lucanus leaflet
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A color photo-catalogue of all known African species and subspeciesof Stag Beetles. This book treats the whole African continent, includingMadagascar and small islands: 125 species of 16 genera from tropicalAfrica and North Africa are listed and figured. For all species andsubspecies the detailed distribution is given, and for some speciesphotos of habitats and living specimens in the field are included.In addition, portraits and short biographical information aboutentomologists and explorers who worked particularly with the Africanlucanid fauna are also given. The species richness of the variouscountries and the perspectives for further research are discussed.For each African country all the known species are listed, togetherwith the number of endemisms. Author, year, publication, synonyms,size, type locality, abundance, remarks, and distribution are given forevery taxon.The label data of all the illustrated specimens are cited,as well as the acronym of the collection where the specimen is held.Many of the type specimens from the most important entomologicalmuseums (Berlin, Pretoria, Eberswalde, London, Paris, Cape Town,Munich, Dresden, Tervuren, Vienna, Florence, Genova and Basel)are illustrated. A large amount of bibliographical references are givenat the end.
More then 400 color photographs of all the speciesand subspecies from the African continent,as well as photos of living specimens and habitats- 192 pages, luxurious hardcover edition, largeformat 23x31, text in English
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fig. 1 fig. 2
fig. 3 fig. 5fig. 4
Hermann Julius KolbeHenry BoileauEugene SéguyBernard BeneshSebastian Endrödy-YoungaHugues Eli BomansMarc DesfontainePhilippe MorettoJean Michel MaesTetsuo MizukamiShinya Kawai
`
fig. 1 - Hermann Julius Kolbe, fig. 2 - Henry Boileau, fig. 3 - Eugene Séguy, fig. 4 - Bernard Benesh, fig. 5 - Sebastian Endrödy-Younga`
11
fig. 6
fig. 7 fig. 8
fig. 9 fig. 10
fig. 6 - Jean Michel Maes, fig. 7 - Hugues Eli Bomans, fig. 8 - Marc Desfontaine (on left) and Philippe Moretto, fig. 9 - Tetsuo Mizukami,fig. 10 - Shinya Kawai (on right) here together with Robert Lízler (on left) and Karl Werner
22
Oonotus ( Macroonotus ) rex Endrödy-Younga
Publication: ENDRÖDY-YOUNGA, 1993: 34
Type locality: South Africa, Transkei, Port St. Jones
Size: m 19.1, f 15.7
Rarity: very rare
Remarks: only the type specimens are known.
The genus Oonotus has been revised by Endrödy-Younga (1993). R. Müller (pers.
comm.), who was with S. Endrödy-Younga when he collected the holotype, told us
that the specimen was slowly wandering on the bark of a tree in thick forest.
Distribution: South Africa (Transkei)
fig. 17 - m, 19.1 mm, South Africa, Transkei, Port St. Jones, 2.XII.1987, Endrödy-Younga leg., TMP, HOLOTYPUS
fig. 18 - f, 15.7 mm, South Africa, Transkei, Port St. Jones, 2.XII.1987, Endrödy-Younga leg., TMP, ALLOTYPUS
fig. 17 fig. 18
37
Figulus sublaevis (Palisot de Beauvois)
Publication: PALISOT DE BEAUVOIS, 1805: 3 (Lucanus)
Synonyms: anthracinus Klug, 1833: 85
ebenus Westwood, 1834: 119
nigritus Westwood, 1838: 261
monilifer Parry, 1862: 113
vulneratus J. Thomson, 1862: 433
lettowi Kriesche, 1921: 113
nossibenus Kriesche, 1921: 114
Type locality: “Regne Oware”
Size: 11.4 -17.4
Rarity: common
Remarks: according to Maes (1992a: 53) F. decipies is a subspecies of F. sublaevis
inhabiting only Sao Thomé Island, but according to Bomans (1992: 251)
and Monte & Bartolozzi (2004, in press) F. decipies is a valid species.
F. anthracinus seems to be a valid species too, but further studies are required.
On several occasions individuals of Figulus sublaevis were collected at light
by the second author and R. Lizler in open forests of southern and eastern Africa.
Distribution: Cameroon, Democratic Republic of the Congo (ex Zaire),Gabon,
Guinea, Malawi, Mozambique, Madagascar, Senegal, Tanzania, Togo
fig. 42 - 15 mm, Principe Isl., AKC fig. 43 - m, 17 mm, ex Coll. Borel, AKC
fig. 44 - m, 13 mm, Tanzania, Morogoro Prov., Uluguru Mts., 8. - 9.XII.1996, Werner & Lizler leg., KWC
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Figulus
fig. 42 fig. 43 fig. 44
26
Amneidus godefroyi Coquerel
Publication: COQUEREL, 1866: 326
Type locality: Is. Bourbon
Size: m 13.0 -15.0, f 12.0-15.0
Rarity: rare
Remarks: interesting remarks on this species are given by Paulian (1959, 1973).
