final resul ts of the mackerel scomber scombrus , l ... otolith...1 working document for the...

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1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final R esul ts of the Mackerel ( Sco mber Exchange scombrus , L.) Programme in Otolith 1994. by Begoña Villamor(l) and Manolo Meixide (2) (1) . Inst ituto Españ ol de Oceano grafía . Centro Ocean ografico de Cantabria. Apdo 240 . 39080 Santander. Spain (2). Instituto Español de Oceanografía . Centro Oceanógrafico de Vigo . Apdo 1552. 3 6280 Vigo . Spain ABSTRAC T The 1993 Working Group on the Assessment of Mackerel, Horse Mackerel, Sardine and Anchovy agreed to carry out a mackerel otolith exchange programme to solve the prob lem s found in readings. A total of 398 pairs of otoliths was exchanged, 200 of which carne from the we stern area (ICES Divisions VIa and VI Ibc) and 198 were from the southern d.l.t:(l (:iCeS i:i.i v.ib.ivil:::. v::L.i::L .. ClHU :LL\a), (lJHl i..t:u .1.1;:: (1 " ::: rs f.LOlfi ulffe.Lt:uL l:VU11Üie took part in the excha nge. The comparison of ot olith readings of the two areas was carried out separately. The results show an unaccepta ble variability in mackerel age determination between the readers from different c ou ntries. The average general agreement between readers was low in the two areas, 47% for the western area and 51% for the so uthern area. Agreement between readers varies between 19% and 71% for the western area and between 4% and 81% for the southern area . Standard deviations by age groups increase cons iderably for age groups >5 years for the western area sample and for age s >3 years for the southe rn area sample .

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Page 1: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

1  

Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995

   

 

Final Resul ts of the Mackerel ( Scomber Exchange

scombrus , L.) Programme in

Otolith 1994.

 

 by

 

Begoña Villamor(l) and Manolo Meixide (2)   

(1). Instituto Español de Oceanografía. Centro Oceanografico de Cantabria. Apdo 240. 39080 Santander. Spain

 

(2). Instituto Español de Oceanografía. Centro Oceanógrafico de Vigo. Apdo 1552. 36280 Vigo. Spain

 

     

ABSTRACT  The 1993 Working Group on the Assessment of Mackerel, Horse Mackerel, Sardine and Anchovy agreed to carry out a mackerel otolith exchange programme to solve the problems found in readings.

 A total of 398 pairs of otoliths was exchanged, 200 of which carne from the western area (ICES Divisions VIa and VIIbc) and 198 were from the southern d.l.t:(l (:iCeS i:i.i v.ib.ivil:::. v::L.i::L .. ClHU :LL\a), (lJHl i..t:u .1.1;:: (1 " :::rs f.LOlfi ulffe.Lt:uL l:VU11Üie

took part in the exchange. The comparison of otolith readings of the two areas was carried out separately.

 The results show an unacceptable variability in mackerel age determination between the readers from different countries.

 The average general agreement between readers was low in the two areas, 47% for the western area and 51% for the southern area. Agreement between readers varies between 19% and 71% for the western area and between 4% and 81% for the southern area.

 Standard deviations by age groups increase considerably for age groups >5 years for the western area sample and for ages >3 years for the southern area sample.

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The bias plots of each reader against the modal age show great variability in the sarnples frorn the two areas, above all in older ages. Furtherrnore, they perrnit the identification of which readers are biased, and at what ages.

 The coefficient of variation, CV, and the Average Percentage Error, APE, of all ageings, with otoliths ordered according to modal age, is very similar in both areas (except in sorne young ages in the southern area) and rernains at around O.l.(Villarnor & Meixide, 1994)

Wilcoxson's test shows that bias exists between rnost readers in both sarnples.

Considering the irnportance of the age readings in the assessrnent, new investigations are necessary to increase agreernent, and to standardize ageings.

  INTRODUCTION

 The last rnackerel otolith exchange was carried out in 1985 (Dawson, 1986) and the results indicated an unacceptably low level of agreernent for fish ages >10 years. A workshop was subsequently held in Lowestoft in June 1987 to resolve these differences (Anon., 1987).

 In the 1993 Working Group on the Assessrnent of Mackerel, Horse Mackerel, Sardine and Anchovy, the separable VPA results of the Western Mackerel Stock suggested that there rnust have been difficulties in interpreting ages of old fish, since a very high fishing rnortality at age 9 appeared in 1992. This is the 1983 age class, which is weak, while the adjacent 1984 age class is strong. Problerns otherwise appeared in the ages of rnackerel frorn Division IXa. (Anon., 1993).

 The Working Group recornmended that a new rnackerel otolith exchange be carried out in 1994, since there are now new otolith readers in sorne laboratories. This exchange was organized by Begoña Villarnor, lEO, Santander (Spain).

The otolith readers who have participated in this otolith exchange are:

Elena Seliverstova PINRO, Murrnansk Russia

 

Helga A. GillInstitute of Marine Res., Bergen Norway

 Kurt Jensen Danish Institute for fish. and Marine Res., Hitshals Denrnark

 Doug Beveridge Marine Laboratory, Aberdeen UK (Scotland)

 

Alee Watson Fisheries Laboratory, Lowestoft UK (England)

 

Elizabeth Barnwall

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Dept. of Marine, Fisheries Res. Centre, Dublin Ireland  

Jan Beintema RIVO, Ijmuidem The Netherlands

 

Iñaki Rico AZTI-SIO, San Sebastian Spain (Basque Country)

 Begoña Villamor IEO, Santander Spain

 

Maria Manuel Martins IPIMAR, Lisbon Portugal

 This paper presents the results and analysis of the 398 otoliths read by the above mentioned readers in the otolith exchange.

 

 MATERIAL AND METHODS

 A sample of 398 pairs of otoliths were examined. Of these otoliths, 200 carne from the western area (ICES Divisions VIa and VIIbc) caught between January and April, covering a length range from 25 to 46 cm; 198 were from the southern area (ICES Divisions VIIIc and IXa), caught from January to April and in September, covering a length range from 18 to 48 cm.

 Only the otoliths and corresponding dates of capture were sent to the readers of each country. Fish lengths were not sent, since this information could have sorne influence on otolith readings.

 The otolith readers gave an absolute value to each pair of otoliths wherever possible. Doubtful ages were indicated by the mark "7" after the age, and otoliths which were too difficult to be aged or which had been damaged in transit were assigned the mark "-". Doubtful ages were included in the analysis. In sorne cases, Norway, Ireland and Russia assigned the age as 15+ at age 15 or greater, and these readings were not included in the analysis.

 Otoliths damaqP.ci in transit and thosP. rearlings in whi eh a mode was not obtained were excluded from the analysis, and so 198 otoliths from the western area and 188 otoliths from the southern area have been analyzed.

 The comparison of otoliths from the two areas was made separately. An Excell spreadsheet was used for this purpose (Eltink, 1994).

