18/10/2008 205670.55 631958.32 553255.121 -50489.857 16.2

LNC03 9 12:19:08 18/10/2008 205673.19 631957.74 553255.116 -50489.599 16.4

LNC03 10 12:20:19 18/10/2008 205688.89 631956.62 553255.075 -50488.109 15.8

LNC03 11 12:20:51 18/10/2008 205695.91 631957.31 553255.101 -50487.450 15.6

LNC03 12 12:21:11 18/10/2008 205700.34 631956.62 553255.106 -50487.028 15.3

LNC03 13 12:21:35 18/10/2008 205705.69 631957.95 553255.142 -50486.526 15.3

LNC03 14 12:22:07 18/10/2008 205713.17 631959.11 553255.199 -50485.817 15.5

LNC03 15 12:23:08 18/10/2008 205727.77 631964.35 553255.489 -50484.440 16.2

LNC03 16 12:23:26 18/10/2008 205732.45 631965.20 553255.558 -50484.008 17.0

LNC03 17 12:23:43 18/10/2008 205736.95 631966.41 553255.629 -50483.577 17.4

LNC03 18 12:24:16 18/10/2008 205745.65 631968.03 553255.746 -50482.764 18.4

LNC03 19 12:24:48 18/10/2008 205754.63 631969.78 553255.865 -50481.920 19.3

LNC03 20 12:25:20 18/10/2008 205764.35 631971.60 553255.987 -50481.007 20.2

LNC03 21 12:26:11 18/10/2008 205781.37 631975.32 553256.225 -50479.403 21.7

LNC03 22 12:26:30 18/10/2008 205787.98 631976.07 553256.300 -50478.790 22.1

LNC03 23 12:27:54 18/10/2008 205816.86 631976.56 553256.435 -50476.047 24.2

LNC03 24 12:28:28 18/10/2008 205829.22 631978.32 553256.524 -50474.879 24.8

LNC03 25 12:29:59 18/10/2008 205855.33 631969.35 553256.195 -50472.342 25.6

LNC03 26 12:30:45 18/10/2008 205867.36 631968.45 553256.143 -50471.217 25.7

LNC03 27 12:31:23 18/10/2008 205875.88 631966.16 553256.086 -50470.380 26.2

LNC03 28 12:32:55 18/10/2008 205899.43 631969.83 553256.257 -50468.177 27.3

LNC03 29 12:33:48 18/10/2008 205915.24 631970.25 553256.276 -50466.673 28.3

LNC03 30 12:34:26 18/10/2008 205926.07 631971.83 553256.394 -50465.652 29.0

LNC03 31 12:35:47 18/10/2008 205948.35 631976.05 553256.659 -50463.553 30.1

LNC03 32 12:37:47 18/10/2008 205979.07 631980.77 553257.014 -50460.659 30.9

LNC03 33 12:38:06 18/10/2008 205983.60 631980.46 553257.037 -50460.239 31.0

LNC03 34 12:40:54 18/10/2008 206027.22 631989.33 553257.545 -50456.110 31.4

LNC03 35 12:42:48 18/10/2008 206061.68 631995.90 553258.030 -50452.883 30.9

LNC04 2 08:02:13 18/10/2008 204449.87 631599.84 553232.724 -50604.166 30.1

LNC04 3 08:02:43 18/10/2008 204464.22 631609.89 553233.275 -50602.823 30.1

LNC04 4 08:03:26 18/10/2008 204487.17 631624.55 553234.131 -50600.718 29.6

Page 99: Mapping of the marine habitats and species of Lamlash Bay, Arran

87

LNC04 5 08:04:16 18/10/2008 204515.91 631636.43 553234.897 -50598.087 27.0

LNC04 6 08:04:43 18/10/2008 204528.74 631645.70 553235.410 -50596.899 24.7

LNC04 7 08:04:57 18/10/2008 204536.21 631648.87 553235.603 -50596.205 23.1

LNC04 8 08:05:08 18/10/2008 204542.77 631650.52 553235.721 -50595.600 22.0

LNC04 9 08:05:45 18/10/2008 204563.05 631658.06 553236.176 -50593.706 19.0

LNC04 10 08:06:01 18/10/2008 204571.86 631661.92 553236.388 -50592.875 18.6

LNC04 11 08:06:20 18/10/2008 204583.29 631665.95 553236.655 -50591.824 18.1

LNC04 12 08:07:05 18/10/2008 204604.67 631681.89 553237.480 -50589.789 18.0

LNC04 13 08:07:16 18/10/2008 204609.03 631685.90 553237.696 -50589.376 18.1

LNC04 14 08:07:29 18/10/2008 204614.51 631690.29 553237.945 -50588.873 18.1

LNC04 15 08:08:15 18/10/2008 204634.88 631706.82 553238.875 -50587.000 18.9

LNC04 16 08:08:30 18/10/2008 204641.55 631712.57 553239.204 -50586.393 19.1

LNC04 17 08:08:47 18/10/2008 204649.63 631718.43 553239.554 -50585.672 19.3

LNC04 18 08:09:27 18/10/2008 204668.82 631730.71 553240.264 -50583.906 19.7

LNC04 19 08:09:40 18/10/2008 204675.45 631733.87 553240.476 -50583.316 19.9

LNC04 20 08:10:13 18/10/2008 204691.57 631741.45 553240.949 -50581.840 20.2

LNC04 21 08:10:49 18/10/2008 204707.66 631748.32 553241.371 -50580.353 20.6

LNC04 22 08:11:27 18/10/2008 204723.59 631755.85 553241.803 -50578.863 20.9

LNC04 23 08:11:52 18/10/2008 204734.44 631760.96 553242.100 -50577.853 21.2

LNC04 24 08:12:15 18/10/2008 204744.67 631766.73 553242.417 -50576.892 21.4

LNC04 25 08:12:34 18/10/2008 204753.62 631771.54 553242.703 -50576.064 21.8

LNC04 26 08:13:20 18/10/2008 204773.84 631782.49 553243.335 -50574.187 22.6

LNC04 27 08:13:46 18/10/2008 204785.79 631787.34 553243.651 -50573.095 23.0

LNC04 28 08:14:46 18/10/2008 204814.63 631798.87 553244.349 -50570.411 24.2

LNC04 29 08:15:30 18/10/2008 204835.93 631806.44 553244.825 -50568.432 25.1

LNC04 30 08:16:10 18/10/2008 204854.01 631811.90 553245.178 -50566.746 25.5

LNC04 31 08:17:39 18/10/2008 204892.90 631829.48 553246.175 -50563.104 25.9

LNC04 32 08:19:07 18/10/2008 204931.91 631845.97 553247.167 -50559.478 26.2

LNC04 33 08:20:49 18/10/2008 204971.62 631868.50 553248.427 -50555.763 26.3

LNC04 34 08:22:07 18/10/2008 205000.00 631887.76 553249.529 -50553.149 26.0

LNC04 35 08:23:20 18/10/2008 205027.23 631908.11 553250.677 -50550.634 25.6

LNC04 36 08:24:09 18/10/2008 205046.41 631925.10 553251.627 -50548.873 25.1

LNC04 37 08:24:27 18/10/2008 205053.96 631930.26 553251.941 -50548.197 24.8

LNC04 38 08:24:46 18/10/2008 205062.15 631934.83 553252.202 -50547.438 24.4

LNC04 39 08:25:10 18/10/2008 205072.08 631941.79 553252.605 -50546.524 23.7

LNC04 40 08:25:23 18/10/2008 205077.61 631945.55 553252.821 -50546.017 23.3

LNC04 41 08:25:49 18/10/2008 205088.43 631953.44 553253.269 -50545.022 22.1

LNC04 42 08:26:11 18/10/2008 205097.63 631959.77 553253.635 -50544.178 21.1

LNC04 43 08:26:29 18/10/2008 205104.93 631964.48 553253.912 -50543.513 20.6

LNC04 44 08:26:49 18/10/2008 205113.25 631969.93 553254.223 -50542.746 20.1

LNC04 45 08:27:10 18/10/2008 205122.56 631975.78 553254.566 -50541.889 19.6

LNC04 46 08:27:23 18/10/2008 205128.38 631979.89 553254.789 -50541.341 19.2

LNC04 47 08:27:58 18/10/2008 205144.16 631989.61 553255.361 -50539.899 18.6

LNC04 48 08:28:20 18/10/2008 205154.69 631995.48 553255.716 -50538.932 18.6

LNC04 49 08:30:09 18/10/2008 205205.81 632019.55 553257.164 -50534.205 20.2

Page 100: Mapping of the marine habitats and species of Lamlash Bay, Arran

88

LNC04 50 08:30:24 18/10/2008 205212.61 632022.37 553257.335 -50533.572 20.5

LNC05 2 08:49:22 18/10/2008 204449.10 631498.37 553227.306 -50603.848 31.2

LNC05 3 08:51:04 18/10/2008 204482.08 631516.75 553228.341 -50600.769 31.5

LNC05 4 08:52:27 18/10/2008 204513.95 631531.11 553229.211 -50597.819 31.5

LNC05 5 08:54:03 18/10/2008 204551.23 631544.21 553230.043 -50594.357 31.2

LNC05 6 08:56:12 18/10/2008 204604.02 631563.86 553231.198 -50589.415 29.6

LNC05 7 08:58:30 18/10/2008 204666.53 631582.66 553232.406 -50583.581 26.0

LNC05 8 08:58:57 18/10/2008 204677.27 631585.57 553232.583 -50582.572 26.0

LNC05 9 08:59:10 18/10/2008 204682.65 631586.63 553232.661 -50582.070 26.0

LNC05 10 09:00:50 18/10/2008 204722.51 631601.03 553233.449 -50578.299 26.4

LNC05 11 09:02:01 18/10/2008 204750.36 631620.03 553234.513 -50575.710 26.5

LNC05 12 09:02:33 18/10/2008 204762.98 631627.97 553234.981 -50574.554 26.5

LNC05 13 09:04:36 18/10/2008 204814.16 631653.94 553236.539 -50569.837 26.9

LNC05 14 09:07:02 18/10/2008 204871.25 631678.64 553237.995 -50564.517 27.4

LNC05 15 09:09:40 18/10/2008 204916.46 631712.66 553239.863 -50560.289 27.7

LNC05 16 09:11:01 18/10/2008 204934.98 631732.50 553240.952 -50558.577 27.8

LNC05 17 09:13:16 18/10/2008 204977.47 631763.68 553242.816 -50554.781 28.4

LNC05 18 09:14:29 18/10/2008 204999.78 631774.12 553243.449 -50552.722 28.8

LNC05 19 09:14:29 18/10/2008 204999.78 631774.12 553243.449 -50552.722 28.8

LNC05 20 09:15:57 18/10/2008 205025.90 631788.95 553244.294 -50550.291 29.2

LNC05 21 09:16:53 18/10/2008 205040.87 631797.59 553244.788 -50548.902 29.3

LNC05 22 09:18:32 18/10/2008 205070.02 631815.16 553245.808 -50546.212 28.9

LNC05 23 09:19:26 18/10/2008 205088.96 631824.90 553246.388 -50544.463 28.4

LNC05 24 09:20:05 18/10/2008 205101.78 631828.17 553246.751 -50543.310 27.7

LNC05 25 09:20:45 18/10/2008 205113.79 631837.58 553247.128 -50542.163 26.8

LNC05 26 09:21:14 18/10/2008 205122.51 631843.16 553247.435 -50541.344 25.9

LNC05 27 09:21:42 18/10/2008 205131.64 631848.32 553247.744 -50540.508 24.7

LNC05 28 09:22:11 18/10/2008 205140.76 631853.42 553248.041 -50539.664 23.6

LNC05 29 09:22:49 18/10/2008 205152.53 631859.86 553248.417 -50538.573 22.8

LNC05 30 09:23:54 18/10/2008 205174.71 631871.58 553249.104 -50536.521 21.6

LNC05 31 09:24:13 18/10/2008 205181.78 631875.38 553249.326 -50535.865 21.5

LNC05 32 09:24:57 18/10/2008 205199.23 631885.18 553249.890 -50534.244 20.9

LNC05 33 09:25:24 18/10/2008 205210.07 631890.29 553250.202 -50533.246 20.8

LNC05 34 09:25:56 18/10/2008 205224.43 631897.23 553250.598 -50531.902 20.7

LNC05 35 09:26:22 18/10/2008 205235.80 631899.32 553250.813 -50530.876 20.5

LNC05 36 09:26:42 18/10/2008 205244.04 631903.83 553251.028 -50530.088 20.5

LNC05 37 09:27:21 18/10/2008 205259.23 631910.16 553251.406 -50528.675 20.3

LNC05 38 09:27:39 18/10/2008 205265.72 631914.16 553251.609 -50528.055 20.2

LNC05 39 09:28:13 18/10/2008 205280.11 631922.33 553252.075 -50526.714 20.0

LNC05 40 09:28:50 18/10/2008 205296.51 631929.77 553252.539 -50525.206 19.4

LNC05 41 09:29:07 18/10/2008 205304.16 631932.83 553252.734 -50524.507 19.2

LNC05 42 09:29:29 18/10/2008 205313.92 631937.07 553252.984 -50523.595 18.9

LNC05 43 09:30:05 18/10/2008 205329.64 631943.00 553253.353 -50522.134 18.6

LNC05 44 09:30:29 18/10/2008 205340.19 631947.50 553253.615 -50521.147 18.5

LNC05 45 09:31:17 18/10/2008 205361.24 631957.16 553254.166 -50519.179 19.0

LNC05 46 09:31:30 18/10/2008 205366.85 631960.08 553254.334 -50518.659 19.3

Page 101: Mapping of the marine habitats and species of Lamlash Bay, Arran

89

LNC05 47 09:31:58 18/10/2008 205378.55 631965.76 553254.678 -50517.580 19.8

LNC05 48 09:32:53 18/10/2008 205400.58 631977.87 553255.355 -50515.515 20.9

LNC05 49 09:33:30 18/10/2008 205415.97 631987.84 553255.927 -50514.096 21.4

LNC05 50 09:33:58 18/10/2008 205427.83 631994.72 553256.330 -50513.003 21.8

LNC05 51 09:34:39 18/10/2008 205443.62 632006.05 553256.949 -50511.518 22.1

LNC05 52 09:35:39 18/10/2008 205466.26 632020.67 553257.818 -50509.464 22.2

LNC05 53 09:35:54 18/10/2008 205471.98 632024.31 553258.022 -50508.928 22.1

LNC07 2 16:32:07 18/10/2008 204286.07 631067.08 553203.856 -50617.531 25.5

LNC07 3 16:33:26 18/10/2008 204306.69 631069.21 553203.959 -50615.599 25.1

LNC07 4 16:35:30 18/10/2008 204333.43 631066.13 553203.942 -50613.020 24.5

LNC07 5 16:35:57 18/10/2008 204336.93 631064.19 553203.880 -50612.655 24.4

LNC07 6 16:46:42 18/10/2008 204322.88 631026.08 553201.969 -50613.209 25.6

LNC07 7 16:47:56 18/10/2008 204331.16 631015.40 553201.395 -50612.727 26.0

LNC07 8 16:51:01 18/10/2008 204352.77 631009.72 553200.924 -50610.946 25.9

LNC07 9 16:53:19 18/10/2008 204363.01 631002.71 553200.606 -50609.924 25.5

LNC07 10 16:55:18 18/10/2008 204370.07 630997.97 553200.536 -50609.003 25.1

LNC07 11 16:56:29 18/10/2008 204377.49 630996.79 553200.363 -50608.502 24.9

LNC07 12 16:58:25 18/10/2008 204381.92 630988.97 553200.070 -50607.881 24.7

LNC07 13 17:00:56 18/10/2008 204386.14 630957.84 553198.469 -50607.020 25.3

LNC07 14 17:01:45 18/10/2008 204400.25 630945.92 553197.788 -50605.999 25.6

LNC07 15 17:02:46 18/10/2008 204412.60 630929.09 553196.936 -50604.718 25.2

LNC07 16 17:03:36 18/10/2008 204426.88 630918.17 553196.304 -50603.457 24.7

LNC07 17 17:04:06 18/10/2008 204431.10 630909.10 553195.895 -50602.881 24.2

LNC07 18 17:04:50 18/10/2008 204444.60 630899.77 553195.399 -50601.618 23.3

LNC07 19 17:05:26 18/10/2008 204453.10 630891.14 553194.930 -50600.824 22.5

LNC07 20 17:07:34 18/10/2008 204469.08 630851.35 553192.847 -50599.100 23.5

LNC07 21 17:08:04 18/10/2008 204476.93 630841.80 553192.372 -50598.296 23.3

LNC07 22 17:08:41 18/10/2008 204482.75 630829.73 553191.807 -50597.400 22.8

LNC07 23 17:09:08 18/10/2008 204488.25 630821.52 553191.345 -50597.010 22.7

LNC07 24 17:09:57 18/10/2008 204498.71 630806.21 553190.555 -50595.920 22.1

LNC07 25 17:10:54 18/10/2008 204501.97 630790.53 553189.735 -50595.344 21.7

LNC07 26 17:11:24 18/10/2008 204502.00 630783.31 553189.348 -50595.268 21.4

LNC07 27 17:11:46 18/10/2008 204501.05 630778.74 553189.098 -50595.271 21.9

LNC07 28 17:11:59 18/10/2008 204500.57 630776.10 553188.954 -50595.284 22.5

LNC07 29 17:12:37 18/10/2008 204498.85 630769.36 553188.576 -50595.377 24.6

LNC07 30 17:13:34 18/10/2008 204496.30 630760.55 553188.087 -50595.553 26.1

LNC07 31 17:13:42 18/10/2008 204495.98 630759.38 553188.024 -50595.576 26.4

LNC07 32 17:14:05 18/10/2008 204495.06 630756.12 553187.845 -50595.642 26.6

LNC07 33 17:14:20 18/10/2008 204494.41 630754.23 553187.738 -50595.686 26.7

LNC07 34 17:15:10 18/10/2008 204492.54 630748.22 553187.407 -50595.823 27.5

LNC07 35 17:15:45 18/10/2008 204491.34 630744.44 553187.199 -50595.925 27.6

LNC07 36 17:17:49 18/10/2008 204487.59 630733.03 553186.580 -50596.252 28.4

LNC07 37 17:18:20 18/10/2008 204487.13 630730.88 553186.465 -50596.293 28.5

LNC08 2 07:54:28 19/10/2008 204424.10 630678.89 553182.936 -50602.424 31.77

LNC08 3 07:55:30 19/10/2008 204433.75 630703.82 553184.320 -50601.681 30.8

LNC08 4 07:55:59 19/10/2008 204438.76 630714.84 553184.923 -50601.225 30.34

Page 102: Mapping of the marine habitats and species of Lamlash Bay, Arran

90

LNC08 5 07:56:55 19/10/2008 204446.20 630735.15 553186.022 -50600.574 29.8

LNC08 6 07:57:39 19/10/2008 204450.80 630750.39 553186.859 -50600.202 29.27

LNC08 7 07:58:53 19/10/2008 204459.15 630778.89 553188.411 -50599.529 28.02

LNC08 8 07:59:44 19/10/2008 204466.07 630797.74 553189.453 -50598.993 26.64

LNC08 9 08:00:03 19/10/2008 204467.59 630805.33 553189.851 -50598.827 26.06

LNC08 10 08:00:15 19/10/2008 204468.70 630810.35 553190.121 -50598.725 25.71

LNC08 11 08:00:27 19/10/2008 204469.59 630814.85 553190.359 -50598.630 25.28

LNC08 12 08:01:02 19/10/2008 204473.62 630828.72 553191.119 -50598.340 24.43

LNC08 13 08:01:13 19/10/2008 204476.86 630832.18 553191.350 -50598.183 24.01

LNC08 14 08:01:33 19/10/2008 204477.74 630841.00 553191.805 -50598.049 23.39

LNC08 15 08:01:49 19/10/2008 204479.45 630846.97 553192.121 -50597.871 22.91

LNC08 16 08:02:22 19/10/2008 204483.19 630861.56 553192.908 -50597.567 21.21

LNC08 17 08:02:30 19/10/2008 204484.09 630865.10 553193.107 -50597.494 20.85

LNC08 18 08:02:42 19/10/2008 204485.79 630870.42 553193.400 -50597.368 20

LNC08 19 08:02:51 19/10/2008 204487.07 630874.60 553193.629 -50597.263 19.47

LNC08 20 08:02:59 19/10/2008 204488.54 630878.43 553193.840 -50597.157 18.41