Distribution: La Réunion Is.
fig. 26 - m, 15 mm, Réunion, I.1996, AKC fig. 27 - f, 15 mm, La Réunion, Kerveguen, 10.I.1962, AKC
fig. 26 fig. 27
53
Nigidius gnu Kriesche
Publication: KRIESCHE, 1921: 109
Type locality: D. Ost Afrika, Pangani, Pugu c/o Daressalam
Size: 13.0 -18.0
Rarity: very rare
Remarks: close to N. bubalus. Bomans redescribed this species
and established the lectotype.
Distribution: Tanzania
fig. 73 - 17.5 mm, D. O. Afrika, Pangani, ZMB, LECTOTYPUS
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Nigidius
fig. 73
82
fig. 119 - Madagascar, Marojejy, March 2001 fig. 120 - Madagascar, Ankarana, March 2001
fig. 121 - Madagascar, “Andasibe (Périnet)”, May 1998
fig. 119
fig. 120 fig. 121
83
fig. 122 - Madagascar, Morondava, April 1998 fig. 123 - Madagascar, Beza Mahafaly, April 1998
fig. 124 - Madagascar, Ranomafana, March 1998 fig. 125 - Madagascar, Ranomafana, March 1998
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Ganelius
fig. 124
fig. 123
fig. 122
fig. 125
84
fig. 126 - m, 25 mm, Ruanda, Ihembe, 11 km E de Dendezi, 2000km, 19.IX.1951, H. RENIER leg, IRSNB, PARATYPUS
fig. 127 - m, 24 mm, Zaire, Kahuzi, km 82, 1-15.IX.1951, legit H. Bomans (in coll. G. Taroni)
Dendezia renieri Basilewsky
Publication: BASILEWSKY, 1952: 37
Type locality: Rwanda, Dendezi
Size: m 23.0 -25.0, f [size not indicated in the description]
Rarity: very rare
Remarks: this species is very peculiar for the presence of a medial cephalic horn.
We are grateful to A. Drumont (IRSNB) for the fine pictures of this rare species.
B. Gill (pers. comm.) found a dead specimen of Dendezia renieri on a dirt road in
the Kibale forest of Uganda in October, 2001.
Distribution: SE Democratic Republic of the Congo (ex Zaire), Rwanda, Uganda
fig. 126
fig. 127
90
fig. 137 - m, 19 mm, Rep. of S. Africa, Cape, Swartberg, IX.1993, Lizler leg., AKC
fig. 137
Colophon berrisfordi Barnard
Publication: BARNARD, 1932: 173
Type locality: South Africa, Zwartberg Range, Meiringspoort Berg
Size: m 19.0 -22.0
Rarity: very rare
Remarks: the genus Colophon has been revised by Endrödy-Younga (1988)
and Mizukami & Kawai (1997). R. Lizler (pers. comm.) collected a single male
of C. berrisfordi among several specimens of C. whitei under stones.
Distribution: South Africa (Swartberg)
92
fig. 140 - m, 17.5 mm, Rep. of S. Africa, Cape, Meiringspoortberg, I.1994, AKC
fig. 141 - f, 16 mm, Rep. of S. Africa, Cape, Meiringspoortberg, I.1994, AKC
fig. 141fig. 140
Colophon cassoni Barnard
Publication: BARNARD, 1932: 172
Type locality: South Africa, Zwartberg Range, Meiringspoort Berg
Size: m 16.0 - 24.0, f 16.0 -17.0
Rarity: very rare
Remarks: the genus Colophon has been revised by Endrödy-Younga (1988)
and Mizukami & Kawai (1997). This species is active only during proper weather
conditions.