 From each otolith mean age, mode, range and standard deviation have been estimated. The percentage of agreement between individual readers, the mean percentage of agreement of each reader compared to all readers and the mean percentage of agreement of all readers have been calculated. Those readers who have a higher mean percentage of agreement with all readers than the mean agreement of all readers are considei ed the "best" readers.

 The cumulative age composition as a percentage has been calculated so that it is independent of the number of otoliths unread by each reader and the age

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cornposition in number of each reader, of all readers (based on the modes obtained from all readers) and that of the best reader (based on the modes obtained for the best readers) have also been calculated. The cumulative age composition shows which readers are closer to agreement and the ages at which agreement is lower.  The percentage of disagreement by one year, by two or more years in the age readings of each reader compared with the rest of the readers have been calculated. Furthermore, the mean disagreement by one year and by more than two years of each reader and the mean disagreernent by one year and by more than two years of all readers have also been calculated.

 The mean length at each mackerel age was estimated by each reader to show differences in growth curves obtained for each reader.

 Functional regressions have been made between age readings of each reader. Readers whose age readings show a clase agreement will have values of 1 and O for the slope and intercept respectively.  The frequency distribution of standard deviations of estimated ages has also been estimated. The standard deviations have been ordered by age groups(0-2, 3-5, 6-8, 9-11, 12-14 and 15+)for all readers and for the best readers.

 As well as all these comparisons made on the spreadsheet, the following methods have been carried out:

 - Age bias plots average age ± 2 standard deviations of each age reader and all age readers is plotted against modal age.

 - To find out the bias between readers Wilcoxson's test was done for each pair of readers.

 These last methods are sorne of the procedures recornmended by the Workshop on Sarnpling Strategies for age and maturity (Anon., 1994).

 - The Notched Box and Whisker plot is reprented, and is an exploratory data analysis that surnrnarizes a set of observations and is particularly usefu+ for comparinq parallel batches of data.

 

 RESULTS

 

1- Western Area otoliths exchange sample  The readings of each reader together with the basic information concerning each otolith (year, sample nurnber, otolith number, fish length and month of capture) are shown in Table 1.1, as are the mean age, rnode, range and standard deviation for each otolith. Of the 198 otoliths exarnined, 163 were assigned an age by all readers and only 33 were assigned the sarne age by all readers.

 The readers from The Netherlands, England, Norway, Spain (lEO), Ireland, Denmark and Russia have a higher average percentage of agreernent with all

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read of agreement (see Table 1.1 and ers than the average general percentage 1.5)and are considered the best readers.

 Table l.2 shows the cumulative age composition as a percentage for each individual reader, for all readers and for the best readers. The reader from Portugal is observed to have a totally different age composition to the rest of the readers from age 3. In the rest of the readers the closest agreement is at ages 2, 3, 4 and 6 and from age 7 there is greater variability (Figure 1.1).

 In Table 1.3 the age composition in number for each reader is shown. The reader from Portugal has not assigned any ages over 8 years. In Figure 1.2 the reading of each reader can be compared with the ages obtained from the modes of readings of the best readers.

 The average length by age of each reader is shown in Table 1.4 and Figure 1.3. It can be seen that the reader from Portugal has assigned much younger ages than the rest of the readers. The rest of the readers show higher agreement at ages 2, 3, 4 and 6 and variability is greater from age 7.

 The percentage of agreement betwren readers is presented in Table 1.5 and varies between 19% and 71%. The average general percentage of agreement of all readers is low (47%). The highest agreements are found in the readers from The Netherlands and Spain (IEO) (71%)and the readers from Ireland and Norway (71%). The readers from Scotland, Spain (AZTI) and Portugal have a lower average percentage of agreement than the average general agreement, the reader from Portugal presenting the lowest agreements with 22% of average agreement with the rest of the readers.

 The average general disagreement by 1 year is 26% (Table 1.6). The highest diagreements are found in the reader from Spain (AZTI)(31%). This indicates an error in reading technique or problems in the interpretation of the hyaline/opaque edge of the otolith.

 The average disagreement by 2 or more years between readers is 27% (Table 1.7).The highest disagreement is found in the reader from Portugal, with 61% of average disagreement with the rest of the readers. This indicates an error in reading technique or little experience of readinq.

 If the otoliths were read using the same criteria the functional regress1on slope should be 1 and the intercept should be O. The values of regression obtained (Tables 1.8 and 1.9)show a high level of agreement between readers, except in the case of the reader from Portugal for whom large deviations are observed.

 Nevertheless, the disagreements are better illustrated by calculating the standard deviations by age group (Tables 1.10 and 1.11). In Figure 1.4 the frequency distribution of the standard deviations has been drawn based on the readings of all readers providing information on the current level of precision of readings by age groups and showing how the standard deviations increase for ages >3 years. In Figure 1.5 the frequency distribution of the standard deviations has been drawn based on the readings of the best readers providing information on the maximum level of precision of readings by age

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groups, and showing that the standard deviations increase for ages >5 years despite the standard deviations diminishing in all age groups in comparison wíth Figure l.4.  By modal age are presented the average age recorded, the standard deviation and number of age readings by reader and of all readers (Table 1.12). The age bias plots of each age reader against the modal age present great variability, above all in older ages in many of the readers (Figures 1.6). The most biased readers are 2, 3, 7 and 8 (Scotland, England, Spain (AZTI} and Portugal). The age bias plots of all age readers is also plotted (Figure 1.7), and show greater bias from age 9.

 The results of Wilcoxson's test from analyzing bias between readers is shown in Table 1.13. As can be seen there is a clear bias between most readers.

 The Notched Box and Whisker plot (Figure 1.8) indicate that if the age readings of all readers were the same the grafhs of all readers would be the same. It can be seen that the age readings from Portugal are completely different to those of the rest of the readers. The readings from Russia and Derunark are observed to be similar, as are those from England and Spain (AZTI).

  2- Southern Area otoliths exchange sample

 The readings of each reader together with the basic information concerning each otolith (year, sample number, otolith number, fish length and month of capture) are shown in Table 2.1, as are the mean age, mode, range and standard deviation for each otolith. Of the 188 otoliths examined, 148 were assigned an age by all readers and only 3 were assigned the same age by all readers.

 The readers from The Netherlands, Scotland, Norway, Spain (IEO), Ireland and Russia have an higher average percentage of agreement with all readers than the average general percentage of agreement (see Tables 2.1 and 2.5)and are considered the best readers.

 Table 2.2 shows the cumulative age composition as a percentage for ei'ir.h individual reader, for all readers and for the best readers. As seen in the western area, the reader from Portugal has a totally different age composition to the rest of the readers from age 2. In the rest of the readers the cumulative age composition shows great variability at all ages (Figure 2.1).

 In Table 2.3 the age composition in number for each reader is shown. The reader from Portugal has not assigned any ages over 7 years. In Figure 2.2 the reading of each reader can be compared with the ages obtained from the modes of readings of the best readers.