LNC08 21 08:03:10 19/10/2008 204490.76 630883.65 553194.132 -50596.991 17.25

LNC08 22 08:03:27 19/10/2008 204494.98 630891.81 553194.589 -50596.678 15.28

LNC08 23 08:03:37 19/10/2008 204497.22 630896.40 553194.848 -50596.485 14.33

LNC08 24 08:03:50 19/10/2008 204500.91 630901.69 553195.157 -50596.196 13.28

LNC08 25 08:03:58 19/10/2008 204502.72 630904.34 553195.305 -50596.045 13.07

LNC08 26 08:04:10 19/10/2008 204505.70 630908.70 553195.550 -50595.788 13.26

LNC08 27 08:04:25 19/10/2008 204509.72 630913.88 553195.848 -50595.447 13.75

LNC08 28 08:04:33 19/10/2008 204511.87 630916.42 553195.995 -50595.268 14.4

LNC08 29 08:04:45 19/10/2008 204515.01 630920.57 553196.224 -50594.995 15.02

LNC08 30 08:05:00 19/10/2008 204518.05 630925.33 553196.476 -50594.714 15.33

LNC08 31 08:05:13 19/10/2008 204520.03 630929.99 553196.718 -50594.497 15.78

LNC08 32 08:05:37 19/10/2008 204523.69 630938.53 553197.172 -50594.150 16.36

LNC08 33 08:06:07 19/10/2008 204527.64 630949.31 553197.758 -50593.796 16.84

LNC08 34 08:06:24 19/10/2008 204530.48 630955.39 553198.094 -50593.575 16.98

LNC08 35 08:06:56 19/10/2008 204535.18 630967.68 553198.763 -50593.153 17.24

LNC08 36 08:07:09 19/10/2008 204536.96 630972.93 553199.043 -50592.994 17.35

LNC08 37 08:07:38 19/10/2008 204540.98 630984.92 553199.701 -50592.659 17.69

LNC08 38 08:07:54 19/10/2008 204543.26 630991.49 553200.064 -50592.464 17.92

LNC08 39 08:08:07 19/10/2008 204545.30 630997.21 553200.374 -50592.296 18.37

LNC08 40 08:08:25 19/10/2008 204548.42 631004.96 553200.800 -50592.052 18.69

LNC08 41 08:08:44 19/10/2008 204551.69 631013.35 553201.260 -50591.778 19.24

LNC08 42 08:08:55 19/10/2008 204553.41 631018.25 553201.531 -50591.621 19.21

LNC08 43 08:09:07 19/10/2008 204556.01 631023.79 553201.837 -50591.419 19.56

LNC08 44 08:09:23 19/10/2008 204559.06 631031.30 553202.249 -50591.152 20.01

LNC08 45 08:09:35 19/10/2008 204561.84 631036.73 553202.552 -50590.936 20.28

LNC08 46 08:09:48 19/10/2008 204564.86 631042.45 553202.872 -50590.689 20.55

LNC08 47 08:10:28 19/10/2008 204573.73 631061.72 553203.932 -50589.927 21.77

LNC08 48 08:10:53 19/10/2008 204580.58 631071.59 553204.498 -50589.378 22.32

LNC08 49 08:11:29 19/10/2008 204589.91 631086.53 553205.324 -50588.546 23.6

LNC08 50 08:12:34 19/10/2008 204606.84 631114.39 553206.849 -50587.005 25.17

Page 103: Mapping of the marine habitats and species of Lamlash Bay, Arran

91

LNC08 51 08:13:29 19/10/2008 204617.97 631142.95 553208.394 -50586.026 26.42

LNC08 52 08:13:55 19/10/2008 204622.62 631155.96 553209.106 -50585.630 26.99

LNC08 53 08:15:02 19/10/2008 204635.76 631187.10 553210.822 -50584.550 27.75

LNC08 54 08:16:13 19/10/2008 204651.42 631221.26 553212.704 -50583.213 27.84

LNC08 55 08:16:32 19/10/2008 204657.24 631229.85 553213.204 -50582.767 27.73

LNC08 56 08:17:21 19/10/2008 204671.79 631251.60 553214.422 -50581.495 26.94

LNC08 57 08:18:30 19/10/2008 204695.50 631282.75 553216.147 -50579.369 25.91

LNC08 58 08:18:58 19/10/2008 204704.74 631295.95 553216.878 -50578.535 25.73

LNC08 59 08:19:44 19/10/2008 204718.98 631317.79 553218.084 -50577.271 25.89

LNC08 60 08:19:54 19/10/2008 204722.19 631322.53 553218.347 -50576.990 25.94

LNC08 61 08:20:26 19/10/2008 204733.17 631336.72 553219.154 -50576.045 26.09

LNC08 62 08:21:16 19/10/2008 204751.24 631357.38 553220.328 -50574.451 26.44

LNC08 63 08:21:42 19/10/2008 204760.74 631367.68 553220.905 -50573.589 26.64

LNC08 64 08:21:52 19/10/2008 204764.48 631371.71 553221.130 -50573.251 26.72

LNC08 65 08:22:23 19/10/2008 204776.08 631383.85 553221.813 -50572.203 26.87

LNC08 66 08:22:56 19/10/2008 204788.33 631396.95 553222.546 -50571.100 27.02

LNC08 67 08:23:14 19/10/2008 204794.36 631404.20 553222.943 -50570.540 27.28

LNC08 68 08:23:32 19/10/2008 204800.23 631412.28 553223.379 -50569.989 27.61

LNC08 69 08:23:54 19/10/2008 204807.82 631422.21 553223.922 -50569.292 28

LNC09 2 13:04:48 18/10/2008 205785.40 632079.89 553262.014 -50479.437 22.0

LNC09 3 13:05:16 18/10/2008 205789.52 632078.00 553261.942 -50479.017 22.3

LNC09 4 13:06:14 18/10/2008 205800.27 632070.96 553261.643 -50477.916 23.0

LNC09 5 13:06:58 18/10/2008 205808.49 632066.54 553261.411 -50477.123 23.4

LNC09 6 13:07:45 18/10/2008 205817.67 632063.49 553261.223 -50476.290 23.7

LNC09 7 13:08:34 18/10/2008 205831.79 632061.37 553261.097 -50474.968 24.0

LNC09 8 13:10:33 18/10/2008 205865.51 632051.91 553260.682 -50471.726 24.8

LNC09 9 13:10:56 18/10/2008 205872.16 632051.21 553260.633 -50471.090 25.1

LNC09 10 13:11:16 18/10/2008 205878.01 632049.92 553260.586 -50470.532 25.3

LNC09 11 13:12:12 18/10/2008 205895.91 632043.51 553260.344 -50468.764 26.3

LNC09 12 13:12:40 18/10/2008 205904.01 632038.04 553260.107 -50467.937 27.0

LNC09 13 13:13:36 18/10/2008 205918.39 632026.59 553259.552 -50466.487 28.4

LNC09 14 13:14:00 18/10/2008 205923.24 632022.26 553259.339 -50465.991 28.8

LNC09 15 13:15:09 18/10/2008 205939.31 632014.90 553258.928 -50464.517 29.7

LNC09 16 13:20:45 18/10/2008 206023.73 631982.98 553257.355 -50456.399 31.8

LNC09 17 13:21:29 18/10/2008 206035.03 631978.52 553257.163 -50455.287 31.9

LNC09 18 13:23:26 18/10/2008 206065.72 631967.09 553256.598 -50452.357 31.8

LNC09 19 13:25:44 18/10/2008 206104.60 631963.45 553256.419 -50448.660 31.3

LNC09 20 13:26:32 18/10/2008 206117.66 631963.61 553256.448 -50447.429 30.9

LNC09 21 13:28:43 18/10/2008 206156.28 631962.31 553256.514 -50443.745 30.2

LNC10 1 14:04:50 18/10/2008 204591.70 631308.03 553217.412 -50589.511 34.7

LNC10 2 14:07:26 18/10/2008 204614.50 631314.91 553217.878 -50587.396 33.6

LNC10 3 14:07:56 18/10/2008 204619.93 631316.54 553217.968 -50586.875 33.2

LNC10 4 14:08:07 18/10/2008 204622.32 631316.94 553218.001 -50586.658 32.9

LNC10 5 14:08:35 18/10/2008 204628.49 631317.73 553218.078 -50586.077 32.6

LNC10 6 14:09:50 18/10/2008 204645.85 631320.43 553218.262 -50584.440 31.7

LNC10 7 14:16:45 18/10/2008 204766.32 631322.02 553218.705 -50573.012 27.3

Page 104: Mapping of the marine habitats and species of Lamlash Bay, Arran

92

LNC10 8 14:18:05 18/10/2008 204787.93 631321.24 553218.706 -50570.954 26.5

LNC10 9 14:18:58 18/10/2008 204801.37 631320.67 553218.702 -50569.680 26.2

LNC10 10 14:21:43 18/10/2008 204839.79 631319.69 553218.747 -50566.027 25.9

LNC10 11 14:22:20 18/10/2008 204848.59 631318.69 553218.736 -50565.185 25.8

LNC10 12 14:23:27 18/10/2008 204862.75 631314.71 553218.613 -50563.796 25.8

LNC10 13 14:24:14 18/10/2008 204872.61 631313.29 553218.515 -50562.876 25.8

LNC10 14 14:25:58 18/10/2008 204895.53 631315.09 553218.629 -50560.719 26.2

LNC10 15 14:28:27 18/10/2008 204926.81 631308.19 553218.429 -50557.675 26.6

LNC10 16 14:28:48 18/10/2008 204930.77 631313.05 553218.538 -50557.362 26.5

LNC10 17 14:29:40 18/10/2008 204941.48 631309.26 553218.468 -50556.319 26.6

LNC10 18 14:29:59 18/10/2008 204945.21 631308.10 553218.437 -50555.951 26.6

LNC10 19 14:30:45 18/10/2008 204954.58 631305.34 553218.328 -50555.041 26.6

LNC10 20 14:31:44 18/10/2008 204966.38 631299.72 553218.068 -50553.886 26.2

LNC10 21 14:33:14 18/10/2008 204981.26 631289.52 553217.629 -50552.352 24.7

LNC10 22 14:35:34 18/10/2008 205001.24 631278.52 553217.050 -50550.450 23.6

LNC10 23 14:35:58 18/10/2008 205005.65 631276.80 553216.954 -50550.041 23.3

LNC10 24 14:38:05 18/10/2008 205024.52 631262.50 553216.293 -50548.106 21.1

LNC10 25 14:40:05 18/10/2008 205047.30 631248.95 553215.597 -50545.919 20.4

LNC10 26 14:40:19 18/10/2008 205049.35 631247.48 553215.524 -50545.718 20.1

LNC11 2 15:01:23 18/10/2008 204429.74 631720.90 553239.462 -50606.626 19.2

LNC11 3 15:02:14 18/10/2008 204437.34 631715.18 553239.221 -50605.826 19.8

LNC11 4 15:02:45 18/10/2008 204441.99 631712.44 553239.065 -50605.396 20.2

LNC11 5 15:03:57 18/10/2008 204453.63 631706.47 553238.753 -50604.284 20.8

LNC11 6 15:04:53 18/10/2008 204464.30 631700.45 553238.481 -50603.225 21.2

LNC11 7 15:05:55 18/10/2008 204474.83 631693.81 553238.162 -50602.180 21.7

LNC11 8 15:07:47 18/10/2008 204495.63 631684.84 553237.668 -50600.232 22.0

LNC11 9 15:08:51 18/10/2008 204509.04 631680.70 553237.470 -50598.940 21.7

LNC11 10 15:09:39 18/10/2008 204518.53 631678.64 553237.360 -50598.041 21.3

LNC11 11 15:11:02 18/10/2008 204534.63 631675.27 553237.206 -50596.508 20.7

LNC11 12 15:11:17 18/10/2008 204537.63 631674.49 553237.177 -50596.214 20.7

LNC11 13 15:13:22 18/10/2008 204562.93 631668.01 553236.924 -50593.763 20.4

LNC11 14 15:15:45 18/10/2008 204589.58 631653.38 553236.268 -50591.104 21.3

LNC11 15 15:16:56 18/10/2008 204601.11 631645.10 553235.844 -50589.984 21.9

LNC11 16 15:18:17 18/10/2008 204611.51 631633.47 553235.287 -50588.874 23.0

LNC11 17 15:19:20 18/10/2008 204620.07 631624.72 553234.842 -50588.022 23.9

LNC11 18 15:20:24 18/10/2008 204630.39 631614.83 553234.337 -50586.997 24.8

LNC11 19 15:22:11 18/10/2008 204645.95 631592.88 553233.249 -50585.273 26.5

LNC11 20 15:22:26 18/10/2008 204647.24 631589.74 553233.087 -50585.115 26.8

LNC11 21 15:25:22 18/10/2008 204662.88 631556.92 553231.337 -50583.553 30.0

LNC11 22 15:26:16 18/10/2008 204668.71 631546.75 553230.805 -50582.964 30.7

LNC11 23 15:30:34 18/10/2008 204706.49 631505.20 553228.647 -50579.251 33.3

LNC12 2 08:45:17 19/10/2008 204519.86 630362.42 553166.144 -50591.996 31.3

LNC12 3 08:45:37 19/10/2008 204523.37 630373.62 553166.747 -50591.702 31.5

LNC12 4 08:45:49 19/10/2008 204525.39 630380.78 553167.133 -50591.533 31.4

LNC12 5 08:46:51 19/10/2008 204534.41 630416.58 553169.083 -50590.814 32.0

LNC12 6 08:47:24 19/10/2008 204539.05 630434.62 553170.062 -50590.428 32.3

Page 105: Mapping of the marine habitats and species of Lamlash Bay, Arran

93

LNC12 7 08:48:45 19/10/2008 204553.35 630480.86 553172.588 -50589.295 33.1

LNC12 8 08:49:23 19/10/2008 204559.25 630505.32 553173.915 -50588.793 33.3

LNC12 9 08:50:15 19/10/2008 204567.53 630538.39 553175.720 -50588.205 32.7

LNC12 10 08:51:22 19/10/2008 204579.34 630583.34 553178.168 -50587.264 31.4

LNC12 11 08:52:46 19/10/2008 204599.98 630635.09 553181.010 -50585.571 29.5

LNC12 12 08:54:00 19/10/2008 204615.45 630678.44 553183.380 -50584.254 27.7

LNC12 13 08:54:43 19/10/2008 204623.64 630704.73 553184.808 -50583.568 26.0

LNC12 14 08:55:06 19/10/2008 204628.74 630720.25 553185.662 -50583.173 24.8

LNC12 15 08:55:16 19/10/2008 204630.68 630727.14 553186.032 -50583.005 24.4

LNC12 16 08:55:39 19/10/2008 204635.41 630742.80 553186.891 -50582.623 23.4

LNC12 17 08:56:07 19/10/2008 204641.72 630762.45 553187.963 -50582.116 22.4

LNC12 18 08:57:14 19/10/2008 204658.75 630805.76 553190.337 -50580.700 21.4

LNC12 19 08:57:34 19/10/2008 204662.94 630819.39 553191.075 -50580.346 21.2

LNC12 20 08:58:17 19/10/2008 204672.16 630849.17 553192.704 -50579.572 21.3

LNC12 21 08:58:59 19/10/2008 204682.28 630878.90 553194.327 -50578.760 21.4

LNC12 22 08:59:36 19/10/2008 204692.52 630903.74 553195.699 -50577.948 21.7

LNC12 23 09:00:50 19/10/2008 204711.20 630951.12 553198.294 -50576.342 22.0

LNC12 24 09:01:30 19/10/2008 204720.11 630979.38 553199.823 -50575.566 21.7

LNC12 25 09:02:09 19/10/2008 204729.43 631006.85 553201.326 -50574.808 21.7

LNC12 26 09:02:41 19/10/2008 204738.04 631029.62 553202.585 -50574.128 21.6

LNC12 27 09:03:04 19/10/2008 204744.84 631044.83 553203.422 -50573.570 21.3

LNC12 28 09:03:21 19/10/2008 204749.71 631056.02 553204.036 -50573.153 21.2

LNC12 29 09:04:10 19/10/2008 204763.97 631089.32 553205.862 -50571.933 20.9

LNC12 30 09:04:22 19/10/2008 204767.47 631097.26 553206.298 -50571.643 20.8

LNC12 31 09:04:40 19/10/2008 204772.37 631109.52 553206.966 -50571.212 20.7

LNC12 32 09:05:10 19/10/2008 204780.16 631131.39 553208.157 -50570.542 20.7

LNC12 33 09:05:29 19/10/2008 204785.12 631145.20 553208.914 -50570.130 20.8

LNC12 34 09:05:50 19/10/2008 204791.40 631158.39 553209.640 -50569.602 20.5

LNC12 35 09:06:15 19/10/2008 204798.83 631177.20 553210.674 -50568.987 20.6

LNC12 36 09:06:38 19/10/2008 204807.52 631194.01 553211.609 -50568.277 20.5

LNC12 37 09:06:53 19/10/2008 204814.10 631203.74 553212.166 -50567.740 20.4

LNC12 38 09:07:09 19/10/2008 204821.56 631212.01 553212.651 -50567.120 20.4

LNC12 39 09:07:33 19/10/2008 204829.23 631233.01 553213.772 -50566.411 20.3

LNC12 40 09:08:28 19/10/2008 204848.22 631270.64 553215.831 -50564.734 20.8

LNC12 41 09:08:48 19/10/2008 204855.06 631284.28 553216.585 -50564.145 21.4

LNC12 42 09:09:17 19/10/2008 204865.18 631305.47 553217.745 -50563.254 22.3

LNC12 43 09:09:36 19/10/2008 204872.26 631319.68 553218.528 -50562.658 22.9

LNC12 44 09:09:57 19/10/2008 204880.80 631334.54 553219.357 -50561.941 23.6

LNC12 45 09:10:27 19/10/2008 204894.34 631352.89 553220.386 -50560.750 24.8

LNC12 46 09:10:40 19/10/2008 204900.87 631361.00 553220.853 -50560.198 25.5

LNC12 47 09:10:57 19/10/2008 204909.42 631371.13 553221.428 -50559.451 26.1

LNC12 48 09:11:39 19/10/2008 204927.22 631400.61 553223.047 -50557.866 27.4

LNC12 49 09:12:19 19/10/2008 204944.73 631421.70 553224.241 -50556.332 28.2

LNC12 50 09:12:33 19/10/2008 204949.62 631431.01 553224.727 -50555.840 28.2

LNC12 51 09:12:57 19/10/2008 204960.44 631447.05 553225.623 -50554.895 28.4

LNC12 52 09:13:10 19/10/2008 204966.79 631455.77 553226.111 -50554.341 28.3

Page 106: Mapping of the marine habitats and species of Lamlash Bay, Arran

94

LNC12 53 09:13:37 19/10/2008 204973.71 631475.11 553227.147 -50553.704 28.4

LNC12 54 09:13:59 19/10/2008 204982.81 631491.71 553228.057 -50552.897 28.4

LNC12 55 09:14:29 19/10/2008 204996.15 631514.67 553229.331 -50551.751 28.4

LNC12 56 09:14:49 19/10/2008 205006.00 631529.04 553230.145 -50550.921 28.6

LNC12 57 09:15:08 19/10/2008 205015.23 631541.98 553230.866 -50550.108 28.8

LNC12 58 09:15:41 19/10/2008 205031.65 631562.50 553232.018 -50548.650 29.1

Page 107: Mapping of the marine habitats and species of Lamlash Bay, Arran

95

Appendix 03: Lamlash Bay Still Photographs log - 2008SeaStar Survey - Project J/08/151

Lamlash SouthPositions are OSGB36 East and North (m) and WGS84 Latitude and Longitude (ddmmss.sss)

Client SNH Job NumberLocation Lamlash South VesselSurvey Lamlash Bay Habitat Mapping Survey Date

Site Photograph Time Date East North Latitude (N) Longitude (W) Depth CommentsName Number (GMT) (m) (m) ddmmss.sss ddmmss.sss (m)LSC01 2 17:21:34 16/10/2008 204895.07 629691.37 553131.341 -50553.814 45.2 Depth correct to below water surfaceLSC01 3 17:22:02 16/10/2008 204904.98 629686.48 553131.095 -50552.862 45.3