Distribution: South Africa (Swartberg)
93
fig. 142 - Swartberg Mts., Rep. of S. Africa, March 1993 fig. 143 - C. haughtoni - m, Rep. of S. Africa, Cape, Hexriver Mts., Matroosberg, I. 1993
fig. 144 - C. eastmani nagaii - m, Rep. of S. Africa, Cape Province, Langeberg, Dassie'shoek, 29. I. 1993,
fig. 145 & 146 - C. izardi - m, Rep. of S. Africa, Cape Province, Langeberg, Boosmansbos, 27. XII. 1992
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Colophon
fig. 142
fig. 145 fig. 146
fig. 143 fig. 144
142
fig. 268 - m, 32 mm, Congo Belge, Kasai, Kondue, Leohard leg., MRAC, PARATYPUS
fig. 269 - m, 28 mm, Congo Belge, Kasai, Kondue, Leohard leg., MRAC, PARATYPUS
Prosopocoilus kasaiensis Maes
Publication: MAES, 1990: 13
Synonym: swanzianus sensu Auctorum (partim)
Type locality: Congo Belge, Dekese, Kasai
Size: m 28.0 -32.0, f 20.0 - 22.0
Rarity: very rare
Remarks: this species was described as a form of P. swanzianus, then elevated
to subspecies by Maes (1992b); according to Desfontaine & Moretto (2003)
it is a valid species, very close to P. demeyeri, and can be separated only
by the examination of the genitalia.
Distribution: Democratic Republic of the Congo (ex Zaire)
fig. 268 fig. 269
143
fig. 270 - m, 34.5 mm, Kamerun, 1968, AKC
fig. 271 - f, 26 mm, Kamerun, 1968, AKC
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Prosopocoilus
Prosopocoilus katanganus Bomans
Publication: BOMANS, 1967a: 392
Type locality: Rép. du Congo, Kafakumba
Size: m 31.0 -34.5, f 20.0 - 26.0
Rarity: very rare
Remarks: this species is close to P. congoensis.
Distribution: Cameroon, Democratic Republic of the Congo (ex Zaire)
fig. 270 fig. 271
170
fig. 334 - m, 69 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC
fig. 335 - f, 38 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC
Prosopocoilus umhangi (Fairmaire)
Publication: FAIRMAIRE, 1891b: 192 (Cladognathus)
Synonyms: hacquarti Oberthür, 1880: 119
wentzelheckmannae Kolbe, 1901: 71 (Metopodontus)
elizabethae Waterhouse, 1905: 375 (Metopodontus)
Type locality: Zanzibar
Size: m 28.2 - 69.1, f 26.0 - 38.0
Rarity: not rare
Remarks: P. umhangi specimens were collected in numbers by the second author
and by R. Lizler on trees of rain forest in central Tanzania.
Distribution: Malawi, Tanzania
fig. 334 fig. 335
171
fig. 336 - m, 34 mm, Tanzania, Usambara, AKC fig. 337 - f, 26 mm, Tanzania, Usambara, AKC fig. 338 - m, 37 mm, Tanzania,
Uluguru Mts., IV.1997, Werner & Lizler leg., KWC fig. 339 - m, 67 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC
fig. 340 - f, 32.5 mm, D.O.A., Nkonda Uguro, 1910, AKC fig. 341 - m, 57 mm, Tanzania, Uluguru Mts., IV.1997, Werner & Lizler leg., KWC
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Prosopocoilus
fig. 339 fig. 341fig. 340
fig. 336 fig. 338fig. 337
180
fig. 360 - m, 45 mm, Italia, Basilicata, Gallipoli-Cognato, 7.VII.1997, B. Cecchi & B. Carletti leg., LBC
fig. 361 - f, 32 mm, Sicilia, San Fratello, Messina Prov., 1.VII.1984, Aliquó leg., LBC
Lucanus tetraodon Thunberg
Publication: THUNBERG, 1806: 188
Synonyms: for the long list of synonyms see Krajcik (2001)
Type locality: Italy
Size: m 26.1-50.0, f 25.8 - 37.2
Rarity: common
Remarks: L. tetraodon is a palaearctic species occurring in oak forests. In North
Africa the species is quoted only from Algeria.
Distribution: Algeria; Balkan Peninsula, France (including Corsica),
S Italy (including Sicily)
fig. 360 fig. 361
181
fig. 362 - m, 38 mm, Italia, Calabria, Capistrano, Catanzaro Prov., 800 – 1000 m, VIII.1990, I. Bonato leg., LBC
fig. 363 - m, 28 mm, Italia, Calabria, Capistrano, Catanzaro Prov., 800 – 1000 m, VIII.1990, I. Bonato leg., LBC
fig. 364 - m, 42 mm, Sicilia, Messina Prov., P. Femmina Morta, 9.VII.1983, Aliquó leg., LBC
Luca Bartolozzi & Karl Werner ILLUSTRATED CATALOGUE OF THE LUCANIDAE FROM AFRICA AND MADAGASCAR (Coleoptera: Lucanidae) Lucanus
fig. 364fig. 362 fig. 363