 The average length by age estimated from the age readings of each reader is shown in Table 2.4 and Figure 2.3. It can be seen that the reader from Portugal has assigned much younger ages than the rest of the readers. The rest of the readers show higher agreement at ages 4, 6 and 9 and show great

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variability in young ages (1 to 3 years).  The percentage of agreement between readers is presented in Table 2.5 and varíes between 4% and 81%. The average general percentage of agreement of all readers is low (51%). The highest agreernents are found in the readers from Norway and Spain (lEO) (81%) and the readers from Russia and Norway (76%). The readers from England, Spain (AZTI), Portugal and Denmark have a lower average percentage of agreernent than the average general agreement, the reader from Portugal presenting the lowest agreements with 18% of average agreement with the rest of the readers.  The average general disagreement by 1 year is 28% (Table 2.6). The highest diagreements are found in the readers from England (34%), Spain (AZTI)(31%) and Denrnark (35%). This indicates an error in reading technique or problems in the interpretation of the hyaline/opaque edge of the otolith.

 The average disagreernent by 2 or more years between readers is 21% (Table 2.7).The highest disagreement is found in the reader from Portugal, with 59% of average disagreement with the rest of the readers. This indicates an error in reading technique or little experience of reading.

 The values of regression obtained (Tables 2.8 and 2.9)show a high level of agreement between readers, except in the case of the reader from Portugal for whom large deviations are observed.

 The disagreements are better illustrated by calculating the standard deviations by age group (Tables 2.10 and 2.11). In Figure 2.4 the frequency distribution of the standard deviations has been drawn based on the readings of all readers providing information on the current level of precision of readings by age groups and showing how the standard deviations are quite high for all age groups, increasing greatly for ages >3 years. In Figure 2.5 the frequency distribution of the standard deviations has been drawn based on the readings of the best readers and providing information on the maximum level of precision of readings by age groups, and showing that the standard deviations increase for ages >5 years despite the standard deviations diminishing in all age groups in comparison with Figure 2.4.

 By modal age are presented the average age recorded, the st ndard neviatinn and nurnber of age readings by reader and of all readers (Table 2.12).The age bias plots of each age reader against respect to the modal age present great variability, above all in older ages in many of the readers(Figure 2.6). The most biased readers are 3, 7, 8, 9 and 10 (England, Spain (AZTI), Portugal, Denrnark and Russia). The age bias plots of all age readers is also plotted (Figure 2.7) and show greater bias frorn age 9.

 The results of Wilcoxson's test from analyzing bias between readers is shown in Table 2.13. As can be seen there is a clear bias between rnost readers. The bias between reader is lower in the sarnple of the southern area than in that of the western area.

 The Notched Box and Whisker plot (Figure l.8) indicate that if the age readings of all readers were the same the grafhs of all readers would be the sarne. It can be seen that the age readings frorn Portugal are completely

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different to those of the rest of the readers. The readings from The Netherlands, Scotland, Spain (lEO), Ireland and Denmark are observed to be similar, as are those from England and Norway, and those from Russia and Spain (AZTI).

  CONCLUSIONS

 The results of the present paper show a low level of agreement between the readings of the different readers in the two samples. Of the 386 otoliths examined, only 311 were assigned ages by all ten readers, and in only 36 otoliths was the same age assigned by all readers.  Portugal has a very different criteria of reading to that of the rest of the readers, constantly assigning much younger ages.This can be seen in both the age composition and in the average lengths estimated in both samples, as well as in the values obtained in the functional regression.

 The average lengths present higher variability from age 7 in both samples, and furthermore, in the southern area the young ages (1 to 3 years) show great variability. The growth curve of the reader from Portugal is totally different to that of the rest of the readers.

 The average disagreements by 1 year between readers are similar in both samples, 28% in the western area and 26% in the southern area.

 The average disagreements by 2 or more years between readers are somewhat higher in the western area (27%)than in the southern area (21%), owing to the fact that in the western area sample there was a greater number of old fish than in that of the southern area. Portugal presents a very high average disagreement with the rest of the readers (61% in the western area and 59% in the southern area), which means that this reader used a very different otolith reading criteria to that of the rest of the readers.

 The values obtained from the functional regression indicate that the otoliths were read using similar criteria with the exception of the reader from Portugal, who presents great deviations in the results.

 The standard deviations estimated by age groups are low for the young ages (O to 2 years) in the western area sample and after these ages they begin to increase. In the southern area sample the standard deviations are high in all age groups. Considering the readings of the best readers the standard deviations diminish. Despite this, the standard deviations are high from age 5.

 The age bias plots of each age reader against the modal age presents great variability in the two samples, above all in old ages in many readers. The age bias plots of all age readers show greater bias from age 9.

 The coefficient of variation, CV, and the Average Percentage Error, APE, of all ageings, with otoliths ordered according to modal age, is very similar in both areas (except in sorne young ages in the southern area)and remains at around O.1 (it may be important for the assessment that CV does

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not increase with age, since sorne tuning methods assume CV constant with age).(Villamor & Meixide, 1994)  Wilcoxson's test shows that bias exists between most readers in the two samples.

 The Notched Box and Whisker plot indicate that quite a lot of differences exist between the age readings of all readers in the two samples, above all in the western area.

 The absence of bias is a minimum requirement and in view of the results and considering the importance of the age reading in the assessment, new investigations are necessary to increase agreement and to standardize ageings.

  REFERENCES

 

Anon., 1987. Report of the Mackerel Age Determination Workshop. Lowestoft, 1-6 June 1987.

 

Anon., 1988. Report of the Mackerel Working Group. ICES C.M. 1988/Assess:12  Anon., 1993. Report of the Working Group on the Assessment of Mackerel, Horse Mackerel, Sardine and Anchovy. ICES, C.M. 1993/Assess:19.

 Anon., 1994. Report of the Workshop on sampling strategies for age and maturity. ICES, C.M. 1994/0:1 Ref:G,H,J.

 Beamish,R.J., and Fournier, D.A., 1981. A method for comparing the precision of a set of age determinations. Can.J.Fish Aquat. Sci. 38: 982-983.

 Chang, W.Y.B., 1982. A statistical method for evaluating the reproducibility of age determination. Can.J.Fish Aquat.Sci. 39: 1208-1210.

 Dawson, W., 1986. Analysis of the Mackerel (Scomber scombrus L.) otolith exchange programme. ICES C.M. 1986/H:25

 Eltink, G., 1994. Comparison of otolith readings. Working document for the Workshop on sampling strategies for age and maturity, 1994 at ICES Copenhagen.

 Villamor, B., Meixide, M., 1994. Results of the Mackerel Otolith Exchange. Working Document for the Working Group on the Assessment of Mackerel, Horse Mackerel, Sardine and Anchovy. Corenhager , Junio 1994.