LSC01 4 17:22:54 16/10/2008 204923.87 629677.95 553130.666 -50551.045 45.4

LSC01 5 17:23:41 16/10/2008 204942.35 629670.43 553130.299 -50549.264 45.4

LSC01 6 17:24:34 16/10/2008 204963.92 629664.07 553129.854 -50547.231 45.1

LSC01 7 17:25:04 16/10/2008 204975.10 629656.67 553129.636 -50546.099 45.2

LSC01 8 17:26:08 16/10/2008 204999.75 629646.45 553129.147 -50543.716 45.1

LSC01 9 17:26:42 16/10/2008 205012.32 629640.80 553128.884 -50542.491 45.2

LSC01 10 17:26:46 16/10/2008 205013.77 629640.10 553128.851 -50542.349 45.2

LSC01 11 17:27:11 16/10/2008 205022.17 629635.37 553128.634 -50541.515 45.3

LSC01 12 17:28:11 16/10/2008 205047.35 629625.24 553128.109 -50539.116 45.2

LSC01 13 17:29:24 16/10/2008 205074.82 629610.81 553127.437 -50536.415 44.3

LSC01 14 17:30:26 16/10/2008 205097.44 629598.20 553126.814 -50534.214 44.2

LSC01 15 17:32:08 16/10/2008 205134.67 629579.65 553125.889 -50530.618 44.8

LSC01 16 17:32:36 16/10/2008 205144.94 629574.99 553125.665 -50529.624 44.9

LSC01 17 17:32:49 16/10/2008 205149.85 629572.93 553125.560 -50529.154 45.0

LSC01 18 17:33:01 16/10/2008 205154.34 629570.89 553125.461 -50528.720 44.9

LSC01 19 17:33:35 16/10/2008 205167.31 629565.57 553125.199 -50527.472 44.9

LSC01 20 17:33:56 16/10/2008 205175.23 629562.35 553125.039 -50526.709 44.5

LSC01 21 17:34:10 16/10/2008 205179.32 629559.57 553124.933 -50526.279 44.4

LSC01 22 17:34:55 16/10/2008 205197.33 629553.66 553124.622 -50524.579 43.9

LSC01 23 17:35:23 16/10/2008 205207.69 629549.53 553124.426 -50523.577 43.5

LSC01 24 17:36:14 16/10/2008 205226.47 629541.93 553124.064 -50521.762 42.9

LSC01 25 17:36:40 16/10/2008 205236.16 629538.08 553123.881 -50520.825 42.5

LSC01 26 17:37:35 16/10/2008 205256.82 629530.55 553123.520 -50518.839 41.9

LSC01 27 17:39:49 16/10/2008 205309.15 629513.71 553122.704 -50513.822 40.8

LSC01 28 17:40:26 16/10/2008 205323.48 629508.89 553122.499 -50512.431 40.4

LSC01 29 17:40:38 16/10/2008 205328.16 629507.45 553122.431 -50511.981 40.4

LSC01 30 17:40:50 16/10/2008 205332.62 629505.97 553122.365 -50511.551 40.3

LSC01 31 17:41:32 16/10/2008 205348.43 629500.45 553122.106 -50510.026 39.9

LSC01 32 17:41:59 16/10/2008 205358.57 629496.87 553121.945 -50509.045 39.6

LSC01 33 17:42:25 16/10/2008 205368.69 629492.58 553121.753 -50508.057 39.3

LSC01 34 17:43:22 16/10/2008 205390.74 629482.61 553121.287 -50505.903 38.6

LSC01 35 17:43:36 16/10/2008 205395.88 629479.36 553121.139 -50505.386 38.4

LSC01 36 17:43:52 16/10/2008 205401.65 629475.44 553120.953 -50504.810 38.2

LSC01 37 17:44:05 16/10/2008 205407.83 629474.73 553120.887 -50504.265 37.9

LSC01 38 17:44:50 16/10/2008 205423.55 629464.69 553120.422 -50502.691 36.9

LSC01 39 17:45:13 16/10/2008 205431.65 629459.40 553120.161 -50501.896 36.2

LSC01 40 17:45:34 16/10/2008 205439.16 629454.28 553119.907 -50501.157 35.5

J/08/151SV Alma Na MaraOctober 2008

Page 108: Mapping of the marine habitats and species of Lamlash Bay, Arran

96

LSC01 41 17:45:56 16/10/2008 205447.07 629449.35 553119.655 -50500.393 34.6

LSC01 42 17:46:24 16/10/2008 205455.69 629442.38 553119.310 -50499.532 33.7

LSC01 43 17:47:04 16/10/2008 205468.74 629435.67 553118.939 -50498.312 33.0

LSC01 44 17:47:20 16/10/2008 205472.07 629427.65 553118.586 -50497.870 32.8

LSC01 45 17:47:50 16/10/2008 205482.71 629422.57 553118.305 -50496.890 32.4

LSC01 46 17:48:06 16/10/2008 205487.52 629419.04 553118.133 -50496.411 32.2

LSC01 47 17:48:34 16/10/2008 205496.30 629411.92 553117.786 -50495.525 32.1

LSC01 48 17:49:20 16/10/2008 205512.37 629401.76 553117.251 -50493.993 31.8

LSC01 49 17:49:36 16/10/2008 205517.97 629398.02 553117.066 -50493.443 31.6

LSC01 50 17:49:36 16/10/2008 205517.97 629398.02 553117.066 -50493.443 31.6

LSC01 51 17:50:19 16/10/2008 205530.93 629387.88 553116.562 -50492.156 31.0

LSC01 52 17:51:30 16/10/2008 205555.32 629371.26 553115.714 -50489.787 29.6

LSC01 53 17:52:10 16/10/2008 205568.99 629360.38 553115.184 -50488.414 29.1

LSC01 54 17:52:36 16/10/2008 205577.73 629353.59 553114.837 -50487.557 29.1

LSC01 55 17:53:07 16/10/2008 205587.76 629345.54 553114.429 -50486.567 29.2

LSC01 56 17:53:21 16/10/2008 205592.42 629342.01 553114.248 -50486.117 30.1

LSC01 57 17:53:31 16/10/2008 205595.69 629339.39 553114.116 -50485.790 30.6

LSC01 58 17:54:38 16/10/2008 205616.88 629322.87 553113.261 -50483.736 31.8

LSC01 59 17:55:15 16/10/2008 205627.78 629312.69 553112.765 -50482.620 31.9

LSC01 60 17:55:24 16/10/2008 205630.47 629310.16 553112.639 -50482.348 32.0

LSC01 61 17:55:48 16/10/2008 205637.50 629302.80 553112.267 -50481.638 32.1

LSC02 2 09:37:39 19/10/2008 204065.97 630022.94 553146.761 -50633.492 39.7

LSC02 3 09:38:36 19/10/2008 204068.94 630046.86 553148.112 -50633.627 38.9

LSC02 4 09:38:56 19/10/2008 204065.72 630059.73 553148.727 -50633.673 38.8

LSC02 5 09:39:36 19/10/2008 204065.42 630077.54 553149.694 -50633.670 38.6

LSC02 6 09:40:14 19/10/2008 204067.96 630093.76 553150.568 -50633.631 38.1

LSC02 7 09:40:49 19/10/2008 204069.97 630109.91 553151.446 -50633.534 37.9

LSC02 8 09:41:20 19/10/2008 204072.43 630124.61 553152.253 -50633.382 37.8

LSC02 9 09:41:41 19/10/2008 204074.84 630132.40 553152.688 -50633.261 37.6

LSC02 10 09:42:25 19/10/2008 204082.70 630146.87 553153.594 -50632.842 37.3

LSC02 11 09:43:10 19/10/2008 204086.82 630170.44 553154.780 -50632.365 37.4

LSC02 12 09:44:57 19/10/2008 204108.83 630221.21 553157.556 -50630.465 38.1

LSC02 13 09:45:49 19/10/2008 204113.64 630245.09 553158.835 -50630.009 38.4

LSC02 14 09:47:40 19/10/2008 204122.05 630303.78 553162.009 -50629.407 37.6

LSC02 15 09:48:00 19/10/2008 204126.78 630312.97 553162.631 -50629.329 37.1

LSC02 16 09:50:29 19/10/2008 204143.51 630397.04 553167.092 -50627.866 33.5

LSC02 17 09:51:10 19/10/2008 204150.42 630420.06 553168.349 -50627.301 32.4

LSC02 18 09:51:37 19/10/2008 204154.94 630433.67 553169.094 -50626.938 31.7

LSC02 19 09:52:11 19/10/2008 204161.01 630451.31 553170.054 -50626.439 30.9

LSC02 20 09:52:46 19/10/2008 204166.72 630469.59 553171.057 -50625.955 30.5

LSC02 21 09:54:14 19/10/2008 204183.30 630510.84 553173.327 -50624.584 30.1

LSC02 22 09:55:03 19/10/2008 204191.16 630535.37 553174.649 -50623.941 30.1

LSC02 23 09:55:53 19/10/2008 204202.42 630563.72 553176.221 -50623.034 30.6

LSC02 24 09:56:51 19/10/2008 204214.96 630592.26 553177.783 -50621.938 31.2

LSC02 25 09:59:59 19/10/2008 204257.19 630697.20 553183.559 -50618.477 30.3

LSC02 26 10:00:21 19/10/2008 204262.99 630709.63 553184.227 -50617.917 30.4

LSC02 27 10:00:36 19/10/2008 204266.16 630718.57 553184.712 -50617.588 30.5

LSC02 28 10:00:55 19/10/2008 204270.17 630729.91 553185.325 -50617.252 30.3

LSC02 29 10:02:29 19/10/2008 204288.91 630783.80 553188.260 -50615.729 30.1

Page 109: Mapping of the marine habitats and species of Lamlash Bay, Arran

97

LSC02 30 10:03:11 19/10/2008 204293.84 630807.03 553189.519 -50615.310 30.1

LSC02 31 10:03:33 19/10/2008 204298.44 630819.92 553190.233 -50614.956 30.0

LSC02 32 10:04:38 19/10/2008 204316.50 630856.16 553192.253 -50613.459 30.2

LSC02 33 10:05:22 19/10/2008 204329.08 630880.99 553193.605 -50612.352 30.1

LSC02 34 10:07:27 19/10/2008 204371.90 630954.10 553197.680 -50608.670 27.0

LSC02 35 10:08:20 19/10/2008 204385.99 630981.86 553199.172 -50607.376 24.9

LSC02 36 10:08:57 19/10/2008 204393.97 631002.64 553200.303 -50606.677 23.5

LSC02 37 10:09:12 19/10/2008 204398.37 631012.11 553200.830 -50606.319 23.0

LSC02 38 10:09:31 19/10/2008 204403.21 631023.90 553201.476 -50605.891 22.3

LSC02 39 10:09:36 19/10/2008 204404.89 631026.56 553201.627 -50605.775 22.2

LSC02 40 10:10:15 19/10/2008 204415.99 631051.17 553202.971 -50604.838 21.7

LSC02 41 10:10:34 19/10/2008 204421.88 631063.81 553203.674 -50604.328 21.9

LSC02 42 10:10:59 19/10/2008 204431.00 631077.88 553204.474 -50603.578 22.1

LSC02 43 10:11:30 19/10/2008 204442.74 631094.54 553205.399 -50602.546 22.9

LSC02 44 10:11:57 19/10/2008 204451.46 631109.16 553206.198 -50601.757 23.5

LSC02 45 10:13:36 19/10/2008 204483.10 631161.47 553209.081 -50598.944 26.5

LSC02 46 10:13:59 19/10/2008 204491.79 631175.49 553209.867 -50598.219 27.3

LSC02 47 10:14:11 19/10/2008 204496.49 631184.28 553210.349 -50597.800 27.8

LSC02 48 10:15:08 19/10/2008 204519.48 631219.47 553212.317 -50595.801 29.7

LSC03 2 10:08:26 20/10/2008 204875.09 628966.79 553091.974 -50552.649 37.2

LSC03 3 10:09:08 20/10/2008 204883.96 628976.75 553092.483 -50551.770 37.4

LSC03 4 10:10:30 20/10/2008 204902.40 628990.40 553093.330 -50550.169 37.5

LSC03 5 10:11:03 20/10/2008 204911.04 628994.38 553093.578 -50549.376 37.5

LSC03 6 10:11:34 20/10/2008 204918.15 628998.70 553093.806 -50548.698 37.5

LSC03 7 10:12:19 20/10/2008 204933.68 629003.31 553094.163 -50547.310 37.8

LSC03 8 10:12:42 20/10/2008 204941.30 629004.99 553094.306 -50546.604 37.5

LSC03 9 10:13:02 20/10/2008 204948.37 629005.54 553094.373 -50545.941 37.4

LSC03 10 10:14:14 20/10/2008 204973.24 629017.19 553094.892 -50543.500 37.5

LSC03 11 10:17:36 20/10/2008 205044.85 629039.80 553096.469 -50536.941 38.0

LSC03 12 10:18:15 20/10/2008 205059.49 629042.91 553096.644 -50535.546 38.4

LSC03 13 10:19:09 20/10/2008 205081.61 629046.92 553096.884 -50533.451 38.7

LSC03 14 10:19:26 20/10/2008 205088.73 629049.24 553097.006 -50532.793 38.8

LSC03 15 10:19:48 20/10/2008 205097.52 629052.41 553097.187 -50531.970 38.8

LSC03 16 10:20:34 20/10/2008 205115.78 629059.93 553097.626 -50530.260 38.9

LSC03 17 10:20:46 20/10/2008 205120.90 629061.89 553097.747 -50529.776 39.0

LSC03 18 10:21:25 20/10/2008 205137.19 629066.14 553098.052 -50528.267 38.9

LSC03 19 10:22:21 20/10/2008 205154.23 629072.80 553098.441 -50526.680 38.8

LSC03 20 10:22:41 20/10/2008 205159.71 629073.76 553098.522 -50526.162 38.8

LSC03 21 10:22:59 20/10/2008 205164.75 629075.48 553098.603 -50525.691 38.7

LSC03 22 10:23:43 20/10/2008 205180.63 629082.24 553098.977 -50524.199 38.7

LSC03 23 10:24:55 20/10/2008 205207.30 629097.77 553099.838 -50521.702 38.7

LSC03 24 10:26:08 20/10/2008 205223.49 629110.88 553100.540 -50520.160 39.2

LSC03 25 10:29:48 20/10/2008 205277.98 629152.59 553102.934 -50515.209 40.2

LSC03 26 10:32:56 20/10/2008 205331.81 629179.31 553104.542 -50510.254 39.2

LSC03 27 10:34:12 20/10/2008 205351.53 629192.14 553105.263 -50508.421 39.1

LSC03 28 10:34:31 20/10/2008 205355.67 629194.43 553105.402 -50508.042 39.0

LSC03 29 10:36:00 20/10/2008 205369.42 629212.67 553106.310 -50506.648 39.0

LSC03 30 10:37:06 20/10/2008 205380.09 629226.15 553107.102 -50505.763 39.2

LSC03 31 10:37:21 20/10/2008 205383.91 629228.72 553107.272 -50505.449 39.2

LSC03 32 10:40:06 20/10/2008 205402.62 629272.36 553109.570 -50503.658 39.6

Page 110: Mapping of the marine habitats and species of Lamlash Bay, Arran

98

LSC03 33 10:41:27 20/10/2008 205406.62 629291.66 553110.596 -50503.171 39.8

LSC03 34 10:41:49 20/10/2008 205406.33 629295.90 553110.828 -50503.198 39.9

LSC03 35 10:42:42 20/10/2008 205409.08 629306.41 553111.406 -50503.079 40.1

LSC03 36 10:42:57 20/10/2008 205410.79 629309.04 553111.564 -50502.999 40.1

LSC03 37 10:43:13 20/10/2008 205411.50 629312.79 553111.773 -50502.907 40.3

LSC04 2 12:26:53 20/10/2008 204730.18 629404.46 553115.291 -50568.312 44.2

LSC04 3 12:27:32 20/10/2008 204743.09 629408.00 553115.498 -50567.100 44.2

LSC04 4 12:27:56 20/10/2008 204750.16 629410.81 553115.639 -50566.423 44.2

LSC04 5 12:29:42 20/10/2008 204774.95 629420.15 553116.117 -50564.021 44.4

LSC04 6 12:30:07 20/10/2008 204781.34 629426.44 553116.436 -50563.378 44.4

LSC04 7 12:30:24 20/10/2008 204786.27 629430.77 553116.696 -50562.961 44.4

LSC04 8 12:31:11 20/10/2008 204800.25 629439.94 553117.275 -50561.746 44.4

LSC04 9 12:31:51 20/10/2008 204812.48 629444.43 553117.599 -50560.659 44.4

LSC04 10 12:32:50 20/10/2008 204828.58 629446.42 553117.826 -50559.174 44.4

LSC04 11 12:33:18 20/10/2008 204834.65 629448.05 553117.898 -50558.586 44.6

LSC04 12 12:35:04 20/10/2008 204856.65 629460.28 553118.498 -50556.475 44.5

LSC04 13 12:35:18 20/10/2008 204859.41 629462.50 553118.610 -50556.215 44.5

LSC04 14 12:35:28 20/10/2008 204860.83 629464.54 553118.706 -50556.047 44.6

LSC04 15 12:37:17 20/10/2008 204882.12 629479.79 553119.587 -50554.125 44.6

LSC04 16 12:37:58 20/10/2008 204891.40 629483.57 553119.888 -50553.314 44.9

LSC04 17 12:38:50 20/10/2008 204905.75 629491.72 553120.325 -50551.967 45.0

LSC04 18 12:39:07 20/10/2008 204912.04 629492.44 553120.444 -50551.422 45.1

LSC04 19 12:39:42 20/10/2008 204923.32 629494.18 553120.603 -50550.389 45.1

LSC04 20 12:40:23 20/10/2008 204934.57 629494.51 553120.663 -50549.327 45.2

LSC04 21 12:42:32 20/10/2008 204966.67 629516.83 553121.813 -50546.281 45.8

LSC04 22 12:42:54 20/10/2008 204972.98 629521.46 553122.050 -50545.675 45.9

LSC04 23 12:45:51 20/10/2008 205014.05 629546.37 553123.515 -50541.918 46.6

LSC04 24 12:49:24 20/10/2008 205059.28 629568.46 553124.916 -50537.786 47.0

LSC04 25 12:52:32 20/10/2008 205105.39 629568.77 553125.033 -50533.421 47.3

LSC04 26 12:54:05 20/10/2008 205134.41 629577.78 553125.568 -50530.669 48.1

LSC04 27 12:55:37 20/10/2008 205156.77 629588.25 553126.110 -50528.557 48.4

LSC04 28 12:55:51 20/10/2008 205160.73 629589.68 553126.221 -50528.217 48.4

LSC04 29 12:56:00 20/10/2008 205163.11 629590.66 553126.287 -50528.014 48.4

LSC04 30 12:57:00 20/10/2008 205179.15 629601.52 553126.884 -50526.500 48.4

LSC04 31 12:57:48 20/10/2008 205197.70 629610.07 553127.391 -50524.803 48.0

LSC04 32 12:58:24 20/10/2008 205214.09 629611.27 553127.619 -50523.291 47.9

LSC04 33 12:59:31 20/10/2008 205242.98 629612.50 553127.760 -50520.571 47.4

LSC04 34 13:00:26 20/10/2008 205260.95 629621.05 553128.170 -50518.861 47.4

LSC04 35 13:02:27 20/10/2008 205289.16 629643.02 553129.352 -50516.190 46.5

LSC04 36 13:02:49 20/10/2008 205295.21 629647.40 553129.607 -50515.638 46.1

LSC04 37 13:03:36 20/10/2008 205308.61 629653.61 553130.025 -50514.457 44.8

LSC04 38 13:04:02 20/10/2008 205315.78 629658.22 553130.308 -50513.804 43.8

LSC04 39 13:05:01 20/10/2008 205334.90 629666.38 553130.786 -50512.003 40.7

LSC04 40 13:05:15 20/10/2008 205338.99 629668.40 553130.893 -50511.629 40.0

LSC04 41 13:05:29 20/10/2008 205342.89 629670.18 553131.002 -50511.277 39.3

LSC04 42 13:05:41 20/10/2008 205345.48 629667.38 553130.982 -50511.058 38.9

LSC04 43 13:06:12 20/10/2008 205351.67 629674.34 553131.239 -50510.432 38.0

LSC04 44 13:06:33 20/10/2008 205355.68 629679.66 553131.473 -50510.014 37.3

LSC04 45 13:06:45 20/10/2008 205357.62 629683.80 553131.673 -50509.813 36.8

LSC04 46 13:07:23 20/10/2008 205368.05 629691.53 553132.158 -50508.921 35.1

LSC04 47 13:07:58 20/10/2008 205378.76 629697.61 553132.556 -50507.986 33.6

Page 111: Mapping of the marine habitats and species of Lamlash Bay, Arran

99

LSC04 48 13:08:20 20/10/2008 205384.99 629702.09 553132.800 -50507.376 32.7

LSC04 49 13:08:49 20/10/2008 205395.95 629705.72 553133.059 -50506.414 30.3

LSC04 50 13:08:58 20/10/2008 205399.12 629705.97 553133.109 -50506.120 29.5

LSC04 51 13:09:06 20/10/2008 205402.17 629706.29 553133.153 -50505.846 29.1

LSC05 2 13:24:51 20/10/2008 204352.91 629762.09 553133.548 -50605.575 44.1

LSC05 3 13:25:04 20/10/2008 204357.93 629764.17 553133.693 -50605.130 44.0

LSC05 4 13:26:42 20/10/2008 204390.70 629768.45 553134.116 -50602.088 43.9

LSC05 5 13:27:18 20/10/2008 204400.17 629769.91 553134.202 -50601.187 44.2

LSC05 6 13:28:16 20/10/2008 204415.10 629773.13 553134.388 -50599.778 44.2

LSC05 7 13:28:43 20/10/2008 204421.70 629774.65 553134.478 -50599.153 44.2

LSC05 8 13:29:54 20/10/2008 204440.55 629778.32 553134.727 -50597.386 44.2

LSC05 9 13:30:25 20/10/2008 204449.06 629782.55 553134.898 -50596.556 44.2

LSC05 10 13:31:49 20/10/2008 204476.22 629788.23 553135.308 -50594.022 44.2

LSC05 11 13:33:57 20/10/2008 204513.82 629796.81 553135.910 -50590.509 44.1

LSC05 12 13:35:06 20/10/2008 204538.70 629800.39 553136.164 -50588.171 44.1

LSC05 13 13:36:19 20/10/2008 204570.97 629804.83 553136.491 -50585.133 44.5

LSC05 14 13:38:40 20/10/2008 204625.00 629802.49 553136.540 -50579.998 45.3

LSC05 15 13:39:28 20/10/2008 204648.99 629801.28 553136.556 -50577.715 45.7

LSC05 16 13:40:57 20/10/2008 204686.09 629790.70 553136.132 -50574.124 46.5

LSC05 17 13:42:14 20/10/2008 204715.84 629786.06 553135.901 -50571.302 46.8

LSC05 18 13:42:46 20/10/2008 204728.12 629785.25 553135.870 -50570.130 47.1

LSC05 19 13:43:11 20/10/2008 204738.65 629783.71 553135.841 -50569.126 47.1

LSC06 2 13:59:32 20/10/2008 204648.97 630101.76 553152.769 -50578.969 44.5

LSC06 3 13:59:46 20/10/2008 204653.96 630100.30 553152.708 -50578.489 44.6

LSC06 4 14:00:16 20/10/2008 204663.47 630098.57 553152.618 -50577.587 44.9

LSC06 5 14:00:33 20/10/2008 204668.41 630098.64 553152.597 -50577.113 45.1

LSC06 6 14:01:22 20/10/2008 204682.29 630099.95 553152.653 -50575.816 45.2

LSC06 7 14:01:54 20/10/2008 204691.32 630100.75 553152.719 -50574.969 45.3

LSC06 8 14:02:31 20/10/2008 204701.08 630103.02 553152.817 -50574.013 45.7

LSC06 9 14:03:12 20/10/2008 204712.89 630104.37 553152.994 -50572.920 45.7

LSC06 10 14:04:26 20/10/2008 204737.58 630110.05 553153.319 -50570.605 45.9

LSC06 11 14:04:58 20/10/2008 204748.27 630110.47 553153.411 -50569.603 46.0

LSC06 12 14:05:08 20/10/2008 204750.83 630112.69 553153.430 -50569.326 46.1

LSC06 13 14:05:42 20/10/2008 204760.70 630109.19 553153.408 -50568.409 46.2

LSC06 14 14:06:43 20/10/2008 204776.32 630112.78 553153.469 -50566.905 46.3

LSC06 15 14:07:38 20/10/2008 204793.25 630123.06 553154.036 -50565.306 46.4

LSC06 16 14:08:07 20/10/2008 204802.79 630125.27 553154.267 -50564.480 46.5

LSC06 17 14:09:29 20/10/2008 204832.73 630132.47 553154.702 -50561.649 46.8

LSC06 18 14:09:50 20/10/2008 204842.78 630133.59 553154.826 -50560.715 46.8

LSC06 19 14:10:23 20/10/2008 204857.45 630131.33 553154.808 -50559.326 46.9

LSC06 20 14:11:02 20/10/2008 204872.70 630129.28 553154.741 -50557.867 47.0