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500 1 0 e 03570 e o 16 3 300 3 o 000 seo e 2 080 540 1 040 5 Ja.? • 2 080

&40 9 1' 5 70 1 o 48 e 580 e 2 080 7 56 7 S 184 550 5 1 050 580 6 o 49 1 1213 13 e le!! 500 S 2 04S 880 g 10 273 840 e S 150

 770 8 155 6 900 9 • 110 7 57 g 168 5 a40 8 • 111 1014 12 8 2 70 e22 10 8 ve 944 12 275 g

3 o 000 300 3 o 000 2.3:) 1 0 48 2 4 56 • 3 098

300 3 o 000 300 3 000 o 300 3 o 000 300 3 o 000 300 o 000 3 390 • 1 030

560 e 2 oso 530 6 ,00 2 017

2 078 '

600 e 000 o 820 8 S 125 967 10 087 2 750 8 e 1 75 856 e 2 088

590 1 030 6

,.11 78 15 11

400 356

5 1 030

300 4 360 • 1 0 49

1 030

300 • 1 030

4 30 4 3 090 638 7 2 070 680 7 3 075 68C 7 3 075 680 7 3 075 800 8 5 145 913 8 3 105 900 11 8 2 38 860 7 5 135 750 7 5 1 43 1200 12 12 316 8 80 4 1087

ll W 11 :l7i.. 2 7 8 e 1 59

1156 11 068

 12.C,t •2 e 150 1356 13 3 096 1213 18 12 4 23 610 G 4 09: 950 10 G 1 57 800 8 5 161 10 4-4 ,1 1 1 63 0

2

1020 o ,600 ? 6 , 51

 1000 10 5 , 41 920 9 8 160 tOOO 11 7 184 S70 9 9 233gao 10 5 133ero 10 o 174 840 s•• 10 3

8 S 138

096 940 10 S ISO 9 3:) 10 5 1 27 94 10 3 0920

15694.0. 10 8

., 271

6 13 7 2 0113

1233 1313

1258 ,. 310

8 12 44 • 117

156

11 200 2 o 000 2JO 2 000 o 200 2 o 000 200 2 000 o 200 2 000 o 200 2 0 00 o 200 2 0 00 o 200 2 o 000 200 2 000 o 200 2 000 o 200 2 o 000 2.00 2 000 o 2 00 2 000 o 200 2 o 000

00 • 14 40. 1  00 • 1e 40e 1 7 1' e 10 g 10 9

 llO

 4

 111 .,.   1  

9

 g

 g  

10

 g

 10

 g

llO • '22 420 1      00 • 24

 

43 7 1 12 13 7 12 11 6

00  

298 2      

  4

• 41 1 • 41 3 4

a 426 •

  425 •

13 420 4 424 4

15 13 12 12 13 1213 15 12 13 1) 1313 14 10 12 '3 13

11 32S 2 4 S

 5

 S • S • 3 5 S • 50

 S

 2  

067

11 379 2 6 S 4 6 5 e S 4 6 6      11

, 16 382 2 6 5 8 6 8 8 5 4 8 8

 

 6

5 70 8 2 084

 11 23 418

 2

     

 1S

 

 15

 

 15

  

g

  

10 10

  

10

 822 1200

 8  

2 4

 083 158

S 5 S 5 5 5 S S 5

13 13

2 2 2

91 13 13 39 2

 91

 13

 18

 409 2

91 13 17 415 2 9'1 13 18 41 2 2

4 4 3 4 •

• 8 2 2G

91 35 19 438 4

' 9

, 1

 8

Tab1.1. WESTERN AREA .MACKEREL rom Dvl .VIa and Vllb,c and rom January • April

  

00 1• 2 3!>0

90 14 • 3S9

90 • e 382 1

 00 11 1• 00 JGO 1

3G8 12

 3 3 3 3 3 3 33 3 3

  5 5 S 5 S S S • 5 4 4 80 040 S 1

 6

  

e

g 10  

 

• 8 10 3 300

 00 11 3D 2 e S S • • 5 S 054

S 5 5 5 400 2 5

00 7 322 2 

 00

 00

 00

  

 00

 90

  

 91 13 2 5 387

    

• 4 • 3 3 • 3 4

 

 820 3 0876

4 4 • 4 4 4 4 • 3 4 4 13 e Je2 2 390 • 1 030

7 7 o 5 o 8 7 7  

5 8 7 7 89 8

g 8 7 5 9 11 10 1

1 10 0 8 8

1 9 3

    

91 13 21

 

 91

4 2 . 1 10 10 11 8 11 10 12 '2 •e 13 13

987 .. 12 12 10 12 12

 

6 8 6 7 6 8 6 5 • 14 11

6

10

G

 35 10 10 10

91 10 11 10 11 e 8

10

11 9 V 11 10 10 9 g 9

1 47 91 7 8 7 6

13 11 12 13 12 11 6 7

91 351100 7 100

 

1' 91

 91 111 35

 91 35 16 432 • 91 35 20 ... o •

10 15 9 9 10 7 ! 9  

 1 80

91 45 s 4 9 5 •29

12 10

91 35 2• 480 • " 92 35 1 3 2 2 2 2 2 2 2

Page 11: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

.e 2 348 2 5 5 5 5 S S 5 3 e 480

e •

• •

3 ,.

,.

" ,.

8 13 S

1 :;ee 3 4 S 5 8 5 e S 4 5 e S20 6 2 on

373 2 S • 5 8 7 4 4 8 533 5 4 33 1

28 1S 428 3 12 .. g 1 10 10 13 9 e 10

6 g"

28

26

,.

" 233

e 20 408 2 12 6 12 11 12 10 5 11 1030 210" " , 7

30 12 <I V 3 10 14 13 1' • 11 , 209,. ,. .. ,." 10

435 3 11 9 g a 9 8 9 7 11!9• 850

'30 14 4n 3 S 8 9 g g g 5 g !> 8'10 ' 9 9

8 8 1 78

30 15 420 3 14 16 14 13 ,. 14 11 5 13 14 1260 ,. 11 2'U"' 30 16 42 2 3 g 9 g g 8 e <O 9 863 • 1111' 30 17 438 3 11 1S 13 11 11 12 S 11 9 1133 ,. 6 176

3S 1S 281 3 ¡ ¡ 1 2 2 2 2 2 2 200 2 e 000 3S 18 287 3 ¡ 1 2 2 2 2 2 ? 2 2 200 2 o 000 3S 17 2111 3 2 2 2 2 2 2 2 2 2 2 200 2 o 000 3S 18 2117 3 2 2 2 2 2 2 2 2 2 2 200 2 o 000 3S 19 211 7 3 2 1 2 2 2 2 2 2 2 2 200 2 o 000 3S 20 2111 3 2 2 2 2 2 2 2 2 2 2 200 2 000 () 3S 21 307 3 2 1 2 2 2 2 2 2 2 2 200 2 000 o 3S 22 308 3 3 3 3 3 3 3 3 3 3 3 300 3 o 0 00 3S 23 310 3 3 3 3 3 3 3 3 3 3 3 200 3 0 00 o 3S 24 31& 3 3 3 3 3 3 3 3 3 3 300 o 0 00 3 3 3S 25 3&4 3 8 6 5 6 4 3 7 e $30 e 4 1 18 8 1 328 2 3 3 3 3 3 3 2 2 3 3 280 3 1 040

S S 3 070 6 3 346 2 4 4 4 4 4 4 3 4 4 390 4 1 030

4 343 2 3 4 4 4 3 3 4 4 3 70 • 1 0 46

6 6 S 352 2 S S 5 5 5 6 S :;es 2 • • 4 • 4 5 5 • 3 6 7 3&3 2 5 S 4 5 5 • 4 5 4 •50 5 1 050

3 2 S 5 480 5 2 080• • 3 70 4 1 064

6 6 37 2 2 5 • 3 4 • • 4 3 4 4

3 2

378 2 8 9 4

S g 8 g 6 4 8 9 e S 168 057

8

,e 10 770

37 5 2 6 7 7 7 7 7 7 5 7 8 seo 7 3 075 37 2 2 9 8 9 9 8 ll S S 4

6 9 9 800 9 155 8 38S 2 6 6 6 8 5 8 3 8 8 558 6 3 OliS 12

,. 380 2 e 9 1 9 8 5 8 8 8 733 8 4 133 e e 1S 391 2 10 11 10 12 10 13 7 8 10 11 1000 10 7 200 8 15 390 2 e 5 5 tS 6 e 5 4 6 8 550 tS 007 2 8 17 393 2 13 12 13 12 13 11 5 13 13 1180 13 o o 18 402 2 9 9 9 7 8 7 e 6 9 780 9 128 8 3 8 19 408 2 8 9 e 7 8 7 7 5 7 7 7 10 7 • 1 ()4

8 21 408 2 9 9 8 9 6 9 8 e 9 g 840 9 3 0112 8 413 2 9 10 9 9 9 9 7 9 9 869 9 3 074 22 8 n 410 2 9 8 11 9 9 8 g 10 9 44 9 8 208

24 43 1 2 11 13 12 13 12 13 5 13 1120 13 8 240 6 25 458 2 17 14 16 15 13 7 18 ,.14 18 10 318 28 28 2 318 3 ; S S S 5 4 5 480 5 1 (S )40 28 3 379 3 6 6 5 6 8 6 5 4 e 560 6 2 o u 8 28 3&8 3 6 6 e 8 e 5 4 6 e $10 S 2 064 6 26 3&7 3 5 6 e 7 e 8 4 7 e SAO S 5 138 28 6 383 3 7 6 8 8 6 7 5 7 8 71() 7 S o 94 7 26 7 385 3 6 6 6 tS e e e 5 7 7 510 6 2 054 28 8 398 3 6 8 8 8 8 6 7 5 7 9 700 6 116 26 9 <400 3 6 6 4 e e 7 5 4 6 550 E 3 092 S 28 10 3&7 3 7 8 6 8 8 9 8 4 a 10 780 8 6 158 28 11 407 3 7 g 1 8 7 9 7 S 8 8 750 7 4 112

26 13 404 3 6 7 e e e 5 7 e 620 2 080 7 6 8 28 14 407 3 8 •o e 8 11 11 5 6 g 867 a 17e 6 26 15 413 3 7 i 7 9 7 7 6 5 9 g 710 7 • 127

1000 10 8 2 37 11 42 4 3 13 13 8 13 13 13 11 e 8 11 10 13 7 2511 13

28 18 •3 S 3 12 15 g 10 12 11 10 4 10 •o 1030 10 11 2&5 26 19 439 3 1< 1' 12 12 12 11 8 11 10 1110 12 8 197 28 20 439 3 12 14 12 13 12 12 e 13 13 1210 12 8 211

21 440 10 11 g 12 11 9 5 12 13 1022 11 8 225 28 22 44 9 3 13 19 11 15 15 13 S 15 1325 15 380 26 23 45 3 3 15 18 15 14 15 7 ,'5. " 1'13 15 11 293 26 24 453 3 13 10 15 15 13 S 1238 13 10 318 28 25 488 3 10 g 8 10 11 Q 5 8 10 869 10 8 168 30 1 lS7 3 S 6 5 S 5 S S 4 5 5 500 S 2 0 45 30 2 388 3 6 6 6 8 5 e 5 4 S 5 550 6 2 067 30 3 392 3 8 7 6 6 e 7 4 7 6 600 6 3 089 5 30 383 3 g 10 9 9 8 Q e S 9 g 850 9 S 1 28 30 5 39 1 3 8 10 g g 9 7 5 8 830 9 S 9 9 13$ 30 6 406 3 g 9 g g g 9 7 4 g 9 830 9 5 · ss 30 7 407 3 9 9 10 g g g 8 5 8 850 9 S 128 9 30 6 405 3 8 g 8 g 8 g 8 5 g 9 820 9 4 117 30 9 .1 o 3 ¡ 7 7 7 7 7 7 S 8 7 890 7 3 070 30 10 40 9 3 7 6 6 7 7 8 7 4 6 7 630 7 3 ovo 30 11 412 3 8 8 8 8 8 8 e 4 8 8 780 8 <20

13 •2 e 3 9 8 11 7 S 10 7 813 6

30

30 18 •3 8 3 9 11 g 10 10 12 8 e 10 11 960 10 6 1 62 30 19 44 o 3 12 13 13 12 12 12 12 12 13 1180 '2 5 1 37 6 30 2() 433 3 9 5 5 7 5 • 6 6 576 5 5 1 40 30 21 9 9 9 30 22 444 15 14 10 14 13 e 5 15 11 , ;a 15 10 3 15 3

30 25 4$8 ,..,3,..

 20 22 20 20 13 18 1683 20 9 as

30 23 •4 2 3 13 17 g 10 12 13 S 10 12 1122 13 12 3 12

l 30 2• ..2 3 16 16 15 15 15 12 8 14 15 1389 16 10 303

, 1 16ll 1\ lit; 1 154 W5- \QO 1SQ. 95 14 116 TOI01 NOT •toe o 9 o 28 • e o 14 3 6

Emp1yiQYh 2 2 2 2 2 2 2 2 2 2 N N y y Aoreemt- tabove 1\'e'IQ y N y y y V

 

Page 12: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

 N•l!\eri Scoc!ond Eond N<>'W>y S<>{tEOj 1•• ond SPIAZ" Po<tu<,;al Oonmort<

Reader 1 R A•.Oer 3 RUdet t RUdf'f S Re er O Rea!Jf'• 1 Uder 8Re.ott $ •011 RRt"'"""'0Atl

r adetsBEST

re acle o 1 2 3• S e 7 & o

,0

12 1,3. 15 18 17 18 19 20 21 22 23

4 25 25 27 28 29 30 31 32 33 34 35 3& 37 3& 39 .00

..... · ,.,. ... ,.,.' 1''!1. , 1 111(,

- ,.,.

' -!0 24.. 27.,.

.. .3. - -24% ..... 3010 39't !);% 33... 31% 2•' 25'0 :4% ... 25% 24...