LSC06 21 14:11:16 20/10/2008 204878.29 630128.23 553154.713 -50557.337 46.9

LSC06 22 14:12:02 20/10/2008 204894.88 630126.41 553154.681 -50555.723 47.0

LSC06 23 14:13:30 20/10/2008 204930.62 630118.41 553154.308 -50552.330 46.5

LSC06 24 14:14:52 20/10/2008 204963.49 630115.91 553154.227 -50549.209 46.2

LSC06 25 14:15:58 20/10/2008 204986.13 630120.37 553154.483 -50547.065 46.0

LSC06 26 14:16:41 20/10/2008 205002.68 630121.78 553154.592 -50545.493 46.0

LSC06 27 14:17:19 20/10/2008 205018.61 630122.05 553154.640 -50543.990 45.9

LSC06 28 14:18:00 20/10/2008 205033.35 630124.49 553154.789 -50542.599 45.6

LSC06 29 14:19:18 20/10/2008 205058.18 630133.99 553155.313 -50540.261 44.5

LSC06 30 14:21:14 20/10/2008 205094.96 630157.87 553156.669 -50536.861 40.6

Page 112: Mapping of the marine habitats and species of Lamlash Bay, Arran

100

LSC06 31 14:22:07 20/10/2008 205113.03 630166.25 553157.168 -50535.176 37.5

LSC07 2 14:36:56 20/10/2008 203811.80 630387.76 553166.088 -50659.649 39.0

LSC07 3 14:37:44 20/10/2008 203836.19 630387.96 553166.163 -50657.332 38.9

LSC07 4 14:37:57 20/10/2008 203842.95 630387.61 553166.159 -50656.693 38.8

LSC07 5 14:38:30 20/10/2008 203859.99 630387.12 553166.151 -50655.072 38.7

LSC07 6 14:38:46 20/10/2008 203867.62 630388.42 553166.214 -50654.354 38.6

LSC07 7 14:39:30 20/10/2008 203888.40 630393.00 553166.480 -50652.388 38.5

LSC07 8 14:40:02 20/10/2008 203904.13 630396.81 553166.719 -50650.910 38.2

LSC07 9 14:40:10 20/10/2008 203908.11 630397.53 553166.773 -50650.542 38.2

LSC07 10 14:40:49 20/10/2008 203928.13 630403.04 553167.101 -50648.655 38.3

LSC07 11 14:41:17 20/10/2008 203943.39 630405.77 553167.302 -50647.223 38.4

LSC07 12 14:41:46 20/10/2008 203959.27 630408.41 553167.500 -50645.734 38.3

LSC07 13 14:42:25 20/10/2008 203980.01 630409.24 553167.633 -50643.778 38.3

LSC07 14 14:42:38 20/10/2008 203986.49 630409.15 553167.648 -50643.163 38.1

LSC07 15 14:43:01 20/10/2008 203997.27 630408.52 553167.646 -50642.138 38.0

LSC07 16 14:43:47 20/10/2008 204016.38 630410.39 553167.766 -50640.334 37.6

LSC07 17 14:44:28 20/10/2008 204036.89 630413.73 553167.971 -50638.389 37.6

LSC07 18 14:44:45 20/10/2008 204045.57 630416.39 553168.088 -50637.553 37.5

LSC07 19 14:47:08 20/10/2008 204120.33 630429.25 553169.020 -50630.532 34.9

LSC07 20 14:48:00 20/10/2008 204145.68 630427.68 553169.051 -50628.126 34.0

LSC07 21 14:48:28 20/10/2008 204160.07 630426.70 553169.051 -50626.754 33.5

LSC07 22 14:49:08 20/10/2008 204181.05 630434.55 553169.440 -50624.798 32.5

LSC07 23 14:49:27 20/10/2008 204192.75 630435.63 553169.540 -50623.690 32.1

LSC07 24 14:49:47 20/10/2008 204205.44 630435.66 553169.588 -50622.496 31.8

LSC07 25 14:50:54 20/10/2008 204239.20 630431.19 553169.488 -50619.265 30.7

LSC07 26 14:51:09 20/10/2008 204246.27 630431.08 553169.477 -50618.600 30.5

LSC07 27 14:51:40 20/10/2008 204260.71 630430.16 553169.458 -50617.224 30.3

LSC07 28 14:51:55 20/10/2008 204268.51 630430.28 553169.460 -50616.477 30.1

LSC07 29 14:52:26 20/10/2008 204287.72 630427.20 553169.370 -50614.642 29.2

LSC07 30 14:52:35 20/10/2008 204293.34 630424.81 553169.238 -50614.093 29.3

LSC07 31 14:53:00 20/10/2008 204306.18 630428.66 553169.467 -50612.911 29.3

LSC07 32 14:54:13 20/10/2008 204346.76 630432.04 553169.725 -50609.068 30.2

LSC07 33 14:55:34 20/10/2008 204390.48 630428.34 553169.693 -50604.893 32.0

LSC07 34 14:56:00 20/10/2008 204403.89 630428.50 553169.700 -50603.628 31.9

LSC07 35 14:56:29 20/10/2008 204418.95 630429.17 553169.752 -50602.203 32.8

LSC07 36 14:57:01 20/10/2008 204437.61 630434.54 553170.012 -50600.439 33.2

LSC07 37 14:57:56 20/10/2008 204464.56 630432.32 553170.036 -50597.885 34.0

LSC07 38 14:58:13 20/10/2008 204471.85 630433.24 553170.085 -50597.202 34.2

LSC07 39 14:58:28 20/10/2008 204478.20 630434.47 553170.152 -50596.604 34.4

LSC07 40 14:59:15 20/10/2008 204499.12 630438.69 553170.414 -50594.630 34.7

LSC07 41 15:00:23 20/10/2008 204538.66 630436.65 553170.522 -50590.868 35.2

LSC07 42 15:02:11 20/10/2008 204597.48 630430.26 553170.247 -50585.272 36.2

LSC07 43 15:05:10 20/10/2008 204691.90 630406.08 553169.244 -50576.183 40.2

LSC07 44 15:05:22 20/10/2008 204697.67 630404.52 553169.182 -50575.629 40.3

LSC07 45 15:05:38 20/10/2008 204705.27 630402.58 553169.097 -50574.904 40.5

LSC07 46 15:06:30 20/10/2008 204733.69 630401.04 553169.016 -50572.216 40.7

LSC07 47 15:07:23 20/10/2008 204762.65 630398.22 553168.984 -50569.446 40.9

Page 113: Mapping of the marine habitats and species of Lamlash Bay, Arran

101

Appendix 03: Lamlash Bay Still Photographs log - 2008SeaStar Survey - Project J/08/151

Lamlash OuterPositions are OSGB36 East and North (m) and WGS84 Latitude and Longitude (ddmmss.sss)

Client SNH Job NumberLocation Lamlash Outer VesselSurvey Lamlash Bay Habitat Mapping Survey Date

Site Photograph Time Date East North Latitude (N) Longitude (W) Depth CommentsName Number (GMT) (m) (m) ddmmss.sss ddmmss.sss (m)

LOC01 2 07:19:47 20/10/2008 204915.28 634504.35 553390.330 -50572.353 18.5 Depth correct to below water surfaceLOC01 3 07:19:58 20/10/2008 204918.78 634504.44 553390.353 -50572.016 19.2

LOC01 4 07:20:11 20/10/2008 204923.37 634504.30 553390.371 -50571.582 20.0

LOC01 5 07:20:34 20/10/2008 204930.14 634504.34 553390.397 -50570.947 21.2

LOC01 6 07:20:58 20/10/2008 204936.72 634504.36 553390.416 -50570.309 22.4

LOC01 7 07:21:20 20/10/2008 204942.68 634502.99 553390.394 -50569.726 23.3

LOC01 8 07:21:36 20/10/2008 204947.96 634500.71 553390.323 -50569.187 24.2

LOC01 9 07:22:03 20/10/2008 204958.59 634496.34 553390.137 -50568.159 25.2

LOC01 10 07:22:33 20/10/2008 204969.26 634491.33 553389.866 -50567.147 26.2

LOC01 11 07:22:53 20/10/2008 204974.66 634487.83 553389.731 -50566.566 26.9

LOC01 12 07:23:23 20/10/2008 204981.97 634483.65 553389.547 -50565.843 27.2

LOC01 13 07:23:40 20/10/2008 204985.97 634481.52 553389.442 -50565.455 27.4

LOC01 14 07:24:15 20/10/2008 204993.52 634476.97 553389.220 -50564.708 27.4

LOC01 15 07:24:37 20/10/2008 204997.58 634473.78 553389.071 -50564.287 27.3

LOC01 16 07:25:13 20/10/2008 205003.84 634469.49 553388.858 -50563.678 27.3

LOC01 17 07:25:23 20/10/2008 205004.32 634467.51 553388.784 -50563.572 27.2

LOC01 18 07:26:50 20/10/2008 205036.20 634453.54 553388.044 -50560.579 27.5

LOC01 19 07:27:20 20/10/2008 205047.30 634448.27 553387.776 -50559.499 27.8

LOC01 20 07:27:42 20/10/2008 205054.30 634443.23 553387.556 -50558.779 27.7

LOC01 21 07:27:59 20/10/2008 205059.95 634438.91 553387.359 -50558.202 27.5

LOC01 22 07:28:12 20/10/2008 205063.51 634436.21 553387.229 -50557.851 27.5

LOC01 23 07:28:26 20/10/2008 205068.07 634433.38 553387.079 -50557.423 27.5

LOC01 24 07:28:36 20/10/2008 205070.52 634431.21 553386.974 -50557.155 27.6

LOC01 25 07:30:40 20/10/2008 205110.52 634397.13 553385.251 -50553.171 27.0

LOC01 26 07:31:01 20/10/2008 205116.88 634390.76 553384.932 -50552.540 27.1

LOC01 27 07:31:29 20/10/2008 205125.82 634381.50 553384.459 -50551.638 27.5

LOC01 28 07:32:00 20/10/2008 205134.97 634371.60 553383.947 -50550.723 27.7

LOC01 29 07:32:20 20/10/2008 205141.51 634364.91 553383.604 -50550.083 28.0

LOC01 30 07:33:09 20/10/2008 205159.30 634348.63 553382.743 -50548.366 28.6

LOC01 31 07:33:37 20/10/2008 205170.27 634342.93 553382.422 -50547.356 28.7

LOC01 32 07:35:30 20/10/2008 205208.49 634323.03 553381.434 -50543.665 28.7

LOC01 33 07:36:41 20/10/2008 205232.88 634309.14 553380.756 -50541.272 28.4

LOC01 34 07:37:31 20/10/2008 205247.50 634299.81 553380.277 -50539.843 27.9

LOC01 35 07:38:15 20/10/2008 205262.30 634289.68 553379.783 -50538.376 27.3

LOC01 36 07:38:40 20/10/2008 205271.88 634283.71 553379.490 -50537.449 27.0

LOC01 37 07:39:15 20/10/2008 205282.81 634277.62 553379.173 -50536.390 26.7

J/08/151SV Alma Na MaraOctober 2008

Page 114: Mapping of the marine habitats and species of Lamlash Bay, Arran

102

LOC01 38 07:40:00 20/10/2008 205296.81 634270.24 553378.816 -50535.027 26.6

LOC01 39 07:40:44 20/10/2008 205312.25 634261.88 553378.389 -50533.544 26.4

LOC01 40 07:41:22 20/10/2008 205326.90 634256.68 553378.104 -50532.175 26.3

LOC01 41 07:42:04 20/10/2008 205336.90 634253.62 553377.969 -50531.194 26.2

LOC01 42 07:42:20 20/10/2008 205340.94 634252.42 553377.910 -50530.815 26.1

LOC01 43 07:42:47 20/10/2008 205347.37 634250.40 553377.819 -50530.212 25.8

LOC01 44 07:43:19 20/10/2008 205356.05 634249.42 553377.736 -50529.380 25.9

LOC01 45 07:43:54 20/10/2008 205365.02 634246.42 553377.623 -50528.505 25.7

LOC01 46 07:44:27 20/10/2008 205373.16 634242.37 553377.472 -50527.709 25.3

LOC01 47 07:44:37 20/10/2008 205376.17 634242.27 553377.429 -50527.415 25.3

LOC01 48 07:44:57 20/10/2008 205382.24 634240.09 553377.337 -50526.818 25.1

LOC01 49 07:45:27 20/10/2008 205392.13 634235.84 553377.153 -50525.871 24.7

LOC01 50 07:45:43 20/10/2008 205396.99 634233.09 553377.042 -50525.378 24.3

LOC01 51 07:45:46 20/10/2008 205397.67 634232.69 553377.022 -50525.297 24.3

LOC01 52 07:46:04 20/10/2008 205401.46 634228.50 553376.867 -50524.813 23.7

LOC01 53 07:46:11 20/10/2008 205404.93 634228.79 553376.830 -50524.607 23.6

LOC01 54 07:46:29 20/10/2008 205410.67 634226.94 553376.715 -50524.063 23.5

LOC01 55 07:46:44 20/10/2008 205415.55 634225.60 553376.640 -50523.607 23.6

LOC01 56 07:47:11 20/10/2008 205422.71 634222.80 553376.530 -50522.922 24.5

LOC01 57 07:47:45 20/10/2008 205429.75 634220.36 553376.406 -50522.243 25.4

LOC01 58 07:47:59 20/10/2008 205431.86 634219.14 553376.366 -50522.013 25.7

LOC01 59 07:49:34 20/10/2008 205450.42 634210.05 553375.968 -50520.159 26.6

LOC01 60 07:49:54 20/10/2008 205456.67 634205.48 553375.763 -50519.488 27.3

LOC01 61 07:50:41 20/10/2008 205474.79 634198.04 553375.338 -50517.830 29.0

LOC01 62 07:51:09 20/10/2008 205484.28 634194.76 553375.169 -50516.944 29.9

LOC01 63 07:51:30 20/10/2008 205490.94 634192.81 553375.074 -50516.297 30.9

LOC01 64 07:52:02 20/10/2008 205501.16 634192.88 553375.023 -50515.381 32.1

LOC01 65 07:52:57 20/10/2008 205516.06 634188.86 553374.894 -50513.920 34.1

LOC01 66 07:54:03 20/10/2008 205531.06 634187.68 553374.814 -50512.510 35.9

LOC01 67 07:55:11 20/10/2008 205541.74 634188.03 553374.711 -50511.497 37.2

LOC01 68 07:55:36 20/10/2008 205544.68 634188.31 553374.747 -50511.217 37.5

LOC01 69 07:56:11 20/10/2008 205548.57 634181.34 553374.655 -50510.706 38.1

LOC01 70 07:56:36 20/10/2008 205553.88 634177.85 553374.476 -50510.191 38.8

LOC01 71 07:56:59 20/10/2008 205559.68 634175.38 553374.320 -50509.683 39.4

LOC01 72 07:57:36 20/10/2008 205568.01 634173.70 553374.113 -50508.945 40.0

LOC01 73 07:58:11 20/10/2008 205577.57 634172.72 553374.039 -50508.030 40.5

LOC01 74 07:58:45 20/10/2008 205586.88 634167.83 553373.900 -50507.108 40.8

LOC01 75 07:59:15 20/10/2008 205596.01 634165.87 553373.823 -50506.245 41.0

LOC01 76 08:00:12 20/10/2008 205611.17 634170.43 553373.958 -50504.814 41.3

LOC01 77 08:00:55 20/10/2008 205625.21 634174.61 553374.203 -50503.495 41.4

LOC01 78 08:01:37 20/10/2008 205641.36 634184.68 553374.724 -50501.974 41.2

LOC01 79 08:02:30 20/10/2008 205653.35 634200.80 553375.525 -50500.743 41.3

LOC02 2 08:18:45 20/10/2008 205713.01 633555.50 553341.083 -50492.528 32.8

LOC02 3 08:19:03 20/10/2008 205719.12 633560.27 553341.319 -50491.964 33.0

LOC02 4 08:19:38 20/10/2008 205731.18 633566.60 553341.687 -50490.851 33.7

LOC02 5 08:19:49 20/10/2008 205734.98 633568.81 553341.808 -50490.485 33.9

Page 115: Mapping of the marine habitats and species of Lamlash Bay, Arran

103

LOC02 6 08:20:13 20/10/2008 205742.11 633573.51 553342.046 -50489.760 34.5

LOC02 7 08:22:56 20/10/2008 205796.96 633598.56 553343.507 -50484.672 35.2

LOC02 8 08:24:04 20/10/2008 205821.34 633611.47 553344.308 -50482.449 35.2

LOC02 9 08:25:36 20/10/2008 205863.68 633623.54 553345.160 -50478.541 34.8

LOC02 10 08:26:22 20/10/2008 205884.65 633628.66 553345.447 -50476.560 34.5

LOC02 11 08:26:47 20/10/2008 205895.78 633632.95 553345.682 -50475.521 34.5

LOC02 12 08:27:19 20/10/2008 205909.94 633637.38 553345.985 -50474.186 35.1

LOC02 13 08:28:00 20/10/2008 205926.19 633644.57 553346.370 -50472.649 35.5

LOC02 14 08:28:29 20/10/2008 205936.63 633649.17 553346.642 -50471.672 35.8

LOC02 15 08:29:55 20/10/2008 205972.24 633664.85 553347.574 -50468.361 38.7

LOC02 16 08:30:12 20/10/2008 205980.06 633666.22 553347.696 -50467.626 39.1

LOC02 17 08:30:28 20/10/2008 205986.30 633667.58 553347.787 -50467.050 39.3 No flashLOC02 18 08:30:50 20/10/2008 205995.28 633670.94 553347.955 -50466.248 39.1 No flashLOC02 19 08:31:23 20/10/2008 206006.11 633677.96 553348.317 -50465.151 39.8 No flashLOC04 2 09:17:26 20/10/2008 206058.82 632401.50 553279.684 -50454.834 23.3

LOC04 3 09:17:44 20/10/2008 206071.70 632409.64 553280.153 -50453.640 22.9

LOC04 4 09:18:02 20/10/2008 206084.61 632417.07 553280.599 -50452.452 22.8

LOC04 5 09:18:15 20/10/2008 206093.70 632422.38 553280.911 -50451.617 22.6

LOC04 6 09:18:30 20/10/2008 206103.97 632428.30 553281.247 -50450.685 22.4

LOC04 7 09:18:44 20/10/2008 206112.83 632433.86 553281.563 -50449.845 22.7

LOC04 8 09:18:53 20/10/2008 206118.77 632437.53 553281.777 -50449.307 22.8

LOC04 9 09:19:08 20/10/2008 206128.23 632443.86 553282.133 -50448.415 22.9

LOC04 10 09:19:16 20/10/2008 206133.50 632447.17 553282.325 -50447.934 23.1

LOC04 11 09:19:34 20/10/2008 206144.38 632455.22 553282.768 -50446.901 23.5

LOC04 12 09:20:00 20/10/2008 206160.52 632467.82 553283.467 -50445.440 23.8

LOC04 13 09:20:11 20/10/2008 206167.12 632473.15 553283.774 -50444.822 23.9

LOC04 14 09:20:26 20/10/2008 206177.02 632480.05 553284.174 -50443.947 24.6

LOC04 15 09:21:02 20/10/2008 206199.83 632496.99 553285.142 -50441.806 26.3

LOC04 16 09:21:29 20/10/2008 206218.90 632510.10 553285.900 -50440.094 27.9

LOC04 17 09:21:49 20/10/2008 206235.39 632520.35 553286.493 -50438.606 29.6

LOC04 18 09:22:03 20/10/2008 206246.57 632526.24 553286.859 -50437.563 31.1

LOC04 19 09:22:20 20/10/2008 206259.66 632532.80 553287.254 -50436.370 32.9

LOC04 20 09:22:42 20/10/2008 206276.13 632541.12 553287.744 -50434.816 35.5

LOC04 21 09:23:27 20/10/2008 206303.77 632563.43 553288.922 -50432.147 40.1

LOC04 22 09:24:00 20/10/2008 206326.00 632577.90 553289.767 -50430.155 43.7

LOC04 23 09:24:20 20/10/2008 206339.27 632591.60 553290.549 -50428.975 45.9

LOC04 24 09:24:43 20/10/2008 206355.79 632603.43 553291.348 -50427.521 48.7

LOC04 25 09:25:18 20/10/2008 206380.20 632628.48 553292.618 -50425.217 51.3

Page 116: Mapping of the marine habitats and species of Lamlash Bay, Arran

104

Appendix 03: Lamlash Bay Sidescan sonar and Bathymetry log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m)

Client SNH Job Number

Location Lamlash Bay Vessel

Survey Lamlash Bay 2009 habitat mapping survey Date

Depth Depth

Date Area SSS Range Line SOL Time EOL Time Easting (m) Northing (m)Corrected (m) Easting (m) Northing (m) Corrected (m) Comments

10/09/2009 A 100m A09 16:42:04 16:48:49 206255.63 632528.73 31.59 205368.29 631863.53 15.22 16:46 - offline for buoy

10/09/2009 A 50m A08 16:51:41 16:59:33 205315.62 631881.19 15.46 206156.97 632528.08 22.18

10/09/2009 A 50m A07 17:08:50 17:15:02 206138.08 632567.14 22.29 205308.60 631950.79 17.46

10/09/2009 A 50m A06 17:18:12 17:26:02 205231.07 631947.56 18.87 206103.25 632611.81 22.48

10/09/2009 A 50m A05 17:35:06 17:42:11 206097.86 632660.37 24.72 205242.93 632021.52 20.37

10/09/2009 A 50m A04 17:45:00 17:52:47 205167.31 632028.83 17.91 206073.45 632701.78 25.34

10/09/2009 A 50m A03 17:56:44 18:04:14 206097.21 632787.17 30.73 205204.02 632106.12 19.49

10/09/2009 A 50m A02 18:06:55 18:14:30 205142.00 632132.74 17.66 206032.57 632799.39 28.44

11/09/2009 A 75m A01 07:06:21 07:14:06 205057.05 632134.29 16.98 206013.70 632861.77 32.07

11/09/2009 G 75m G06 07:26:48 07:30:42 206173.13 631758.09 26.36 205688.37 631404.23 4.14

11/09/2009 G 75m G05 07:34:40 07:36:52 205907.48 631609.71 11.28 205633.29 631413.08 4.00 Line cut short - too shallow

11/09/2009 G 75m G04 07:40:31 07:43:30 205933.69 631669.34 19.87 205551.79 631410.30 5.27

11/09/2009 G 75m G03 07:47:26 07:50:38 205863.58 631698.44 13.05 205450.13 631396.84 4.70

11/09/2009 G 75m G02 07:55:46 07:59:50 205823.36 631733.48 12.53 205300.23 631348.54 4.59

11/09/2009 G 75m G01 08:05:02 08:11:18 205785.58 631765.83 12.68 204972.89 631165.99 14.55

11/09/2009 F 75m F01 08:24:23 08:31:47 205080.92 631288.31 17.10 205104.66 630189.49 31.44 08:27 - offline to avoid anchored yacht

11/09/2009 F 75m F02 08:35:15 08:42:43 205052.74 630133.81 40.48 205029.13 631288.79 18.25

11/09/2009 F 75m F03 08:45:08 08:52:49 204982.62 631350.48 25.11 205006.32 630142.25 42.30

11/09/2009 H 75m H06 08:56:49 09:03:55 205139.65 629990.97 39.13 205872.71 629247.80 32.14

11/09/2009 H 75m H05 09:05:29 09:12:06 205875.85 629176.07 32.03 205168.11 629901.75 43.41

11/09/2009 H 75m H04 19:14:41 09:23:06 205093.81 629896.04 44.73 205941.45 629045.65 30.79

11/09/2009 H 75m H03 09:28:08 09:35:53 205925.64 628987.07 28.74 205094.07 629831.66 46.49

11/09/2009 H 75m H02 09:39:18 09:49:06 205050.21 629802.02 45.39 206000.23 628834.14 23.09

11/09/2009 H 75m H01 10:01:54 10:10:29 205968.70 628799.21 27.23 205020.42 629756.53 45.41

11/09/2009 B 75m B06 10:39:17 10:52:34 204106.43 631453.36 18.52 205751.18 632679.33 7.02

11/09/2009 B 75m B05 10:54:51 11:16:16 205697.19 632703.48 5.87 203060.01 630757.23 23.74

11/09/2009 B 75m B04 11:18:00 11:38:00 203098.05 630840.53 25.25 205558.43 632672.51 4.31

11/09/2009 B 75m B03 11:39:05 11:56:36 205503.30 632683.63 4.09 203308.60 631058.08 19.61

11/09/2009 B 75m B02 11:58:05 12:15:26 203230.74 631072.66 14.08 205365.01 632640.73 3.54

11/09/2009 B 75m B01 12:18:00 12:33:24 205356.37 632705.01 3.14 203476.49 631310.60 19.57 12:24 - offline for mooring buoys, 12:30 - offline for passing yacht

11/09/2009 E 75m E10 13:09:38 13:22:30 205759.25 627492.05 30.98 207459.38 628309.72 53.96