..-.. · 31%

4 5

S

- - - 100'14

O'!'

-$-5'14 s

-m.

- 98%

-- t-7% · 100'10. -- -

010 0'10 '1'4 ,;,. 23%

......53%

7:¡.,. 8()'!(,

SS% a.."%

-93'14

-10010.10010 10010.1()0% 100'14100!0 100% 100.. 100% 10010 10010.1()0% 100'!1.100% IOO'!E. 100% 100% IOO'lo

 11'4

 23... 31% 45'11. 55% e;¡.,.

- 84...

113%

- - 100...  

100% 10010 100'14 10010 100'\0 100!0 10010 100!to IOQ!r,

10010 10010 10010 100'14 100'14 10010 100'11> 10010 100'1.

. oe:ono NO<Wiy "Pii.OJ_ "•••no " OttV;a -lJiñm•rK u5J e RU(fe.! 1 Re,a,(Mr 2 Rudtr 3 Reader ¡j Re;u:J•IS Ruder 6 Rudet 1 ;( ad'r 8 RuMor $ Reder 10

All

re.ad• SIOST

read•rs

o 1 2 3 4 5 8 7 8 9

,0

12 1,3. 15 •& 17 18 19 20 21 22 23 24 25 2\l 27 26 :!11 XI 31 32

3,3. 35 3ll 37 3ll 39 40

 22 22

15 9 12 1! 13 1'

22 22 22 22 23 23 21 21

,. ,. 12 12 16 12 12 12 15 31 14 13 48

17 13 15 10 se 17 13 18 :11

,. 2< 2• z• 28 ·e 21 24 XI •2

16 9 15 13 18 24 5 17 15

\6 12 18 " 8 2C 2 li 18

13 15 18 22 16 :1 ,Q. 17 17

10 8 9 12 10 4 8 3 8 7 g 6 8 8 10 3 11 2 • 3 8

2 5

 22 12 12 15 26 17 15 19 16 10 8 9 4 S

 22 12 12 18 27 1i 14 20 18 7 10 8 3 8

18lr 198 169 1 - 198 198

010 ()'!(, 010()'!(,

 20'0

0'10  

'8'1.

0'10  

18'0

"" '"'OO.

Tabel 1.2. WESTERN AREA . Cumulative age composition by aoe rt'ader in percent.age. Cumula1lve age compositoi n of all and bect readers obt.ained from the modea.

 ¡m

0.. ()'!(, 010 010 010 010

010 ()'!(,

 •ll'loo

5'!1.  411"0

2l!'!lo '"'

  

0'10

""'

96'!1. % S<%

&4%

'""' 18'4

     31%

  010

""' 11'4

45% 47% 53 Só... s SO'!'

MIO &5% 7 &4%

 

97% 94'14

;¡.,. 48'4 54%

SS'ti>

 

... 8"'

$!>"'

7:2'4 78'14 71... &8'14 82'< 7&11. Wl4 74.,. 7 1 &7 &5% &4%

- -

 

,.,.

71... 100'!1. 84% 100'!1.

7:¡.,. 81.,.

- 100'\0 100'l4

- &:N 84'14 e3'l4 110'14 eolio 96')1 10010 nll g;¡.,.

90% i!>lfo

10010 96'!1. 100% 100% IOO'l4 100'14 10010 100'!1. 100'10 100% 100... 100... 100% 100% 100% 100% 100% 100% 100% 100%

100% 10010 100... 100% 100% 100% 100'1> 100'l4 100% 100'l4 100'!1. 100'!1.

2 100...

10010 10010 100'14 10010 0010 100'1\  

          

Tablr 1.3n. WESTERN AREA . Agr composltion by age ruder (in num berc ol otolith• read Age composotion ol all and best readers obtained from the modes.

r"'A!i'r'   

'3

  

20 19 21 19 11  

:1$  

25

2

24  

22

12 14 7 6 8 10   6 1

 5

 5    

3    5      

Page 13: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 NeU>oif - and tng and Nor.,.y Sp(IOI lreland ::.ow 11 >'O!tVgal Uinmari<

Reedet 1 Re•d-e1 2 Rud•t 3 Re•der 4 Re.dr S Re•doer 8 quclet 1 Reader $ Ruder RU$$13

8 Reader 1(ALL

reade,.a<:ST

re•de o 1 2

S 6 7 8 g 10 11

12 1,.3 1S 16 17 18 19 20 21 22 23 24 25 26 27 26 29 30 31 32 33 34 35 36 37 36 39 40

3·1g 319 328 319 31 g 319 J.& S 320

363 378 .167 36 1 38S 37 9 393 42 367 0 403 385 40 3 393 40 3 40 o 403 43 5 39 7 •o s 398 41 e 401 405 400 41 4 42 2 41 S 112 409 41 7 40 4 .,7 407 430 41 5 429 414 419 •2e 41 7 41 e 42 3 42 4 419 42 2 ••o 425 41 e 432 428 430 435 <20 432 42 7 43 3 434 431 •3 3 ••e 432 433 429 435 428 432 43 3 428 433 438 45 8 44 9 444 41 5 43e AS 2 A3 2 45 3 44 0 450 453 450 44 6 44 8 •• 5 44 2 458 •s s 458 458 44 2

<53 458 •• 9

458 •se 458

458

 27 g 317 358 359 38 9 404 405 408 426 421 44 5 42 9 44 1 442

 280 31 g 353 370 362 402 408 408 42 7 •2 1 439 43 S 439 44 8 451

 

 458

  

3

 

      

Table 1.4. WESTERN AREA.Mean lengttl ala¡¡e by otolilh r (without correclion lor asure<! below) Age moduof all and best readers used for 11M catultion of mun lenglh.

 

19

 38 4 400 408 41 o 428 418 439 432 439 •• 8 458

 

 458

r""m""  

280 280 280 280 280 280 282 28 4 279  

356 .167 3 3H 3e7389 37 3 377 358 388 368 377 •o g 37 1

280  353 368

 

Page 14: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

.

·

:; ts.... 2•"'

',,.

R ;: d6 :

:

::!•', 1·

Table 1.6. WESTERN AREA. Agr ment ('1.) of otolith readings.

 el'oie4!heerfi t R·u;:er 2

EOQlOnd Ru<$e·.c "'.os Ze =; R.\:$1.1

Ruderl · 56'11> !>6% 71% 5<% 7."!1. $2. 55%

Qu4e·3 Rtao.r5 '1. .6<,10

4211, 81% .a... .. lll'lo

.. :20% S?%

Reader 57" 3&" 47" 5211.

-:20%

 

o.. 23%

Reader 3 47% '"' •e" 2T" 5<% 48%

Re.dcr 4 81% 71,_ ., eaO.r 5 57'!j,

21.. 57... u.. 58"

Re-•8 41.. :20% S3% R.-.7 27% 47% Rude•e

q.. Reade1 10

';5..