11/09/2009 E 75m E09 13:25:20 13:39:18 207447.66 628357.13 56.57 205702.43 627525.01 25.77

11/09/2009 E 75m E08 13:41:51 13:54:55 205723.46 627585.75 25.15 207531.98 628432.02 61.18

11/09/2009 E 75m E07 13:55:55 14:11:14 207532.99 628511.62 62.71 205662.36 627610.67 19.09

11/09/2009 E 75m E06 14:29:39 14:42:06 205676.29 627685.23 20.63 207382.69 628495.25 49.78

11/09/2009 E 75m E05 14:54:13 15:08:34 207340.20 628524.73 43.01 205637.76 627713.58 8.33

11/09/2009 E 75m E04 15:14:03 15:26:43 205636.33 627780.31 6.57 207349.44 628582.37 43.74

11/09/2009 E 75m E03 15:30:06 15:44:28 207300.90 628642.80 39.94 205617.47 627809.69 4.75

11/09/2009 E 75m E02 15:45:21 15:57:44 205609.21 627866.74 3.20 207305.29 628674.98 40.09

11/09/2009 E 75m E01 16:09:26 16:23:29 207441.94 628795.09 54.52 205656.07 627935.28 4.36

Bathy and Sidescan Sonar Lines Time (GMT)

J/09/182

SV Otarie

September 2009

SOL Position EOL Position

Page 117: Mapping of the marine habitats and species of Lamlash Bay, Arran

105

11/09/2009 D 75m D01 16:37:20 16:40:47 206978.86 629189.04 16.53 206773.19 629671.85 5.58 Line stopped - too shallow and line D02 started 16:40

11/09/2009 D 75m D02 16:40:59 16:42:55 206774.93 629704.30 6.30 206661.87 629960.72 3.47 Line stopped - too shallow and line D03 started 16:43

11/09/2009 D 75m D03 16:43:38 16:56:55 206661.01 630079.09 17.53 205824.17 631848.6 19.74

12/09/2009 D 75m D04 07:39:36 07:58:50 205891.60 631828.24 22.76 207095.72 629269.78 36.43

12/09/2009 D 75m D05 08:01:54 08:20:42 207114.63 629343.72 37.37 205907.50 631910.47 25.35

12/09/2009 D 75m D06 08:29:17 08:47:53 205945.46 631951.49 27.39 207093.67 629503.88 37.22

12/09/2009 D 75m D07 08:52:14 09:10:19 207072.37 629669.84 38.02 205979.73 631986.66 27.57

12/09/2009 D 75m D08 09:21:41 09:29:03 206625.53 630736.22 41.11 207069.15 629791.54 41.47 09:25 - offline for pots

12/09/2009 D 75m D09 09:31:38 09:47:00 207038.59 629992.94 44.09 206061.67 632051.54 26.45 towfish on a short tow at the start of line to avoid pots

12/09/2009 D 75m D10 09:48:52 10:02:38 206091.90 632109.84 24.61 207026.16 630115.56 46.29

12/09/2009 D 75m D11 10:04:34 10:19:54 207009.17 630276.97 50.98 206146.60 632105.45 24.00 10:15 - offline for passing yacht

12/09/2009 D 75m D12 10:22:13 10:34:46 206183.47 632171.13 22.63 206996.49 630422.86 51.60

12/09/2009 D 75m D13 10:37:08 10:49:26 206984.12 630563.82 51.24 206232.80 632156.69 25.65

12/09/2009 D 75m D14 10:51:56 11:02:38 206260.91 632236.03 29.12 206957.98 630736.10 52.78

12/09/2009 D 75m D15 11:04:33 11:14:20 206939.44 630895.12 51.62 206300.16 632245.27 32.67

12/09/2009 D 75m D16 11:16:58 11:24:59 206388.03 632176.36 37.85 206920.56 631054.56 34.63

12/09/2009 D 75m D17 11:26:59 11:34:48 206904.32 631204.56 28.15 206396.55 632274.67 40.23

12/09/2009 D 75m D18 11:37:27 11:42:35 206464.97 632251.29 41.58 206791.70 631559.05 49.21

13/09/2009 I 75m I06 11:01:53 11:17:26 205726.19 633159.86 25.07 204500.90 634987.43 11.35

13/09/2009 I 75m I05 11:18:46 11:32:50 204451.77 634965.52 6.76 205685.70 633127.96 15.67

13/09/2009 I 75m I04 11:37:43 11:51:39 205510.80 633316.70 6.93 204415.93 634928.75 4.77

13/09/2009 I 75m I03 11:53:05 12:01:03 204407.91 634893.16 3.49 205031.47 633931.50 7.95 No lines run inshore of I03 - too shallow

13/09/2009 C 75m C09 12:38:22 12:49:59 202944.90 630892.10 10.10 204155.74 629680.74 38.42

13/09/2009 C 75m C08 12:53:01 13:04:15 204088.81 629660.01 32.17 202936.78 630846.70 12.19 12:58:30 - offline for pot marker

13/09/2009 C 75m C07 13:05:36 13:16:51 202908.47 630796.91 9.02 204059.06 629630.83 31.44

13/09/2009 C 75m C06 13:20:23 13:30:37 203918.48 629700.57 24.41 202887.27 630753.05 3.36 Line started to short to keep clear of mussel farm

13/09/2009 C 75m C05 13:32:56 13:37:50 202884.06 630676.99 2.62 203391.56 630172.59 31.64 No lines run inshore of C05 because of buoys and pot markers

13/09/2009 D 75m D03a 14:07:58 14:14:54 207044.84 629267.48 31.23 206630.23 630145.84 18.46 Southern end infill

13/09/2009 D 75m D02a 14:26:15 14:31:35 207011.17 629211.75 25.32 206700.91 629879.96 5.86 Southern end infill

13/09/2009 D 75m D02b 14:56:17 15:01:12 205780.73 631821.86 15.13 206089.38 631162.66 4.90 Northern end infill

Page 118: Mapping of the marine habitats and species of Lamlash Bay, Arran

106

Appendix 03: Lamlash Bay Video log - 2009SeaStar Survey - Project J/09/182

Client SNH Job Number J/09/182

Location Lamlash Bay Vessel SV Otarie

Survey Date September 2009

DATE Site Location Time Video DVD Number Depth DVD

Name (GMT) Duration No. of Photos East (m) North (m) (m) Number

09/09/2009 NTZ01 SOL 13:12:52 0:11:22 1 17 205275.91 632650.18 3.8

EOL 13:24:14 205442.23 632596.01 4.5

09/09/2009 NTZ02 SOL 13:36:14 0:12:24 1 15 205573.60 632384.17 5.3

EOL 13:48:38 205722.95 632259.92 8.4

09/09/2009 NTZ03 SOL 14:18:34 0:18:44 1 22 205756.69 631614.51 9.6

EOL 14:37:18 205892.03 631448.54 7.4

09/09/2009 NTZ05 SOL 14:50:59 0:15:22 2 21 204919.86 631163.64 17.5

EOL 15:06:21 205010.66 631008.30 17.1

09/09/2009 NTZ04 SOL 15:25:14 0:17:28 2 22 204368.72 631857.45 5.7

EOL 15:42:42 204548.56 631707.49 14.3

12/09/2009 A01 SOL 13:43:28 0:25:01 3 27 205286.16 632379.48 8.0

EOL 14:08:29 205102.30 632448.70 4.2

12/09/2009 A02 SOL 14:23:20 0:14:57 3 15 205890.23 632633.40 17.7

EOL 14:38:17 205822.25 632627.42 10.3

12/09/2009 A03 SOL 14:46:07 0:08:50 4 9 205830.36 632763.66 11.3

EOL 14:54:57 205772.94 632727.29 3.4

12/09/2009 H01 SOL 15:32:36 0:38:33 4 40 205405.94 629542.12 36.3

EOL 16:11:09 205284.42 629947.14 17.0

Positions are OSGB36 East and North (m)

Lamlash Bay 2009 habitat mapping survey

Position

Page 119: Mapping of the marine habitats and species of Lamlash Bay, Arran

107

13/09/2009 F01 SOL 07:50:06 0:15:20 5 18 205121.62 630532.47 2.7

EOL 08:05:26 205044.79 630436.39 18.5

13/09/2009 E01 SOL 08:31:42 0:21:10 5 27 206487.85 628176.72 10.8

EOL 08:52:52 206540.62 627852.28 12.7

13/09/2009 E02 SOL 09:11:42 0:10:42 5 15 207041.95 628592.35 13.1

EOL 09:22:24 206965.58 628387.62 16.0

13/09/2009 D01 SOL 09:52:20 0:11:53 5 11 206633.76 630578.21 45.6

EOL 10:04:13 206605.89 630483.40 42.7

13/09/2009 D03 SOL 10:24:17 0:13:53 6 15 206316.36 630960.11 12.0

EOL 10:38:10 206276.79 630861.24 8.8

13/09/2009 D02 SOL 15:50:32 0:11:06 6 9 206566.06 631666.42 47.6

EOL 16:01:38 206522.49 631616.79 45.5

13/09/2009 B01 SOL 16:26:32 0:09:46 6 11 203540.33 631204.80 23.0

EOL 16:36:18 203513.67 631219.10 22.2

Page 120: Mapping of the marine habitats and species of Lamlash Bay, Arran

108

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

NTZ01 1 deck 09/09/2009

NTZ01 2 13:13:20 09/09/2009 205282.87 632649.34 4.9

NTZ01 3 13:14:19 09/09/2009 205298.53 632645.99 4.8

NTZ01 4 13:15:01 09/09/2009 205309.02 632643.19 4.8

NTZ01 5 13:16:34 09/09/2009 205332.34 632638.12 4.7

NTZ01 6 13:16:50 09/09/2009 205335.10 632635.94 4.4

NTZ01 7 13:18:00 09/09/2009 205351.07 632627.70 4.8

NTZ01 8 13:18:09 09/09/2009 205354.01 632627.53 4.7

NTZ01 9 13:18:38 09/09/2009 205361.66 632627.76 4.6

NTZ01 10 13:19:04 09/09/2009 205367.47 632628.43 4.7

NTZ01 11 13:19:56 09/09/2009 205378.96 632623.82 4.9

NTZ01 12 13:20:44 09/09/2009 205384.32 632620.19 4.7

NTZ01 13 13:21:08 09/09/2009 205392.23 632609.26 5.0

NTZ01 14 13:21:40 09/09/2009 205400.18 632603.79 4.9

NTZ01 15 13:22:14 09/09/2009 205411.05 632600.30 5.0

NTZ01 16 13:22:59 09/09/2009 205423.82 632599.01 5.0

NTZ01 17 13:23:50 09/09/2009 205436.17 632597.95 5.4

NTZ02 1 deck 09/09/2009

NTZ02 2 13:38:47 09/09/2009 205614.87 632362.58 6.5

NTZ02 3 13:39:08 09/09/2009 205618.72 632361.89 7.9

NTZ02 4 13:40:06 09/09/2009 205630.70 632355.62 6.9

NTZ02 5 13:40:43 09/09/2009 205640.92 632349.55 7.3

NTZ02 6 13:41:12 09/09/2009 205645.20 632348.14 7.3

NTZ02 7 13:41:26 09/09/2009 205647.20 632348.02 8.2

NTZ02 8 13:41:47 09/09/2009 205650.24 632346.91 7.5

NTZ02 9 13:42:45 09/09/2009 205662.73 632339.16 8.6

SNH

No Take Zone

Lamlash Bay 2009 habitat mapping survey

Page 121: Mapping of the marine habitats and species of Lamlash Bay, Arran

109

NTZ02 10 13:43:05 09/09/2009 205666.41 632338.34 8.4

NTZ02 11 13:43:33 09/09/2009 205670.37 632338.68 7.7

NTZ02 12 13:43:55 09/09/2009 205673.50 632335.82 8.8

NTZ02 13 13:45:51 09/09/2009 205696.59 632303.42 9.5

NTZ02 14 13:47:44 09/09/2009 205714.14 632271.79 10.4

NTZ02 15 13:48:02 09/09/2009 205716.43 632268.74 9.7

NTZ03 1 deck 09/09/2009

NTZ03 2 14:03:21 09/09/2009 Line stopped

NTZ03a 1 deck 09/09/2009

NTZ03a 2 14:20:34 09/09/2009 205775.33 631592.09 8.2

NTZ03a 3 14:21:49 09/09/2009 205779.18 631582.49 7.9

NTZ03a 4 14:22:36 09/09/2009 205782.90 631574.04 7.7

NTZ03a 5 14:23:12 09/09/2009 205786.97 631569.68 7.5

NTZ03a 6 14:23:32 09/09/2009 205786.62 631567.95 7.8

NTZ03a 7 14:24:20 09/09/2009 205791.66 631561.60 7.9

NTZ03a 8 14:25:34 09/09/2009 205801.09 631551.17 8.1

NTZ03a 9 14:26:13 09/09/2009 205805.77 631550.07 7.9

NTZ03a 10 14:27:02 09/09/2009 205809.52 631541.07 7.7

NTZ03a 11 14:28:26 09/09/2009 205818.86 631527.10 8.2

NTZ03a 12 14:28:59 09/09/2009 205822.84 631519.65 8.3

NTZ03a 13 14:29:50 09/09/2009 205828.98 631511.41 8.3

NTZ03a 14 14:30:56 09/09/2009 205837.10 631498.25 8.2

NTZ03a 15 14:31:35 09/09/2009 205838.82 631492.09 8.1

NTZ03a 16 14:32:13 09/09/2009 205842.80 631485.27 / Bad sounding data.

NTZ03a 17 14:32:50 09/09/2009 205847.79 631479.28 8.0

NTZ03a 18 14:33:27 09/09/2009 205851.46 631471.94 8.1

NTZ03a 19 14:34:24 09/09/2009 205860.81 631466.54 8.2

NTZ03a 20 14:35:14 09/09/2009 205869.10 631459.42 8.2

NTZ03a 21 14:35:48 09/09/2009 205875.79 631456.98 8.2

NTZ03a 22 14:36:34 09/09/2009 205884.64 631452.45 8.2

NTZ05 1 deck 09/09/2009

NTZ05 2 14:51:28 09/09/2009 204919.57 631158.16 18.7

NTZ05 3 14:52:00 09/09/2009 204918.55 631150.72 18.6

NTZ05 4 14:52:44 09/09/2009 204921.84 631142.20 18.4

NTZ05 5 14:53:20 09/09/2009 204921.75 631136.96 18.3

NTZ05 6 14:53:56 09/09/2009 204922.48 631131.38 18.2

NTZ05 7 14:54:28 09/09/2009 204923.53 631124.66 18.1

Page 122: Mapping of the marine habitats and species of Lamlash Bay, Arran

110

NTZ05 8 14:55:03 09/09/2009 204926.06 631117.06 18.0

NTZ05 9 14:55:59 09/09/2009 204929.22 631105.97 17.8

NTZ05 10 14:56:52 09/09/2009 204931.59 631096.11 17.7

NTZ05 11 14:57:33 09/09/2009 204936.33 631087.20 17.6

NTZ05 12 14:58:37 09/09/2009 204939.11 631074.61 17.5

NTZ05 13 14:59:10 09/09/2009 204942.90 631065.47 17.5

NTZ05 14 15:00:06 09/09/2009 204951.21 631053.57 17.6

NTZ05 15 15:00:49 09/09/2009 204956.72 631044.44 17.6

NTZ05 16 15:01:32 09/09/2009 204965.71 631037.75 17.7

NTZ05 17 15:01:57 09/09/2009 204968.14 631035.69 17.7

NTZ05 18 15:02:44 09/09/2009 204972.38 631031.38 17.8

NTZ05 19 15:03:30 09/09/2009 204979.38 631026.21 17.9

NTZ05 20 15:04:15 09/09/2009 204988.27 631020.44 18.0

NTZ05 21 15:05:58 09/09/2009 205006.75 631010.19 17.9

NTZ04 1 deck 09/09/2009

NTZ04 2 15:26:36 09/09/2009 204387.88 631842.97 6.9

NTZ04 3 15:27:08 09/09/2009 204398.66 631840.31 6.9

NTZ04 4 15:27:46 09/09/2009 204407.88 631838.90 6.8

NTZ04 5 15:28:26 09/09/2009 204416.10 631834.19 6.9

NTZ04 6 15:30:00 09/09/2009 204436.81 631826.12 7.0

NTZ04 7 15:30:52 09/09/2009 204445.52 631820.08 7.3

NTZ04 8 15:31:55 09/09/2009 204459.50 631814.65 7.6

NTZ04 9 15:32:32 09/09/2009 204465.66 631808.07 8.0

NTZ04 10 15:33:19 09/09/2009 204476.76 631795.04 8.4

NTZ04 11 15:34:27 09/09/2009 204487.39 631782.08 8.8

NTZ04 12 15:35:03 09/09/2009 204494.61 631778.32 8.9

NTZ04 13 15:36:00 09/09/2009 204501.42 631772.09 9.2

NTZ04 14 15:36:19 09/09/2009 204503.27 631769.50 9.5

NTZ04 15 15:36:55 09/09/2009 204510.36 631766.71 9.8

NTZ04 16 15:37:30 09/09/2009 204514.65 631762.51 10.2

NTZ04 17 15:38:30 09/09/2009 204521.88 631755.39 10.9

NTZ04 18 15:39:22 09/09/2009 204529.81 631745.84 11.6

NTZ04 19 15:40:17 09/09/2009 204536.88 631734.96 12.6

NTZ04 20 15:40:51 09/09/2009 204541.67 631728.79 13.0

NTZ04 21 15:41:48 09/09/2009 204544.88 631716.99 14.3

NTZ04 22 15:42:24 09/09/2009 204547.21 631710.84 15.0

Page 123: Mapping of the marine habitats and species of Lamlash Bay, Arran

111

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

A01 1 deck 12/09/2009

A01 2 13:44:06 12/09/2009 205280.62 632381.77 8.5

A01 3 13:45:27 12/09/2009 205271.95 632390.09 8.1

A01 4 13:45:47 12/09/2009 205269.56 632388.54 7.9

A01 5 13:46:42 12/09/2009 205264.46 632391.26 7.7

A01 6 13:47:49 12/09/2009 205259.33 632394.34 7.6

A01 7 13:48:09 12/09/2009 205257.66 632395.20 7.5

A01 8 13:48:44 12/09/2009 205253.84 632398.73 7.2

A01 9 13:49:52 12/09/2009 205247.02 632404.27 6.9

A01 10 13:50:50 12/09/2009 205238.47 632406.89 7.2

A01 11 13:51:41 12/09/2009 205231.36 632409.72 7.1

A01 12 13:52:33 12/09/2009 205223.75 632412.85 7.0

A01 13 13:53:53 12/09/2009 205211.84 632417.83 7.0

A01 14 13:54:54 12/09/2009 205203.00 632419.41 6.9

A01 15 13:56:14 12/09/2009 205191.10 632422.82 6.9

A01 16 13:57:03 12/09/2009 205184.58 632425.11 / Bad sounding data.

A01 17 13:58:10 12/09/2009 205175.66 632430.23 6.0

A01 18 13:58:44 12/09/2009 205171.64 632430.72 6.3

A01 19 13:59:57 12/09/2009 205163.65 632434.08 6.7

A01 20 14:00:50 12/09/2009 205158.11 632431.12 5.7

A01 21 14:01:58 12/09/2009 205149.48 632433.23 5.8

A01 22 14:02:55 12/09/2009 205141.15 632436.65 6.7

A01 23 14:03:30 12/09/2009 205136.95 632437.17 6.6

A01 24 14:05:06 12/09/2009 205124.36 632440.77 6.5

A01 25 14:06:13 12/09/2009 205116.06 632442.25 6.4

SNH

Area A

Lamlash Bay 2009 habitat mapping survey

Page 124: Mapping of the marine habitats and species of Lamlash Bay, Arran

112

A01 26 14:07:12 12/09/2009 205110.05 632444.90 6.4

A01 27 14:08:22 12/09/2009 205103.00 632448.17 6.3

A02 1 deck 12/09/2009

A02 2 14:24:47 12/09/2009 205886.59 632631.59 15.7

A02 3 14:26:00 12/09/2009 205882.91 632629.15 17.8

A02 4 14:27:12 12/09/2009 205880.80 632627.02 16.5

A02 5 14:28:41 12/09/2009 205877.42 632625.00 16.5

A02 6 14:30:06 12/09/2009 205873.14 632626.65 16.3

A02 7 14:31:09 12/09/2009 205870.94 632624.86 16.5

A02 8 14:33:03 12/09/2009 205858.33 632624.77 15.5

A02 9 14:34:42 12/09/2009 205833.45 632629.37 12.7

A02 10 14:35:17 12/09/2009 205831.01 632627.55 12.3

A02 11 14:35:56 12/09/2009 205828.08 632625.46 11.8

A02 12 14:36:21 12/09/2009 205826.10 632625.15 11.7

A02 13 14:37:06 12/09/2009 205824.87 632626.11 11.5

A02 14 14:37:40 12/09/2009 205823.06 632626.77 11.0

A02 15 14:38:05 12/09/2009 205822.47 632627.33 11.2 Line A02 stopped

A03 1 deck 12/09/2009

A03 2 14:47:11 12/09/2009 205826.51 632766.19 10.6

A03 3 14:48:24 12/09/2009 205817.89 632768.71 9.4

A03 4 14:48:48 12/09/2009 205814.11 632769.58 9.4

A03 5 14:49:43 12/09/2009 205807.32 632763.64 9.6

A03 6 14:50:37 12/09/2009 205798.53 632755.87 9.4

A03 7 14:51:22 12/09/2009 205794.18 632749.68 / Bad sounding data.

A03 8 14:51:54 12/09/2009 205793.09 632741.80 / Bad sounding data.

A03 9 15:52:51 12/09/2009 205794.93 632733.45 7.2

Page 125: Mapping of the marine habitats and species of Lamlash Bay, Arran

113

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

B01 1 deck 13/09/2009

B01 2 16:27:40 13/09/2009 203536.99 631208.30 22.9

B01 3 16:28:50 13/09/2009 203536.02 631212.42 22.9

B01 4 16:29:44 13/09/2009 203532.64 631213.28 23.7

B01 5 16:30:52 13/09/2009 203529.31 631212.96 21.5

B01 6 16:31:59 13/09/2009 203526.38 631216.22 22.9

B01 7 16:32:39 13/09/2009 203525.55 631218.45 23.7

B01 8 13:33:55 13/09/2009 203520.43 631219.72 23.0

B01 9 16:34:12 13/09/2009 203519.59 631219.77 22.9

B01 10 16:35:26 13/09/2009 203515.92 631220.45 22.9

B01 11 16:36:03 13/09/2009 203514.51 631219.17 22.9

SNH

Area B

Lamlash Bay 2009 habitat mapping survey

Page 126: Mapping of the marine habitats and species of Lamlash Bay, Arran

114

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

D01 1 deck 13/09/2009

D01 2 09:52:58 13/09/2009 206632.60 630573.26 46.4

D01 3 09:54:19 13/09/2009 206630.17 630562.63 45.7

D01 4 09:55:17 13/09/2009 206628.19 630555.01 45.2

D01 5 09:55:57 13/09/2009 206626.88 630549.55 45.0

D01 6 09:56:51 13/09/2009 206624.61 630541.54 / Bad Sounding.

D01 7 09:58:08 13/09/2009 206620.72 630530.16 45.1

D01 8 09:59:23 13/09/2009 206617.06 630519.54 45.3

D01 9 10:01:19 13/09/2009 206612.16 630504.35 44.7

D01 10 10:02:09 13/09/2009 206610.73 630498.29 44.5

D01 11 10:04:01 13/09/2009 206606.55 630485.02 43.6

D03 1 deck 13/09/2009

D03 2 10:24:43 13/09/2009 206312.80 630961.02 12.7

D03 3 10:25:44 13/09/2009 206310.83 630950.19 12.4

D03 4 10:26:56 13/09/2009 206307.28 630941.07 11.9

D03 5 10:28:17 13/09/2009 206302.75 630930.40 / Bad Sounding.

D03 6 10:29:09 13/09/2009 206299.30 630923.80 10.6

D03 7 10:29:58 13/09/2009 206297.04 630918.21 10.4

D03 8 10:30:46 13/09/2009 206295.83 630913.67 10.3

D03 9 10:31:36 13/09/2009 206294.26 630908.69 10.2

D03 10 10:32:39 13/09/2009 206291.07 630901.41 10.1

D03 11 10:33:37 13/09/2009 206288.20 630894.60 10.0

D03 12 10:34:40 13/09/2009 206285.38 630887.02 9.9

D03 13 10:35:13 13/09/2009 206284.20 630883.22 9.8

D03 14 10:36:00 13/09/2009 206283.26 630878.28 9.8

D03 15 10:37:51 13/09/2009 206278.09 630863.92 9.7

SNH

Area D

Lamlash Bay 2009 habitat mapping survey

Page 127: Mapping of the marine habitats and species of Lamlash Bay, Arran

115

D02 1 deck 13/09/2009

D02 2 15:51:26 13/09/2009 206563.78 631661.66 48.5

D02 3 15:52:47 13/09/2009 206558.79 631655.01 48.4

D02 4 15:53:23 13/09/2009 206555.94 631652.62 48.3

D02 5 15:55:09 13/09/2009 206550.01 631645.12 48.1

D02 6 15:57:48 13/09/2009 206539.54 631632.75 47.1

D02 7 15:58:32 13/09/2009 Nav frozen

D02 8 15:58:32 13/09/2009 Nav frozen

D02 9 15:58:32 13/09/2009 Nav frozen

Page 128: Mapping of the marine habitats and species of Lamlash Bay, Arran

116

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

E01 1 deck 13/09/2009

E01 2 08:32:34 13/09/2009 206493.03 628168.64 11.6

E01 3 08:33:15 13/09/2009 206496.89 628162.51 11.0

E01 4 08:33:49 13/09/2009 206500.25 628157.08 11.6

E01 5 08:34:28 13/09/2009 206503.95 628151.02 10.9

E01 6 08:35:31 13/09/2009 206509.69 628138.62 10.8

E01 7 08:37:03 13/09/2009 206516.55 628118.49 10.7

E01 8 08:37:03 13/09/2009 206516.55 628118.49 10.7

E01 9 08:38:29 13/09/2009 206521.08 628096.39 11.0

E01 10 08:39:15 13/09/2009 206523.89 628084.99 10.7

E01 11 08:40:20 13/09/2009 206527.29 628070.00 10.9

E01 12 08:41:29 13/09/2009 206528.62 628048.68 11.3

E01 13 08:42:18 13/09/2009 206530.72 628035.52 11.6

E01 14 08:43:08 13/09/2009 206531.82 628020.35 12.0

E01 15 08:44:08 13/09/2009 206534.35 628003.78 12.2

E01 16 08:44:29 13/09/2009 206534.66 627996.51 12.1

E01 17 08:44:45 13/09/2009 206534.95 627990.74 12.4

E01 18 08:46:13 13/09/2009 206536.43 627963.95 12.8

E01 19 08:46:44 13/09/2009 206537.16 627954.19 12.9

E01 20 08:47:26 13/09/2009 206538.63 627941.94 13.1

E01 21 08:47:50 13/09/2009 206539.43 627935.32 13.2

E01 22 08:48:15 13/09/2009 206540.45 627928.20 13.1

E01 23 08:48:43 13/09/2009 206540.94 627920.36 12.7

E01 24 08:49:23 13/09/2009 206542.47 627911.39 12.7

E01 25 08:49:50 13/09/2009 206541.98 627901.69 12.1

E01 26 08:50:30 13/09/2009 206542.33 627889.51 12.4

E01 27 08:51:35 13/09/2009 206543.83 627871.47 13.0

SNH

Area E

Lamlash Bay 2009 habitat mapping survey

Page 129: Mapping of the marine habitats and species of Lamlash Bay, Arran

117

E02 1 deck 13/09/2009

E02 2 09:12:17 13/09/2009 207038.31 628579.24 14.6

E02 3 09:13:02 13/09/2009 207033.93 628562.53 15.2

E02 4 09:13:15 13/09/2009 207032.71 628557.68 15.4

E02 5 09:14:29 13/09/2009 207025.68 628532.07 15.4

E02 6 09:15:15 13/09/2009 207021.33 628516.73 15.4

E02 7 09:15:49 13/09/2009 207017.38 628505.97 15.4

E02 8 09:16:57 13/09/2009 207008.74 628484.95 15.3

E02 9 09:17:32 13/09/2009 207004.08 628474.89 15.4

E02 10 09:18:07 13/09/2009 206999.07 628464.92 15.3

E02 11 09:18:47 13/09/2009 206993.04 628453.00 14.9

E02 12 09:19:46 13/09/2009 206983.41 628436.91 14.7

E02 13 09:20:27 13/09/2009 206977.71 628425.24 15.1

E02 14 09:21:51 13/09/2009 206968.81 628398.23 16.4

E02 15 09:22:30 13/09/2009 206965.00 628385.79 17.1

Page 130: Mapping of the marine habitats and species of Lamlash Bay, Arran

118

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

F01 1 deck 13/09/2009

F01 2 07:50:55 13/09/2009 205117.35 630526.26 3.7

F01 3 07:51:48 13/09/2009 205114.36 630520.48 4.0

F01 4 07:52:43 13/09/2009 205110.91 630515.79 4.1

F01 5 07:53:23 13/09/2009 205107.21 630513.28 4.2

F01 6 07:53:48 13/09/2009 205104.73 630511.63 4.3

F01 7 07:54:46 13/09/2009 205098.72 630507.25 4.7

F01 8 07:56:06 13/09/2009 205089.54 630501.54 6.1

F01 9 07:57:01 13/09/2009 205083.92 630495.58 7.3

F01 10 07:58:02 13/09/2009 205078.54 630489.83 8.6

F01 11 07:58:58 13/09/2009 205074.39 630483.85 9.9

F01 12 08:00:04 13/09/2009 205068.37 630476.04 11.4

F01 13 08:00:38 13/09/2009 205066.49 630472.67 12.4

F01 14 08:01:13 13/09/2009 205064.96 630463.29 13.4

F01 15 08:02:01 13/09/2009 205061.26 630458.10 14.6

F01 16 08:02:55 13/09/2009 205057.30 630452.29 15.4

F01 17 08:03:44 13/09/2009 205053.35 630446.85 17.5

F01 18 08:05:15 13/09/2009 205045.79 630437.61 19.8

SNH

Area F

Lamlash Bay 2009 habitat mapping survey

Page 131: Mapping of the marine habitats and species of Lamlash Bay, Arran

119

Appendix 03: Lamlash Bay still photography log - 2009SeaStar Survey - Project J/09/182

Positions are OSGB36 East and North (m) E/S draft:0.9m

Client Job Number J/09/182

Location Vessel SV Otarie

Survey Date September 2009

Site Photograph Time Date East North Corrected Depth Comments

Name Number (GMT) (m) (m) (m)

H01 1 deck 12/09/2009

H01 2 15:33:10 12/09/2009 205402.51 629549.17 37.0

H01 3 15:34:17 12/09/2009 205399.05 629560.75 36.5

H01 4 15:34:50 12/09/2009 205398.11 629566.98 36.3

H01 5 15:35:32 12/09/2009 205396.22 629575.21 35.9

H01 6 15:36:23 12/09/2009 205393.92 629585.42 35.3

H01 7 15:37:23 12/09/2009 205394.70 629594.82 34.1

H01 8 15:38:28 12/09/2009 205388.53 629610.16 32.5

H01 9 15:39:14 12/09/2009 205386.16 629619.46 30.7

H01 10 15:39:55 12/09/2009 205384.17 629627.89 29.2

H01 11 15:40:26 12/09/2009 205382.56 629633.97 28.7

H01 12 15:40:58 12/09/2009 205380.96 629639.46 28.9

H01 13 15:41:39 12/09/2009 205379.12 629646.81 30.2

H01 14 15:42:50 12/09/2009 205374.14 629661.22 31.5

H01 15 15:43:25 12/09/2009 205371.11 629668.34 32.0

H01 16 15:44:04 12/09/2009 205367.97 629675.91 32.4

H01 17 15:44:34 12/09/2009 205365.41 629681.83 32.6

H01 18 15:45:19 12/09/2009 205361.62 629690.25 32.8

H01 19 15:46:07 12/09/2009 205358.61 629699.22 33.0

H01 20 15:46:59 12/09/2009 205355.08 629709.16 33.2

H01 21 15:48:19 12/09/2009 205348.51 629724.48 33.6

H01 22 15:49:26 12/09/2009 205342.78 629735.70 33.8

H01 23 15:50:17 12/09/2009 205339.97 629745.73 33.9

H01 24 15:51:31 12/09/2009 205335.69 629759.01 33.7

SNH

Area H

Lamlash Bay 2009 habitat mapping survey

Page 132: Mapping of the marine habitats and species of Lamlash Bay, Arran

120

H01 25 15:52:32 12/09/2009 205333.13 629768.51 33.3

H01 26 15:53:40 12/09/2009 205331.38 629780.52 32.6

H01 27 15:54:59 12/09/2009 205328.52 629793.21 31.9

H01 28 15:56:05 12/09/2009 205324.90 629803.88 31.6

H01 29 15:57:14 12/09/2009 205321.38 629814.59 31.4

H01 30 15:58:29 12/09/2009 205317.63 629827.42 30.9

H01 31 15:59:55 12/09/2009 205313.03 629841.33 30.7

H01 32 16:00:44 12/09/2009 205311.39 629849.50 30.3

H01 33 16:01:52 12/09/2009 205309.67 629859.85 29.9

H01 34 16:03:04 12/09/2009 205307.04 629871.54 29.7

H01 35 16:04:34 12/09/2009 205304.05 629886.71 29.1

H01 36 16:05:57 12/09/2009 205301.25 629898.97 28.2

H01 37 16:07:03 12/09/2009 205298.77 629908.79 26.9

H01 38 16:08:21 12/09/2009 205294.14 629921.15 25.2

H01 39 16:09:18 12/09/2009 205291.09 629929.91 23.3

H01 40 16:10:33 12/09/2009 205286.34 629941.20 19.9

Page 133: Mapping of the marine habitats and species of Lamlash Bay, Arran

121

Appendix 04. Particle Size Analysis (PSA)

Appendix 04: Lamlash Bay PSA - 2008SeaStar Survey - Project J/08/151

Particle Size Analysis -

NTZ01#01 NTZ01#01 NTZ02#02 NTZ02#02 NTZ03#01 NTZ03#01 NTZ04#01 NTZ04#01 NTZ05#01 NTZ05#01

Sieve Size Weight Cumul. Weight Cumul. Weight Cumul. Weight Cumul. Weight Cumul.

(mm) % % % % % % % % % %

16.000 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

8.000 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.20 0.20

4.000 0.00 0.00 0.00 0.00 0.02 0.02 0.00 0.00 0.00 0.20

2.000 0.04 0.04 0.03 0.03 0.02 0.04 0.06 0.06 0.07 0.27

1.000 0.38 0.42 0.18 0.21 0.22 0.26 0.19 0.25 0.57 0.84

0.500 1.55 1.97 0.65 0.86 0.75 1.01 0.95 1.20 1.32 2.15

0.250 7.08 9.05 3.49 4.35 3.55 4.56 4.28 5.48 5.77 7.92

0.125 53.04 62.09 37.14 41.49 26.68 31.24 16.85 22.33 41.50 49.43

0.063 32.71 94.80 48.78 90.27 57.66 88.89 65.34 87.67 41.36 90.78

< 0.063 5.44 100.24 10.00 100.28 11.29 100.18 12.10 99.77 9.35 100.13

Process Differential -0.24 -0.28 -0.18 0.23 -0.13

% Gravel 0.04 0.03 0.04 0.06 0.27

% Sand 94.76 90.24 88.85 87.61 90.52

% Silt 5.44 10.00 11.29 12.10 9.35

% Total 100.24 100.28 100.18 99.77 100.13

Dry Weight (g) 128.42 140.63 174.29 102.59 89.73

NTZ06#03 NTZ06#03 NTZ07#01 NTZ07#01 NTZ08#02 NTZ08#02 NTZ09#01 NTZ09#01 NTZ10#01 NTZ10#01

Sieve Size Weight Cumul. Weight Cumul. Weight Cumul. Weight Cumul. Weight Cumul.

(mm) % % % % % % % % % %

16.000 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00 0.00

8.000 0.00 0.00 0.29 0.29 5.13 5.13 1.49 1.49 4.39 4.39

4.000 0.00 0.00 1.07 1.36 3.45 8.58 0.00 1.49 3.92 8.31

2.000 0.14 0.14 3.97 5.33 7.20 15.77 0.00 1.49 3.63 11.93

1.000 0.72 0.86 9.15 14.48 7.36 23.14 0.09 1.57 4.96 16.90

0.500 2.31 3.16 6.77 21.25 5.84 28.98 1.28 2.86 6.19 23.09

0.250 9.72 12.88 19.06 40.31 13.03 42.01 7.28 10.14 17.79 40.88

0.125 50.42 63.30 42.63 82.94 38.11 80.12 38.91 49.05 39.31 80.19

0.063 30.62 93.92 10.84 93.78 12.86 92.98 42.12 91.17 13.60 93.79

< 0.063 6.17 100.09 6.36 100.14 7.09 100.06 8.86 100.03 6.32 100.12

Process Differential -0.09 -0.14 -0.06 -0.03 -0.12

% Gravel 0.14 5.33 15.77 1.49 11.93

% Sand 93.78 88.45 77.20 89.68 81.86

% Silt 6.17 6.36 7.09 8.86 6.32

% Total 100.09 100.14 100.06 100.03 100.12

Dry Weight (g) 121.03 147.03 126.99 126.99 152.76

Page 134: Mapping of the marine habitats and species of Lamlash Bay, Arran

122

Appendix 05. Modified FOLK sediment classification (see Connor et al., 2004)

Page 135: Mapping of the marine habitats and species of Lamlash Bay, Arran

123

Appendix 06. Lamlash Bay Day grab sample species list

Appendix 06: Lamlash Bay Day Grab Sample Analysis - 2008SeaStar Survey - Project J/08/151

Sediment Sample Anaylsis Results

MCSA MCSN Taxon Species Authority Qualifier NTZ 01#01 NTZ 02#02 NTZ 03#01 NTZ 04#01 NTZ 05#01 NTZ 06#03 NTZ 07#01 NTZ 08#02 NTZ 09#01 NTZ 10#01

D 583 ANTHOZOA spp. 1 2

D 632 Cerianthus llodyii Gosse, 1859 1 1 2

D 766 Edwardsia claparedii (Panceri, 1869) 1 2 1

G 1 NEMERTEA spp. indet. 2 3 3 2 1 2 5 5

G 39 Cerebratulus sp. Renier, 1804 2 2

G 26 Tubulanus sp. Renier, 1804 1

N 1 SIPUNCULA sp. 1

N 13 Golfingia sp. Lankester, 1885 1

N 14 Golfingia elongata (Keferstein, 1862)

N 17 Golfingia vulgaris (de Blainville, 1827)

O 1 ECHIURA sp. indet.

P 16 Aphroditoidea sp.

P 19 Aphrodita aculeata Linnaeus, 1758 2

P 25 Polynoidae sp. juv. 1

P 49 Gattyana cirrosa (Pallas, 1766)

P 50 Harmothoe spp. indet. 1 3 4 2

P 50 Harmothoe impar Kinberg, 1855

P 64 Harmothoe cf. imbricata (Linnaeus, 1767)

P 66 Harmothoe ljungmani (Malmgren, 1867)

P 68 Malmgreniella arenicolae (Saint-Joseph, 1888)

P 78 Lepidasthenia argus Hodgson, 1900

P 82 Lepidonotus squamatus (Linnaeus, 1758)

P 91 Pholoe sp. Johnston, 1839

P 91 Pholoe assimilis Oersted, 1845

P 91 Pholoe baltica Oersted, 1843

P 92 Pholoe inornata Johnston, 1839

P 94 Pholoe synophtalmica Claparède, 1868 20 6 11 2 6 2 2 4 10 3

P 107 Sthenelais boa (Johnston, 1839)

P 109 Sthenelais limicola (Ehlers, 1864) 2 1 1 2

P 114 Phyllodocidae sp. 3 1

P 116 Eteone sp. Savigny, 1820 1

P 118 Eteone longa (Fabricius, 1780) agg 1 3 1 1

P 127 Mysta picta (Quatrefages, 1866)

P 142 Anaitides lineata (Claparède, 1870) 1

P 144 Anaitides maculata (Linnaeus, 1767)

P 146 Anaitides rosea (McIntosh, 1877) 1 2 2

P 163 Eumida sp. juv.

P 164 Eumida cf. bahusiensis Bergstrom, 1914

P 167 Eumida sanguinea (Oersted, 1843) 1 1

P 191 Sige fusigera Malmgren, 1865

P 253 Glyceroidea sp.

P 255 Glycera sp. Savigny, 1818 indet. 1 1 2 1

P 256 Glycera alba (O F Müller, 1776)

P 257 Glycera celtica O'Connor, 1987

P 258 Glycera dayi O�Connor, 1987 1

P 260 Glycera lapidum Quatrefages, 1866 agg. 3 1

P 268 Glycinde nordmanni (Malmgren, 1866) 1

P 271 Goniada maculata Oersted, 1843 2 2 1 1

P 293 Hesionidae sp. indet. 1 1

P 305 Psamathe fusca (Keferstein, 1862) Syn. Kefersteinia cirrata

P 310 Nereimyra punctata (O F Müller, 1788) 3

P 312 Ophiodromus flexuosus (Chiaje, 1827) 1

Page 136: Mapping of the marine habitats and species of Lamlash Bay, Arran

124

P 319 Podarkeopsis capensis (Day, 1963) Syn. G. helgolandicus

P 349 Ehlersia cornuta (Rathke, 1843) Syn. Syllis 1

P 368 Typosyllis hyalina (Grube, 1863) Syn. Syllis 1 1

P 371 Typosyllis cf. variegata (Grube, 1860) Syn. Syllis

P 388 Odontosyllis gibba Claparède, 1863

P 423 Exogone verugera (Claparède, 1868) 2 1

P 430 Sphaerosyllis taylori Perkins, 1980 1

P 434 Autolytus sp. Grube, 1850 1

P 473 Nereis sp.

P 475 Nereis longissima Johnston, 1840

P 484 Platynereis dumerilii (Audouin & Milne-Edwards, 1833) 1

P 494 Nephtys spp. juv. 3 1 2 5 10 3 3 2 6

P 496 Nephtys caeca (Fabricius, 1780)

P 498 Nephtys cirrosa Ehlers, 1868 2 2 2 1 5 2

P 499 Nephtys hombergii Savigny, 1818 7 3 6 4 3 3 8 1

P 502 Nephtys kersivalensis McIntosh, 1908 1 2 11 2 3 1 4 1 3

P 539 Aponuphis bilineata (Baird, 1870) Syn. Hyalinocea bilineata 4 3

P 558 Eunice pennata (O F Müller, 1776) 1

P 564 Marphysa bellii (Audouin & Milne-Edwards, 1833)

P 566 Marphysa sanguinea (Montagu, 1813)

P 568 Nematonereis unicornis (Grube, 1840) 1

P 572 Lumbrineris sp. de Blainville, 1828 indet. 1

P 577 Scoletoma fragilis (O F Müller, 1776) Syn. Lumbrinereis fragilis 1

P 579 Lumbrineris gracilis (Ehlers, 1868) 1 1 2 2 7 7 10

P 597 Notocirrus scoticus McIntosh, 1869 1

P 643 Schistomeringos rudolphi (Chiaje, 1828)

P 672 Scoloplos armiger (O F Müller, 1776)

P 699 Paradoneis lyra (Southern, 1914) 1 12 2 1

P 708 SPIONOIDEA sp. indet./juv. 1 2

P 718 Poecilochaetus serpens Allen, 1904

P 722 Aonides oxycephala (M Sars, 1862) 3 6 1

P 723 Aonides paucibranchiata Southern, 1914

P 731 Laonice sp. Malmgren, 1867

P 733 Laonice bahusiensis Soderstrom, 1920 1

P 737 Malacoceros fuliginosus (Claparède, 1868)

P 738 Malacoceros tetracerus (Schmarda, 1861) 1 1 1

P 747 Minuspio cirrifera (Wiren, 1883) Syn. Prionospio cirrifera 3

P 754 Dipolydora flava (Claparède, 1870) Syn. Polydora flava 1 1

P 763 Prionospio sp. Malmgren, 1867 1 1 1 1

P 765 Prionospio fallax Soderstrom, 1920 25 3 19 18 31 35 5 4 27 5

P 772 Pseudopolydora antennata (Claparède, 1870) 1 2

P 774 Pseudopolydora pulchra (Carazzi, 1895)

P 776 Pygospio elegans Claparède, 1863 1

P 790 Spio decorata Bobretzky, 1870 1 3 3 2

P 794 Spiophanes bombyx (Claparède, 1870) 1 1 1 1 1

P 796 Spiophanes kroyeri Grube, 1860 2 1 1 1 2

P 803 Magelona sp. F Müller, 1858 indet. 1

P 804 Magelona alleni Wilson, 1958 1 1 1

P 822 Cirratulidae sp. indet. 1 1

P 823 Aphelochaeta sp. Blake, 1991 1 1

P 829 Caulleriella alata (Southern, 1914)

P 830 Caulleriella bioculata/viridis

P 833 Chaetozone gibber Woodham & Chambers, 1994 1 2 1 1 3

P 834 Chaetozone setosa Malmgren, 1867 1 4

P 845 Tharyx spp. indet. 1 1

P 846 Tharyx killariensis (Southern, 1914) 3 2 1

P 878 Diplocirrus glaucus (Malmgren, 1867) 2 1 2 2 1

P 902 Capitellidae sp.