 Table 1.6. WESTERN AREA .Oisagrment of one year (%) beiWHn otohlh readlngs.Thls eouldbe dueto

mlsintetpretatlon of !he hyalin<!lopaque tdge of lhe otolllh).  

Nethe" Uand €1\g!ano Nof'w1!1y Sl>(lt<>i. PO!tugal Denmark _ Russ a R der 1 Fteader 2 R Ruder 4 Re.a Re ra Re r9 Re-10 uder 3 tNrS

2¡,. 2'"" : 22". 30% ''"' I N 3010

.Re.ldef 28"' 18"' 3010 Re-ti

24"'24... 32'0. 20'01. 3,. Ruc:ter3 33.,

Ruder 4 77"' :20% IS% 27,;, 24"' R(l•der 31% :20% 3010 Read•re 34.. 18"' 18"' R.ud•r7 21% 32'% 3N Ro-.8 16'0 Readfo•.

Rea6<o< 1Q  

  

Table 1.7. WESTERN Af'·EA .DisaQrHment of two or more years (%) beiWMn otolith readings

Ne:l'le:1 · n Eng nd OIW>y S<:(tu 1 Portogal 1\.1'\'Urk _ RI.ISS.I .

Reaaer 1 Reader 2 Reader 3 Readr 4 Reaoer $ Reader 8 Ruder 9 Reader 10

Reader \ 27% 20'01. 15% 7% 16% 23% 51% 17% Reacer 2 31% 15% lnl 851!. 24% Z•'ll Rea<1e" 1t11. : SS IS Rt:aott 4 1,. 9'!0 64% la% 13"' Readef 1211. 111'0. 60'!1. 13'1(, Re.adtr6 25% 64% 18% 13% RNde: 5<."' 17'!j, 7 21"' Readtr e 6111 85'1(, Reade• uo•f 1C

15"'

  

Table 1.8. WESTERN AREA. Slope from funetional linear regresslons of age determinatlons (ahoudl be close to 1}.A devlation lrom 1indicates that eer1ain age groups are aged older or younger.

 

y.,.. ""lMII .. Eng.ond """"'"v _so ::o ireta.nd SO\ ;; i'ol!ugal !?"M'.ark RUlolola Sccr.a.X-a>a• Ruder \ •.a., 2 Rea:Oer 3 Rude14 Re er Read:er 5 Rtadoet7 Re.acer & Rnder S Reade•.IO

Reao•r \ 116 o 103 103 3 037 Qg(j o • 08Re.Jdtr 2 086 080 0 89 086 o 73 033 0 61 082

Re.ot·• t96 1 12 O el! ReMr3 109 112 115 OllO 0 41 101 101 108

ose 101 064 OAO 086 09-0 React.• S 101 1 17 Qg(j 10< 10< 0 64 03& 0 94 8 00Re-:6 097 114 087 0 99 0 96 082 037 0 91 o e.aoe' 1:2'1 1 37 1\1 119 122 044 112 , 10 7 f<eadtll a 2 70 307 2 •• 250 285 173 228 251 2 •6

108 123 OQ9 106 , 10 0 <1() , 02 106 1 22 099 106 •O< 109 Gt 041 098

  

Table 1. 9. WESTERN AREA .lntercept lrom fur.ctóonallni ear regressions of age determinations (should be elose toO). A d!!vlation lrom O lndicates a consisten! misreadnl g of 1or more years or lhat certain

"""' ·. age group$ are atJe<l older or younger.

 x.....

S<locono E na ....,_., Sc>(<EO ..,..,. Po<t..ga .,....,. Rusw Ruder'\ R•ader 2 RNder3 Ru .,4 qeaoet S R._:ade'& R•aW 7 Re;Kt*' & Re.ader 9 Road••1C

R•ed•l1 -069 -0 41 -001 -013 055 1 49 8 014 010 02Reaar2 059 oss 012 055 0 54 090 185 o 75 081

Read•r -011 -086 -052 -0(); .() 42 Read., • OtO .() 15 048 o•• 022 06-0 136

 

Page 15: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

082 055 RulHf o .()<54 008 .O <E .on 051 02< OC$ 147 R. .d••! 013 -082 037 -02:! 011 OS3 ue 035 023 Reac•r 7 .067 123 -051 .() 76 -080 -Oe$ 130 -032 -022

 

 o 15 0 69 OOt 051 013 026 .().,

-Re-ader

 -028 .o llO -020 -000 .()26 ·039 o 138 -015

 .() 16

Ruoer8 .- ro 506 ·34't -3 39 -38Q ·•03 ·2 95 ·3 47 ·328 ·372

1 .() 1& .() 7 4 .()l . -0 38 .() 10 -0 020 '33 o 14 -001

 

Page 16: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

Table 1.10. WESTERN AREA. Frequency dislfbi ution of STOEV frorr. all readers.

Silt tnt rv¡!

oso i AGE ";! AGE A<.it Gt 3-5 9-11 "·14 15•

g '5, 120

7 2.0

H 300 39 45 420

3110

51 •ao 57 5.0 63 600 69 eao

:r. a o o o 6 33 g 2 o o 2 ,g "' 5 e

o o o • 6 2

,o o o o o o o o o o o o o o o o o

o o o o o o

 

...!!U

Sllt t !f't.I'VJ

oso ().2 A :" 12·14 !'   AGE AGE

000 080

15 120 21 160 27 2 40 33 H 380

300

45 4 20 SI • ao57 ;.o 63 600 69 680

2,1 2• 7

,,,3

, o 22 3 10

• 25 10 o •,

•o 2

', 6

o o , o o o o o

o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o

 2,1 , 1 o o

·ao

-""'--

• • o o o 10 • o o o o o o o o 2

o o

Jable 1.11. WESTERN AREA.Frequen<:y distrbi utlon of STOEV from lhe rudera wilh en agreement above avere.

 

   

5 3 2

 

 

· Z1 10

2< .o 61 45 , g o. "" eo

 

Page 17: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

l ,,,e _ _]

 

MACKEREL (Div. Vla and Vllc) EXCHANGE SA,\fPLE First rea<Hn  

Tabl !.f¿ By modal age •1e- pruenthe rvu•g• "11'r i!!Corded, thedt!v and numbef Of

ag dln9s by re.'ldt-r an<l or att l'l!iln;-  

 REAOER 1

             

 T

1 1 • --- 1J;¡.'ll--'':7.T-;:;'f: -7:::T .:=::..¡

,ct I,CC 1 UO 1. 75 1 1,54 ---1 ---1 ¡3;;3; ;_,...""'=-:...;-=.:..-..;::,;.=-+-":'::;_¿_

\& ,. . l t;::,;.=-+-":'::;_¿_

\& ,. . l t _.-:'..:---'---'-- ;;._--=---L - -- 

          

,. 15 11,()() ,1Ji0

 

           

T.-.- -REAOER 10

¡ ·..• • o 1 , 2_ 1 ., .t.a• n-<:or<•• :. -I----HH:.5 1::.o -•:.''=n';-:-¡.--é_:)&-:--=t-.',_7::e:.--,;..,7:".'J:'::, -'--e:.-,7:·-;a".&;;;-

. ·. , • • .. 1: 0 " . 1z 't ,, · -

-*

113.:: :::

5. ro,7e · 1 .oow oo "r7

1 12

rmr - . . e_ ., oo o1a. 2·o 1.8.51.7o _ _!.n ,40 u '".....!-,..j4-•!! . 2n -....1_j_ o 12 · •<.... 1& 20 1 l t 11 1 10 -o t 1 O 4

   

            

Page 18: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 G00 /100 OIA '0'3'1 HC6 t B S t C-.Q, fll: n S6 . z o ,r.o

 

 

Page 19: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

      

Table 1.13. test of the sample of Results of Wilcoxson's Divisions VIa and VIIbc.