P 903 Capitellidae sp. A

P 917 Heteromastus filiformis (Claparède, 1864)

Page 137: Mapping of the marine habitats and species of Lamlash Bay, Arran

125

P 919 Mediomastus fragilis Rasmussen, 1973 1 2 3 1 10 10 3

P 921 Notomastus latericeus M Sars, 1851 2 1 2 1 1

P 938 Maldanidae spp. indet. 1 2

P 955 Clymenura tricirrata Bellan & Reys, 1967

P 960 Euclymene sp. Verrill, 1900

P 963 Euclymene lumbricoides (Quatrefages, 1866) 1 1 1

P 964 Euclymene oerstedii (Claparède, 1863) 1 2 1 3 2 4 3

P 971 Praxillella affinis (M Sars, 1872) 5 3 1 1

P 973 Praxillella praetermissa (Malmgren, 1865) 1

P 999 Ophelia borealis Quatrefages, 1866

P 1014 Ophelina acuminata Oersted, 1843 1 3

P 1026 Scalibregma celticum Mackie, 1991 1

P 1027 Scalibregma inflatum Rathke, 1843 1 6 3

P 1093 Galathowenia oculata Zaks, 1922 1

P - Galathowenia fragilis Nilsen & Holthe, 1985 1

P 1098 Owenia fusiformis Chiaje, 1842 1 2 2 1 15 8 6 2 14 15

P 1099 TEREBELLIDA sp. indet./juv. 1 2 1

P 1102 Amphictene auricoma (O F Müller, 1776) 2 3 1 2 3 1 1

P 1107 Lagis koreni (Malmgren, 1866) 1

P 1117 Sabellaria spinulosa Leuckart, 1849

P 1117 Ampharetidae sp. see Holte, 1975

P 1124 Melinna palmata Grube, 1869 2 2 3 5 5 4 4

P 1125 Ampharetinae spp. indet.

P 1139 Ampharete lindstroemi Hessle, 1917 1 16 4 1

P 1171 Trichobranchidae sp.

P 1176 Trichobranchus sp. Malmgren, 1866 2 1 1

P 1177 Trichobranchus glacialis Malmgren, 1866 1

P 1178 Trichobranchus roseus (Malm, 1874) 5 1 2 8 1 2 1 2 6

P 1175 Terebellides stroemi M Sars, 1835 1 1 1

P 1180 Amphitritinae sp. indet. 1

P 1182 Amphitrite cirrata O.F. Muller, 1771 3 1 1 2

P 1189 Eupolymnia nebulosa (Montagu, 1819)

P 1195 Lanice conchilega (Pallas, 1766)

P 1209 Nicolea sp. Malmgren, 1866 juv. 1

P 1218 Pista lornensis (Pearson, 1969) Syn. Scionella lornensis. 1 1

P 1243 Polycirrus cf. norvegicus Wollebaek, 1912

P 1254 Thelepus cincinnatus (Fabricius, 1780) 1

P 1280 Euchone rubrocincta (M Sars, 1861) 1

P 1272 Demonax brachychona (Claparède, 1870) 1

P 1290 Jasmineira elegans Saint-Joseph, 1894

P 1307 Perkinsiana rubra (Langerhans, 1880)

P 1316 Pseudopotamilla reniformis (Bruguiere, 1789)

P 1340 Pomatoceros lamarcki (Quatrefages, 1866) 1

P 1341 Pomatoceros triqueter (Linnaeus, 1758) 1

R 2412 OSTRACODA sp. indet. 2

S 18 Meiosquilla desmaresti Manning, 1977

S 97 AMPHIPODA sp.

S 129 Monoculodes tuberculatus Boeck, 1871 1

S 131 Perioculodes longimanus (Bate & Westwood, 1868) 1

S 137 Synchelidium haplocheles (Grube, 1864) 1

S 140 Westwoodilla caecula (Bate, 1856) 1 1 4

S 177 Leucothoe incisa Robertson, 1892 1 2

S 246 Urothoe sp. Dana, 1852 indet. 1

S 248 Urothoe elegans (Bate, 1856) 4 1 4 3 3 2 5 1

S 249 Urothoe marina (Bate, 1857)

S 253 Harpinia antennaria Meinert, 1890 5 1 1 1

S 271 Lysianassidae sp.

S 275 Acidostoma obesum (Bate & Westwood, 1861) 1 1

S 305 Lysianassa plumosa Boeck, 1871 2 2

S 321 Orchomene nanus (Kröyer, 1846) 3

Page 138: Mapping of the marine habitats and species of Lamlash Bay, Arran

126

S 413 Atylus vedlomensis (Bate & Westwood, 1862) 1 4 1

S 423 Ampelisca sp. Kröyer, 1842 juv./indet. 1 1 2 1 1 1

S 427 Ampelisca brevicornis (Costa, 1853) 2 2 4 4 2

S 429 Ampelisca diadema (A Costa, 1853) 2

S 438 Ampelisca spinipes Boeck, 1861

S 440 Ampelisca tenuicornis Liljeborg, 1855 2 1 4 3 1 2 8 6 1

S 442 Ampelisca typica (Bate, 1856) 1 1 6 2 1

S 498 Abludomelita obtusata (Montagu, 1813) Syn. Melita obtusata 3 1 1

S 503 Cheirocratus sp. Norman, 1867

S 552 Photis longicaudata (Bate & Westwood, 1862)

S 558 Ischyroceridae sp. indet.

S 564 Ericthonius punctatus (Bate, 1857)

S 577 Aoridae sp. 1

S 621 Unciola cf. crenatipalma (Bate, 1862)

S 790 ISOPODA sp. indet.

S 792 Gnathiidae sp. Pranizae 1

S 796 Gnathia oxyuraea (Liljeborg) 1

S 951 Arcturella dilatata (G O Sars, 1882) 1

S 1142 Tanaopsis graciloides (Liljeborg, 1864) 1 4 4 3 4 6 2

S 1248 Diastylis bradyi Norman, 1879 2 1 1

S 1385 Crangon crangon (Linnaeus, 1758) 1

S 1415 Callianassa subterranea (Montagu, 1808)

S 1419 Upogebia deltaura (Leach, 1815)

S 1421 Upogebia stellata (Montagu, 1808)

S 1445 Paguridae sp. juv./indet. 1 2 1 1

S 1448 Anapagurus hyndmanni (Bell, 1845)

S 1460 Pagurus cuanensis Bell, 1845 1

S 1462 Pagurus prideaux Leach, 1815

S 1463 Pagurus pubescens Kröyer, 1838 1

S 1470 Galathea sp. Fabricius, 1793 juv. 5

S 1472 Galathea intermedia Liljeborg, 1851 1

S 1482 Pisidia longicornis (Linnaeus, 1767)

S 1508 Ebalia tuberosa (Pennant, 1777)

S 1580 Liocarcinus depurator (Linnaeus, 1758)

S 1584 Liocarcinus pusillus (Leach, 1815)

S 1636 Pinnotheres sp. indet.

W 9 Chaetoderma nitidulum Lovén, 1844 1 1

W 53 Leptochiton asellus (Gmelin, 1791) 1 2

W 439 Crepidula fornicata (Linnaeus, 1758)

W 459 Trivia arctica (Pulteney, 1799)

W 745 Hinia reticulata (Linnaeus, 1758)

W 747 Hinia incrassata (Ström, 1768)

W 1569 Nucula nitidosa Winckworth, 1930 7 8 6 2 2 1 2

W 1570 Nucula nucleus (Linnaeus, 1758) 6

W 1577 Nuculoma tenuis (Montagu, 1808) 1 1 1 3

W 1773 Aequipecten opercularis (Linnaeus, 1758) 1 1

W 1837 Thyasira flexuosa (Montagu, 1803) 12 13 14 9 16 17 5 4 20 1

W 1865 Diplodonta rotundata (Marshall, 1895)

W 1875 Kellia suborbicularis (Montagu, 1803) 3 4 5 2 2 1

W 1902 Tellimya ferruginosa (Montagu, 1808) 2 1

W 1906 Mysella bidentata (Montagu, 1803) 261 192 212 125 38 23 1 86 2

W 1937 CARDIACEA sp.

W 1952 Parvicardium scabrum (Philippi, 1844)

W 1975 Spisula elliptica (Brown, 1827)

W 1977 Spisula solida (Linnaeus, 1758)

W 1943 Acanthocardia echinata (Linnaeus, 1758) 1 1

W 2001 Ensis siliqua (Linnaeus, 1758)

W 2006 Phaxas pellucidus (Pennant, 1777) 1 1 1

W 2007 TELLINACEA sp. indet. 1

W 2021 Moerella donacina (Linnaeus, 1758) 1 1 1

Page 139: Mapping of the marine habitats and species of Lamlash Bay, Arran

127

W 2231 Thracia phaseolina (Lamarck, 1818) 3 6 1 4 1 2

W 2239 Cochlodesma praetenue (Pulteney, 1799) 3 2

W 2058 Abra sp. Lamarck, 1818 juv. 2

W 2059 Abra alba (W Wood, 1802) 2 1 2 3 4 1

W 2062 Abra prismatica (Montagu, 1808)

W 2095 Gouldia minima (Montagu, 1803) 5 1

W 2098 Chamelea striatula (Linnaeus, 1758) Syn. Venus striatula 2 2 1 1 1 3 2

W 2104 Timoclea ovata (Pennant, 1777) 1 2

W 2113 Tapes rhomboides (Pennant, 1777)

W 2128 Dosinia lupinus (Linnaeus, 1758) 6 8 4 1 2 5 5 11

W 2157 Corbula gibba (Olivi, 1792)

ZA 3 Phoronis spp. 5 3

ZB 100 Asterias rubens Linnaeus, 1758

ZB 121 Ophiuroidea sp. juv. / damaged 7 6 2 2 1 2 2 3 4

ZB 154 Amphiura filiformis (O F Müller, 1776) 64 59 98 59 27 18 2 1 24 18

ZB 161 Amphipholis squamata (Chiaje, 1829)

ZB 165 Ophiuridae sp. 1

ZB 167 Ophiocten affinis (Lütken, 1858) 1 5 1 1 3 1

ZB 168 Ophiura albida Forbes, 1839

ZB 193 Psammechinus miliaris (P. L. S. Müller, 1771)

ZB 212 Echinocyamus pusillus (O F Müller, 1776) 2 1

ZB 223 Echinocardium cordatum (Pennant, 1777) 4 5 5 1 1 1 1 4

ZD 102 Pelonaia corrugata Forbes & Goodsir, 1841 2

ZG 455 Gobiidae sp. 1

Page 140: Mapping of the marine habitats and species of Lamlash Bay, Arran

128

Appendix 07. Lamlash Bay still photography analysis – species list (2008)

Lamlash Bay Habitat Mapping Survey J/08/151

Photographic analysis 2008

MCS

alpha

MCS

num

Genus Species

ALGAE

ZM 1 RHODOPHYCOTA Red algae

ZM 255 Phytomatolithon calcareum - live

ZM 255 Phytomatolithon calcareum - dead

ZM 443 Plocamium cartilagineum

ZR 1 CHROMOPHYCOTA Brown algae

ZR 341 Alaria esculenta

ZR 351 Laminaria hyperborea

ZR 354 Laminaria saccharina

ZS 1 CHLOROPHYCOTA Green algae

C 1 PORIFERA sp.

C - Encrusting sp.

C 414 Suberites sp.

C 651 Halichondria panicea

C 758 Amphilectus fucorum

D 1 CNIDARIA

D - Hydroid sp. SAT

D - Hydroid sp. LAT

D 409 Abietinaria abietina

D 455 Kirchenpaueria pinnata

D 462 Nemertesia sp.

D 463 Nemertesia antennina

D 466 Nemertesia ramosa

D 583 Anthozoa sp.

D 597 Alcyonium digitatum

D 632 Cerianthus llodyii

D 710 Metridium senile

P 2 POLYCHAETA sp.

P 2 POLYCHAETE sp. (tube)

P 2 Faecal casts (coiled - A, marina like)

P - Faecal mounds

P - Burrow vertical

P - Burrow horizontal

P 811 Chaetopterus tubes

P 1188 ?Eupolymnia burrow

P 1195 Lanice conchilega

P 1257 Sabellidae sp. (tube)

P 1300 Myxicola infundibulum

P 1324 Serpulidae sp.

P 1328 Ditrupa sp.

P 1339 Pomatoceros sp.

P 1343 Serpula vermicularis

P 1360 Salmacina sp.

S 1 CRUSTACEA

S - Red shrimp ?Pandalus montagui?

S 1 Crustacea sp.

S 1276 Decapod sp.

S 1402 Nephrops norvegicus

S 1445 Paguridae (Hermit crab) sp

S 1478 Munida rugosa

S 1525 Inachus sp.

Page 141: Mapping of the marine habitats and species of Lamlash Bay, Arran

129

S 1529 Macropodia sp.

S 1566 Cancer pagurus

S 1577 Liocarcinus sp

S 1580 Liocarcinus depurator

S 1589 Liocarcinus puber

W 1 MOLLUSCA

W 88 Gastropoda sp.

W 270 Turritella communis

W 708 Buccinum undatum

W 1560 Bivalvia (PELECYPODA) sp.

W 1768 Pectinidae sp.

W 1771 Pecten maximus

W 1773 Aequipecten opercularis

Y 1 BRYOZOA

Y 187 Flustra foliacea

Y - Membranipora membranacea

ZB - ECHINODERMATA

ZB 1 Crinoidea sp.

ZB 10 Antedon bifida

ZB 15 Leptometra celtica

ZB 18 Asteroidea sp.

ZB 22 Luidia ciliaris

ZB 26 Astropecten irregularis

ZB 54 Porania pulvillus

ZB 74 Crossaster papposus

ZB 100 Asterias rubens

ZB 104 Marthasterias glacialis

ZB 105 Ophiuridae sp.

ZB 128 Ophiocomina nigra

ZB 149 Amphiura sp.

ZB 166 Ophiura sp.

ZB 170 Ophiura ophiura

ZB 181 Echinoidea sp.

ZB 198 Echinus esculentus

ZB 229 Holothuroidea sp.

ZB 82? Henricia sp.

ZD 1 TUNICATA

ZD 2 ASCIDIACEA sp.

ZD 20 Polyclinidae sp.

ZD 71 Ciona intestinalis

ZD 83 Ascidiella sp.

ZD 89 Ascidia mentula

ZF - EUCHORDATA

ZF 29 Scyliorhinus stellaris

ZG 7 TELEOSTEI (Fish indet.) sp.

ZG 105 Gadidae sp.

ZG 121 Melanogrammus aeglefinus

ZG 245 Syngnathus acus

ZG 262 Aspitrigla cuculus

ZG 282 Taurulus sp.

ZG 386 Labridae (Wrasse indet.) sp.

ZG 397 Ctenolabrus rupestris

ZG 398 Labrus bimaculatus

ZG 452 Callionymus lyra

ZG 455 Gobiidae sp.

ZG 479 Pomatoschistus minutus

ZG 545 PLEURONECTIFORMES sp.

ZG 564 Pleuronectidae sp.

Page 142: Mapping of the marine habitats and species of Lamlash Bay, Arran

130

Appendix 07. Lamlash Bay still photography analysis – species list (2009)

Lamlash Bay Habitat Mapping Survey J/09/182

Photographic analysis 2009

MCS

alpha

MCS

num

Genus Species

ALGAEZM 1 RHODOPHYCOTA Red algae

ZM 1 Encrusting pink algae

ZM 170 Palmaria palmata

ZM 255 Phytomatolithon calcareum

ZM 255 Phytomatolithon calcareum

ZM 328 Dilsea carnosa

ZM 344 Chondrus crispus

ZM 417 Polyides rotundus

ZM 443 Plocamium cartilagineum

ZM 616 Phycodrys rubens

ZR 1 CHROMOPHYCOTA Brown algae

ZR 140 Dictyosiphon sp.

ZR 341 Alaria esculenta

ZR 346 Chorda filum

ZR 349 Laminaria sp.

ZR 354 Laminaria saccharina

ZS 1 CHLOROPHYCOTA Green algae

ZS 179 Ulva lactuca

C 1 PORIFERA sp.

D 1 CNIDARIAD Hydroid sp. SAT

D Hydroid sp. LAT

D 463 Nemertesia antennina

D 583 Anthozoa sp.

D 632 Cerianthus llodyii

D 679 Anemonia viridis

D 783 Caryophyll ia smithii

P 2 POLYCHAETA sp.

P 2 POLYCHAETE sp. (tube)

Faecal mounds

Burrow vertical

Burrow horizontal

P 811 Chaetopterus tubes

P 929 Arenicola faecal casts

P 1188 ?Eupolymnia burrow

P 1195 Lanice conchilega

P 1257 Sabellidae sp. (tube)

P 1324 Serpulidae sp.

Page 143: Mapping of the marine habitats and species of Lamlash Bay, Arran

131

CRUSTACEA

R 14 Cirripedia sp.

S 1276 Decapod sp.

S 1445 Paguridae (Hermit crab) sp

S 1478 Munida rugosa

S 1525 Inachus sp.

S 1566 Cancer pagurus

S 1577 Liocarcinus sp

S 1580 Liocarcinus depurator

S 1589 Liocarcinus puber

W 1 MOLLUSCA

W 88 Gastropoda sp.

W 157 Gibbula sp.

W 182 Calliostoma zizyphinum

W 270 Turritella communis

W 1560 Bivalv ia (PELECYPODA) sp.

W 1768 Pectinidae sp.

W 1771 Pecten maximus

W 1773 Aequipecten opercularis

Y 1 BRYOZOA sp.

Y 4 Crisiidae sp.

Y 300 Cellaria fistulosa

Membranipora membranacea

ZB ECHINODERMATA

ZB 10 Antedon bifida

ZB 22 Luidia ciliaris

ZB 54 Porania pulvillus

ZB 82 Henricia sp.

ZB 100 Asterias rubens

ZB 104 Marthasterias glacialis

ZB 128 Ophiocomina nigra

ZB 149 Amphiura sp.

ZB 166 Ophiura sp.

ZB 198 Echinus esculentus

ZD 1 TUNICATAZD 2 ASCIDIACEA sp.

ZD 7 Clavelina lepadiformis

EUCHORDATAZG 262 Aspitrigla cuculus

ZG 440 Pholis gunnellus

ZG 455 Gobiidae sp.

ZG 479 Pomatoschistus minutus

Page 144: Mapping of the marine habitats and species of Lamlash Bay, Arran

132

Appendix 08. Lamlash Bay video analysis – 2008 summary

Line Line S/EOH Time (GMT) Easting (m) Northing (m) Habitat

SOH 16:53:35 206276.21 632392.83 s M sh

EOH 17:15:31 206577.06 632368.17 SS.SMU.CSaMu

SOH 09:55:46 205606.55 631833.85 s m G sh

EOH 10:15:46 205898.05 631907.49 SS.SMU.CSaMu

SOH 10:15:46 205898.05 631907.49 s M g sh

EOH 10:18:46 205968.82 631898.91 SS.SMU.CSaMu

SOH 12:13:51 205578.92 631946.86 s m G sh r and mearl

EOH 12:32:51 205884.56 631972.79 SS.SMPMrl

SOH 12:32:51 205884.56 631972.79 s M g sh

EOH 12:44:03 206071.6 632001.27 SS.SMU.CSaMu

SOH 08:00:30 204384.63 631576.34 s M g sh

EOH 08:04:47 204516.8 631646.02 SS.SMU.CSaMu

SOH 08:04:47 204516.8 631646.02 s m G sh

EOH 08:09:30 204656.8 631728.8 SS.SMx.CMx

SOH 08:09:30 204656.8 631728.8 s M g sh

EOH 08:14:30 204792.72 631794.96 SS.SMU.CSaMu

SOH 08:14:30 204792.72 631794.96 s m G sh

EOH 08:24:30 205042.02 631925.64 SS.SMx.CMx

SOH 08:48:22 204416.68 631496.22 s M g sh

EOH 09:19:22 205073.68 631822.61 SS.SMU.CSaMu

SOH 09:19:22 205073.68 631822.61 s m G sh

EOH 09:35:22 205445.78 632014.3 SS.SMx.CMx

SOH 16:30:24 204265 631083.02 s M g sh

EOH 17:03:30 204415.5 630930.46 SS.SMU.CSaMu

SOH 17:03:30 204415.5 630930.46 s m g R sh

EOH 17:15:24 204503.4 630755.99 CR.LCR.BrAsAmenCio

SOH 17:15:24 204503.4 630755.99 s M g sh

EOH 17:18:33 204496.9 630739.64 SS.SMU.CSaMu

SOH 07:53:58 204413.44 630653.56 s M sh

EOH 08:00:05 204460.33 630794.24 SS.SMU.CSaMu

SOH 08:00:05 204460.33 630794.24 s m G sh

EOH 08:02:38 204476.6 630857.51 SS.SMx.CMx

SOH 08:02:38 204476.6 630857.51 s m g sh R

EOH 08:04:58 204505.47 630916.61 CR.LCR.BrAsAmenCio.Bri

SOH 08:04:58 204505.47 630916.61 s m g R sh

EOH 08:11:28 204577.8 631078.42 CR.LCR.BrAsAmenCio

SOH 08:11:28 204577.8 631078.42 s M g sh

EOH 08:24:05 204799.93 631418.73 SS.SMU.CSaMu

SOH 13:04:00 205764.55 632088.74 s m G sh and Maerl

EOH 13:10:00 205845.06 632061.62 SS.SMPMrl

SOH 13:10:00 205845.06 632061.62 s m G sh

EOH 13:16:00 205943.27 632022.16 SS.SMx.CMx

SOH 13:16:00 205943.27 632022.16 s M sh

EOH 13:28:54 206147.65 631970.63 SS.SMU.CSaMu

SOH 14:03:10 204562.86 631290.92 s M sh

EOH 14:22:10 204833.53 631325.27 SS.SMU.CSaMu

SOH 14:22:10 204833.53 631325.27 s M g sh

EOH 14:32:10 204961.42 631307.79 SS.SMU.CSaMu

SOH 14:32:10 204961.42 631307.79 s m G sh

EOH 14:43:38 205045.67 631256.91 SS.SMx.CMx

SOH 14:58:24 204397.64 631728.14 s m G sh and Maerl

EOH 15:15:24 204585.96 631655.93 SS.SMPMrl

SOH 15:15:24 204585.96 631655.93 s m G sh

EOH 15:20:24 204640.74 631603.14 SS.SMx.CMx

SOH 15:20:24 204640.74 631603.14 s M sh

EOH 15:38:38 204778.64 631439.91 SS.SMU.CSaMu

SOH 08:45:02 204508.27 630344.1 s M sh

EOH 08:54:56 204617.94 630702.43 SS.SMU.CSaMu

Biotope 1

Biotope 1

Biotope 2

Biotope 1

Biotope 2

Biotope 1

Biotope 2

Biotope 3

Biotope 4

Biotope 1

Biotope 2

Biotope 1

Biotope 2

Biotope 3

Biotope 1

Biotope 2

Biotope 2

Biotope 3

Biotope 4

Biotope 5

Biotope 1

Biotope 3

Biotope 1

Biotope 2

Biotope 3

Biotope 1

Biotope 3

Biotope 1

Biotope 2

LNC01

LNC02

LNC02

LNC03

LNC03

LNC04

LNC04

LNC04

LNC04

LNC05

LNC05

LNC07

LNC07

LNC07

LNC08

LNC08

LNC08

LNC08

LNC08

LNC09

LNC09

LNC09

LNC10a

LNC10a

LNC10a

LNC11

LNC11

LNC11

LNC12

Page 145: Mapping of the marine habitats and species of Lamlash Bay, Arran

133

SOH 08:54:56 204617.94 630702.43 s m G sh

EOH 08:56:32 204639.24 630768.71 SS.SMx.CMx

SOH 08:56:32 204639.24 630768.71 s M g sh

EOH 09:05:02 204769.41 631114.53 SS.SMU.CSaMu

SOH 09:05:02 204769.41 631114.53 s m G sh

EOH 09:11:22 204908.13 631382.67 SS.SMx.CMx

SOH 09:11:22 204908.13 631382.67 s M g sh

EOH 09:15:46 205022.07 631557.7 SS.SMU.CSaMu

SOH 17:21:34 204879.34 629699.64 s M sh

EOH 17:46:00 205433.04 629456.85 SS.SMU.CSaMu

SOH 17:46:00 205433.04 629456.85 s m G sh

EOH 17:53:31 205581.78 629350.39 SS.SMx.CMx

SOH 17:53:31 205581.78 629350.39 s m g R sh

EOH 17:54:10 205593.97 629340.64 CR.LCR.BrAsAmenCio.Bri

SOH 17:54:10 205593.97 629340.64 s m G sh

EOH 17:56:24 205629.11 629309.78 SS.SMx.CMx

SOH 09:36:59 204057.25 629996.69 s M sh

EOH 10:08:34 204380.7 630978.23 SS.SMU.CSaMu

SOH 10:08:34 204380.7 630978.23 s m G sh

EOH 10:12:39 204454.12 631120.87 SS.SMx.CMx

SOH 10:12:39 204454.12 631120.87 s M sh

EOH 10:15:13 204510.13 631214.82 SS.SMU.CSaMu

SOH 10:07:42 204848.93 628955.36 s M sh

EOH 10:43:42 205408.02 629304.2 SS.SMU.CSaMu

SOH 12:25:46 204689.53 629397.17 s M g sh

EOH 13:02:06 205271.37 629632.79 SS.SMU.CSaMu

SOH 13:02:06 205271.37 629632.79 s m G sh

EOH 13:08:46 205380.59 629704.36 SS.SMx.CMx

SOH 13:23:45 204320.76 629740.41 s M sh

EOH 13:43:15 204727.57 629790.26 SS.SMU.CSaMu

SOH 13:58:48 204621.31 630112.25 s M sh

EOH 14:22:24 205105.02 630168.57 SS.SMU.CSaMu

SOH 14:36:37 203787.86 630394.24 s M sh

EOH 14:50:46 204223.68 630438.64 SS.SMU.CSaMu

SOH 14:50:46 204223.68 630438.64 s m g R sh

EOH 14:53:17 204301.97 630432.9 CR.LCR.BrAsAmenCio

SOH 14:53:17 204301.97 630432.9 s M g sh

EOH 15:07:19 204747.67 630403.34 SS.SMU.CSaMu

SOH 07:17:56 204866.1 634484.42 s m G sh and Maerl

EOH 07:32:56 205147.06 634364.8 SS.SMPMrl

SOH 07:32:56 205147.06 634364.8 s m G sh

EOH 07:37:56 205247.71 634306.06 SS.SMx.CMx

SOH 07:37:56 205247.71 634306.06 s m g R sh

EOH 07:57:36 205554.72 634177.24 CR.LCR.BrAsAmenCio

SOH 07:57:36 205554.72 634177.24 s M g sh

EOH 08:02:44 205642.66 634198.79 SS.SMU.CSaMu

SOH 08:18:30 205692.99 633552.76 s M g sh

EOH 08:31:43 205999.85 633682.51 SS.SMU.CSaMu

SOH 08:45:25 205634.51 Est 633321.83 Est s m G sh and Maerl

EOH 08:48:30 205732.15 Est 633377.15 Est SS.SMPMrl

SOH 08:48:30 205732.15 Est 633377.15 Est s m g R sh

EOH 08:49:55 205777.00 Est 633402.61 Est CR.LCR.BrAsAmenCio.Bri

SOH 08:49:55 205777.00 Est 633402.61 Est s m G sh and Maerl

EOH 08:56:21 205982.08 Est 633517.41 Est SS.SMPMrl

SOH 08:56:21 205982.08 Est 633517.41 Est s M g sh

EOH 08:59:45 206089.70 Est 633577.86 Est SS.SMU.CSaMu

SOH 09:17:16 206038.38 632393.77 s m G sh

EOH 09:22:25 206250.22 632531.04 SS.SMx.CMx

SOH 09:22:25 206250.22 632531.04 s M g sh

EOH 09:25:27 206373.68 632629.04 SS.SMU.CSaMu

Biotope 2

Biotope 3

Biotope 4

Biotope 5

Biotope 1

Biotope 2

Biotope 3

Biotope 1

Biotope 2

Biotope 3

Biotope 1

Biotope 1

Biotope 2

Biotope 1

Biotope 1

Biotope 2

Biotope 1

Biotope 3

Biotope 1

Biotope 2

Biotope 2

Biotope 4

Biotope 2

Biotope 3

Biotope 4

Biotope 1

Biotope 3

Biotope 4

Biotope 1

Biotope 1

LNC12

LNC12

LNC12

LNC12

LSC01

LSC01

LSC01

LSC01

LSC02

LSC02

LSC02

LSC03

LSC04

LSC04

LSC05

LSC06

LSC07

LSC07

LSC07

LOC01

LOC01

LOC01

LOC01

LOC03

LOC04

LOC04

LOC02

LOC03

LOC03

LOC03

Page 146: Mapping of the marine habitats and species of Lamlash Bay, Arran

134

Appendix 08. Lamlash Bay video analysis – 2009 summary

Lamlash Bay Habitat Mapping Survey J/09/182

Video analysis 2009

Line S/EOH Time (GMT) Easting (m) Northing (m) HabitatSOH 13:12:52 205275.91 632650.18 s G

EOH 13:24:15 205441.04 632596.12 SS.SMP.KSwSS.LsacR.CbPb

SOH 13:36:14 205573.6 632384.17 m s G sh

EOH 13:48:49 205722.95 632259.92 SS.SMPMrl.Pcal

SOH 14:18:34 205756.73 631614.32 g S sh

EOH 14:18:48 205759.74 631612.41 SS.SMPMrl.Pcal

SOH 14:18:48 205759.74 631612.41 g R sh

EOH 14:31:01 205837.73 631497.46 SS.SMP.KSwSS.LsacR.CbPb

SOH 14:31:01 205837.73 631497.46 m s G sh

EOH 14:37:20 205892.03 631448.54 SS.SMPMrl.Pcal

SOH 15:25:14 204368.72 631857.45 g S sh

EOH 15:28:54 204421 631831.09 SS.SMP.KSwSS.LsacR.Sa

SOH 15:28:54 204421 631831.09 s G sh

EOH 15:34:21 204486.04 631782.44 SS.SMP.KSwSS.LsacR.CbPb

SOH 15:34:21 204486.04 631782.44 m s G sh

EOH 15:41:59 204544.51 631714.87 SS.SMPMrl.Pcal

SOH 15:41:59 204544.51 631714.87 m s G sh

EOH 15:42:44 204548.56 631707.49 SS.SMPMrl.(Pcal)

SOH 14:50:59 204919.86 631163.64 m S G sh

EOH 14:55:45 204928.87 631108.92 SS.SMx.CMx.ClloMx(Nem)

SOH 14:55:45 204928.87 631108.92 m S sh

EOH 15:06:21 205010.66 631008.3 SS.SSa.CMuSa.(AbraAirr)

SOH 13:43:28 205286.16 632379.48 m s G sh

EOH 14:08:09 205104.44 632447.62 SS.SMPMrl.Pcal

SOH 14:23:20 205890.23 632633.4 m s G sh

EOH 14:30:20 205872.88 632626.09 SS.SMPMrl.Pcal

SOH 14:30:20 205872.88 632626.09 m s G sh

EOH 14:33:00 205859.13 632624.45 SS.SMPMrl.(Pcal)

SOH 14:33:00 205859.13 632624.45 s G sh

EOH 14:35:00 205831.68 632628.93 SS.SMPMrl.Pcal

SOH 14:35:00 205831.68 632628.93 m s G sh

EOH 14:38:18 205822.25 632627.42 SS.SMPMrl.Pcal

SOH 14:46:06 205830.36 632763.66 m s G sh

EOH 14:49:36 205808.17 632764.45 SS.SMPMrl.Pcal

SOH 14:49:36 205808.17 632764.45 m s G sh r

EOH 14:54:59 205772.94 632727.29 SS.SMP.KSwSS.LsacR.CbPb

SOH 15:32:36 205405.94 629542.12 m s G sh

EOH 15:39:12 205386.26 629619.07 SS.SMx.CMx.ClloMx(Nem)

CAM A02 Bio 4

CAM A03 Bio 1

CAM H01 Bio 1

CAM A03 Bio 1

CAM NTZ04 Bio 2

CAM NTZ04 Bio 3

CAM NTZ04 Bio 4

CAM NTZ05 Bio 1

CAM NTZ05 Bio 2

CAM A01

CAM A02 Bio 1

CAM A02 Bio 2

CAM A02 Bio 3

CAM NTZ04 Bio 1

CAM NTZ01

CAM NTZ02

CAM NTZ03 Bio 1

CAM NTZ03 Bio 2

CAM NTZ03 Bio 3

Page 147: Mapping of the marine habitats and species of Lamlash Bay, Arran

135

SOH 15:39:12 205386.26 629619.07 m s G r sh

EOH 15:45:40 205360.25 629694.22 CR.LCR.BrAs.AmenCio

SOH 15:45:40 205360.25 629694.22 m s G sh

EOH 16:11:10 205284.42 629947.14 SS.SMx.CMx.ClloMx(Nem)

SOH 07:50:06 205121.62 630532.47 S g sh

EOH 07:53:36 205103.96 630510.99 SS.SMP.KSwSS.LsacR.CbPb

SOH 07:53:36 205103.96 630510.99 m S g sh

EOH 07:58:36 205076.02 630486.25 SS.SMP.KSwSS.LsacR.Sa

SOH 07:58:36 205076.02 630486.25 m s G sh

EOH 08:05:27 205044.79 630436.39 SS.SMx.CMx.ClloMx(Nem)

SOH 08:31:42 206487.85 628176.72 s G sh

EOH 08:42:22 206530.59 628034.23 SS.SMP.KSwSS.LsacR.CbPb

SOH 08:42:22 206530.59 628034.23 m s G sh

EOH 08:45:02 206534.9 627984.74 SS.SMPMrl.Pcal

SOH 08:45:02 206534.9 627984.74 m s G sh

EOH 08:45:17 206535.23 627980.2 SS.SMPMrl.Pcal

SOH 08:45:17 206535.23 627980.2 m s G sh

EOH 08:47:34 206538.99 627939.68 SS.SMPMrl.Pcal

SOH 08:47:34 206538.99 627939.68 m s G sh

EOH 08:52:54 206540.62 627852.28 SS.SMX.CMx.OphMx

SOH 09:11:42 207041.95 628592.35 m s G sh

EOH 09:15:57 207016.35 628503.45 SS.SMPMrl.Pcal

SOH 09:15:57 207016.35 628503.45 m s G sh

EOH 09:17:32 207004.08 628474.89 SS.SMx.CMx.ClloMx(Nem)

SOH 09:17:32 207004.08 628474.89 m s G sh

EOH 09:21:42 206969.9 628401.34 SS.SMX.CMx.OphMx

SOH 09:21:42 206969.9 628401.34 m s G sh

EOH 09:22:24 206965.58 628387.62 SS.SMx.CMx.ClloMx(Nem)

SOH 09:52:20 206633.76 630578.21 m S

EOH 10:04:14 206605.89 630483.4 SS.SSa.CMuSa.(AbraAirr)

SOH 15:50:32 206566.06 631666.42 m S

EOH 15:59:50 206522.49 631616.79 SS.SSa.CMuSa.(AbraAirr)

SOH 12:24:17 206316.36 630960.11 m s G sh

EOH 10:38:10 206276.79 630861.24 SS.SMPMrl.Pcal

SOH 16:26:32 203540.33 631204.80 m S

EOH 16:36:18 203513.67 631219.10 SS.SSa.CMuSa.(AbraAirr)CAM B01

CAM E02 Bio 2

CAM E02 Bio 3

CAM E02 Bio 4

CAM D01

CAM D02

CAM D03

CAM E01 Bio 2

CAM E01 Bio 3

CAM E01 Bio 4

CAM E01 Bio 5

CAM E02 Bio 1

CAM H01 Bio 2

CAM H01 Bio 3

CAM F01 Bio 1

CAM F01 Bio 2

CAM F01 Bio 3

CAM E01 Bio 1

Page 148: Mapping of the marine habitats and species of Lamlash Bay, Arran

136

Appendix 09. Faunal densities for noteworthy species

Still photography analysis 2008

Munida rugosa

Cancer pagurus

Pectinidae Sp.

Pecten maximus

Aequipecten opercularis

Total 9.4 0.5 1.5 4.4 2.0

Total (N) 80 4 13 37 17

LNC 8.8 0.5 1.2 4.7 2.0

LOC 16.3 0.0 2.0 2.6 0.7

All

LSC 6.6 0.7 1.7 4.9 2.8

NTZ 6.6 0.7 1.8 3.3 2.6

NTZ (N) 18 2 5 9 7

NTZ

NTZ LNC 6.6 0.7 1.8 3.3 2.6

SaMu Total 8.3 0.4 0.4 1.5 1.5

Total (N) 22 1 1 4 4

LNC 7.9 0.0 0.6 1.8 2.4

LOC 22.0 0.0 0.0 0.0 0.0

LSC 0.0 1.7 0.0 1.7 0.0

CMx Total 10.6 0.5 1.0 7.0 3.0

Total (N) 21 1 2 14 6

LNC 5.6 0.8 0.0 4.0 3.2

LOC 24.5 0.0 0.0 4.1 2.0

LSC 7.7 0.0 7.7 26.9 3.8

Mrl.Pcal Total 1.7 0.8 5.0 7.6 0.0

Total (N) 2 1 6 9 0

LNC 1.3 1.3 3.9 10.5 0.0

LOC 2.3 0.0 7.0 2.3 0.0

LSC - - - - -

AmenCio Total 27.5 0.0 1.4 10.1 0.0

AmenCio Total (N) 19 0 1 7 0

LNC 35.7 0.0 2.4 7.1 0.0

LOC 15.8 0.0 0.0 5.3 0.0

LSC 12.5 0.0 0.0 37.5 0.0

MuSa Total 0.0 8.9 0.6 1.7 1.7

Total (N) 0 15 0 3 3

LNC - - - - -

LOC - - - - -

LSC 0.0 8.9 0.6 1.7 1.7

SaMu SS.SMu.CSaMu.(AfilMysAnit) CMx SS.SMx.CMx.ClloMx(Nem) Mrl.Pcal SS.SMP.Mrl.Pcal AmenCio CR.LCR.BrAs.AmenCio AmenCio CR.LCR.BrAs.AmenCio.Bri MuSa SS.SSa.CMuSa.(AbraAirr)

The approximate densities (ind/100 m2) of scallops and two other species present in the still photographs from the 2008 Lamlash Bay survey with total densities and densities for each classified biotope (total: total number of individuals/100m2; total (N): total number of individuals; NTZ: individuals/100m2 in the NTZ area; LNC: total number of individuals/100m2 in northern Lamlash Bay; LOC: total number of individuals/100m2 in Outer Lamlash Bay; LSC: total number of individuals/100m2 in southern and central Lamlash Bay; Pectinidae sp. are unidentified individuals i.e. not the total of all scallops).

Page 149: Mapping of the marine habitats and species of Lamlash Bay, Arran

137

Video footage analysis 2008

Pectinidae sp.

Cancer pagurus

Nephrops norvegicus

Pleuronectidae sp.

Total 2.3 0.4 0.1 0.0

Total (N) 342 61 9 2

LNC 2.7 0.3 0.0 0.0

LOC 2.0 0.7 0.2 0.0

All

LSC 1.9 0.2 0.0 0.0

SaMu Total 2.3 0.5 0.1 0.0

CMx Total 1.8 0.5 0.0 0.0

Mrl.Pcal Total 2.5 0.1 0.0 0.0

AmenCio

AmenCio Total 4.4 0.1 0.0 0

MuSa Total * * * *

SaMu SS.SMu.CSaMu.(AfilMysAnit)

CMx SS.SMx.CMx.ClloMx(Nem)

Mrl.Pcal SS.SMP.Mrl.Pcal

AmenCio CR.LCR.BrAs.AmenCio

AmenCio CR.LCR.BrAs.AmenCio.Bri

MuSa SS.SSa.CMuSa.(AbraAirr)

The approximate densities (ind/100 m2) of scallops and three other species present in the video footage from the 2008 Lamlash Bay survey (with the approximate densities in the different survey areas and biotope; total: total number of individuals/100m2; total (N): total number of individuals; Pectinidae sp. are the total number of scallop individuals as identification of scallops species is difficult on video footage).

Page 150: Mapping of the marine habitats and species of Lamlash Bay, Arran

138

Still photography analysis 2009

Munida rugosa

Cancer pagurus

Pectinidae sp.

Pecten maximus

Aequipecten opercularis

Total 0.0 0.0 0.0 0.0 0.0 All Total (N) 6 1 2 5 1

NTZ 0.0 0.0 0.0 0.0 0.0 NTZ NTZ (N) 0 0 1 2 0

A02 & A03 Total 0.0 0.0 0.0 0.0 0.0

B01 & H01 Total 0.0 0.0 0.0 0.0 0.0

D Total 0.0 0.0 0.0 0.0 0.0

E Total 0.0 0.0 0.0 0.0 0.0

CMx Total 5.8 0.0 1.9 3.8 1.9

Total (N) 3 0 1 2 1

Mrl.Pcal Total 0.0 0.8 0.8 2.3 0.0

Total (N) 0 1 1 3 0

AmenCio Total 0.0 0.0 0.0 0.0 0.0

AmenCio Total (N) 0 0 0 0 0

MuSa Total 8.1 0.0 0.0 0.0 0.0

Total (N) 3 0 0 0 0

OphMx Total 0.0 0.0 0.0 0.0 0.0

Total (N) 0 0 0 0 0

LsacR Total 0.0 0.0 0.0 0.0 0.0

LsacR Total (N) 0 0 0 0 0

AmenCio CR.LCR.BrAs.AmenCio

AmenCio CR.LCR.BrAs.AmenCio.Bri

CMx SS.SMx.CMx.ClloMx(Nem)

OphMx SS.SMX.CMx.OphMx

Mrl.Pcal SS.SMPMrl.Pcal

Mrl.Pcal SS.SMPMrl.(Pcal)

MuSa SS.SSa.CMuSa.(AbraAirr)

LsacR SS.SMP.KSwSS.LsacR.CbPb

LsacR SS.SMP.KSwSS.LsacR.Sa

The approximate densities (ind/100 m2) of scallops and two other species present in the video footage from the 2009 Lamlash Bay survey (with the approximate densities in the different survey areas and biotope; total: total number of individuals/100m2; total (N): total number of individuals; Pectinidae sp. are the total number of scallop individuals as identification of scallops species is difficult on video footage).

Page 151: Mapping of the marine habitats and species of Lamlash Bay, Arran

139

Video footage analysis 2009

Cancer pagurus

Nephrops norvegicus

Munida rugosa

Pectinidae sp.

Pecten maximus

Total 0.3 0.3 3.0 0.1 1.1 All

Total (N) 7 9 82 2 31

Total 0.2 0.0 0.0 0.1 0.5 NTZ

Total (N) 3 0 0 1 6

A02 & A03 Total 0.0 0.0 0.0 0.0 1.4

B01 & H01 Total 0.2 0.0 5.1 0.2 3.1

D Total 1.1 3.3 21.7 0.0 0.4

E Total 0.0 0.0 0.0 0.0 1.5

CMx Total 0.0 0.0 4.4 0.2 2.4

Mrl.Pcal Total 0.6 0.0 0.0 0.0 0.6

AmenCio

AmenCio Total 1.3 0.0 1.3 0.0 6.3

MuSa Total 0.0 2.8 18.2 0.3 0.9

OphMx Total 0.0 0.0 0.0 0.0 2.6

LsacR

LsacR Total 0.0 0.0 0.0 0.0 0.0

AmenCio CR.LCR.BrAs.AmenCio

AmenCio CR.LCR.BrAs.AmenCio.Bri

CMx SS.SMx.CMx.ClloMx(Nem)

OphMx SS.SMX.CMx.OphMx

Mrl.Pcal SS.SMPMrl.Pcal

Mrl.Pcal SS.SMPMrl.(Pcal)

MuSa SS.SSa.CMuSa.(AbraAirr)

LsacR SS.SMP.KSwSS.LsacR.CbPb

LsacR SS.SMP.KSwSS.LsacR.Sa

The approximate densities (ind/100 m2) of scallops and two other species present in the video footage from the 2009 Lamlash Bay survey (with the approximate densities in the different survey areas and biotope; total: total number of individuals/100m2; total (N): total number of individuals; Pectinidae sp. are the total number of scallop individuals as identification of scallops species is difficult on video footage).

Page 152: Mapping of the marine habitats and species of Lamlash Bay, Arran

www.snh.gov.uk© Scottish Natural Heritage 2010ISBN: 978-1-85397-673-5

Policy and Advice Directorate, Great Glen House,Leachkin Road, Inverness IV3 8NWT: 01463 725000

You can download a copy of this publication from the SNH website.