  

READERS 2  

** ** ** - ** ** ** - - 1  

  ** - ** ** ** ** ** ** 2  

    ** ** ** * ** ** ** 3  

      ** ** ** ** ** ** 4  

        ** ** ** - - 5  

          ** ** ** * 6  

          ** ** **

       

 7  

    ** ** 8  

                * 9  

       10

3 4 5 6 7 8 9 10

   

A* : Possibility of bias

bsence of bias

** Certainty of bias

 

Page 20: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

f,., fs:> C.!di Yeaf -pie no ..,!h mon:!t Rucet • ea4er2R.-3 q...,.,. R R..óe<6 - q:UO.f8 a9R•aoe< IC

"""'G<' .\Od<! R ST

1 1 2:•8 3 93 1 3 316 3 93 1 4 24 2 3 93 1 5 309 3 93 1 6 248 3 93 1 7 23 4 3

so

93 1 8 238 3 93 1 9 23 2 3 999

3 10 24 3 3

1

1 ,3 1 12 239 3

,12 ,

3

, 93 1 18 462 3 99

3 1 17 3.6 3

1

4443 16 4 3 1

93 , 20 437 3

2

93 1 21 41 a 3

93 1 22 421 3

93 1 23 454 3 93 1 24 •37 3 93 1 2e 439 3 93 1 27 42 4

2

93 1 28 42 5 3 113 1 29 42 4 3 113 1 30 370 3

9

113 1 31 465 3

113 32 418 3 1

93 34 431 3, 93 1 33 429 3  

2

93 1 3$ 334 3 93 1 38 3-03 3 93 1 37 2< 2 3 93 1 38 25 6 3 93 1 3 24 8 3 93 1 234 3

40

93 1 4.2. 44 6 3

93 1 41 44 : 3

1

 

3 80

 

93 1 45 3-02 3 93 1 48 3CS 3 93 2 1 389 2 93 2 2 395 2 93 2 3 388 2 93 2 4 382 2 93 2 S 403 2 93 2 8 37 S 2 93 2 7 358 2 93 2 6 391 2 93 1 9 :398 2 93 2 10 37 3 2 93 2 \1 381 2 93 2 12 39.1 2 93 2 13 301 2 93 2 14 383 2 93 2 15 397 2 93 2 16 386 '2 93 2 385 2 17 93 2 •a 370 2 93 2 2'.) :!77 93 2 21 331 2 93 2 22 33S 2 93 2 23 367 2 93 2 24 365 2 93 2 26 384 2 93 2 27 377 2 93 2 28 411 2

93 . JV .><o < 93 2 31 389 2 93 2 32 386 2 113 2 33 383 2 93 2 34 355 2 93 2 3S 370 2

2 38 39. 2 6

93 2 :!11 · 2

2 37 5 2  

 93 2 4' 38 " 2 ¡¡:¡ 2 2 •2

93 2 36 2... 93 2 43 362 2 0

93 2 45 37 o 2 93 ' 46 37 6 2 93 2 41 302 2

93 3 , 42. 3 113 3 2 3 420 93 3 3 3

.,( 19

, 4

 93 3 6 15 3 93 3 1 42 . 3 93 3 8 392 3 93 3 2 5 3 9 4

3 3 , 425 3911

3 3 413 312

93 3 407 3,. 93 3 13 421 3  

919

3 3 15 3 (3 4

13 3 18 .,o 3 3 3 17 370 3

93 3 18 400 3 93 3 t9 43 8 3 93 3 20 443 3 93 3 21 417 3 93 3 22 41 8 3 93 3 23 418 3 93 3 24 421 3 90 3 25 408 3

3

2 2 1 1 2 1 2 3, , 3 3 3 3 3 3 2 2 3 3

3 2 3 2 2 2 2 2 2 3 2 2 1 1 2 2 1 1 31

2 1 1 1 2 1 2 1

,2 1 1 1 2 1 ,

2 1 1 1 2 , 1, 2 1,

2 1 1 1 1 1 2

, g , , 6 , 10 4

13 8

, 10 10

, ,9 10 10 10 10 9 10 6 10 10

1 g 13 7 11 10

11 12 12 11

13 o 10

, , , , ,8 9 10 10 9 9 4 12 9 4

10 11 10 10 10 12 oo • 11 10

1 u 13 15 12 5 13 ,12 10

,12 12 12 12 1 , 5 1211 1 1 9 1 11 13 19 9 9 g g 9 5 8 9

9 9 9 9 $ 9 9 9 9 9 4 4 4 • • 3 • 4 • 4

12 12 , , 12

1' 11 S ,. 11 , 11 12 11 \1 • 11 g

• • • • 3 • 3 • • 3

2 1 , 2 1 2 1

1 , , 2 1 , , 2 2 , 1

2 1 1 1 2 1 2

1 1

15 13 ,. 15 12 ,. ,2 1 1 1 1 2 1 1 3 1

2 1 1 , 2 1 1 2 1 1

5 • 4 4 4 4 3 8 S S 5 S 5 5 $ 5 3 5 5 8 6 8 6 8 6 4 6 8

9 9 S 9 g g 7 3 9 9

9 13 6 ,, 8

3 5 5 4 5 $ S 5 4 5 5 S S S 5 3 5 S $ 6 S S 6 e 6 6 4 S 8

& 6 5 e e 8 6 • 6 6

5 S 6 6 S 4 6 6 6 4 3 • 4 4 7 3 4 4 4

. 6 S e 6 6 6 6 ! 6 6 7 7 7 7 7 7 7 4 76 $ 6 6

• • 3 e 8 5 4 6 • 4 • 4 4 4 •' • o S 8 ' o 4 ' ' 6 S S

. . J . 4 3 3 4 3

S 8 6 S • S 6

4

, 10 , 8 3 10 e S 10 10 9 3 10 10 9 9 g 9

• • • • • 4 • 3 4 4

e 6 • 6 6 8 6 • 5 6

$ 5 5 • S S 4 5 S 5 8 ' o 5 5 o ' 3 S ' S S 4 S S S S 3 • 5

4 • 4 • • • • 3 4

• 3 3 • 4 • 4 3 4 3 7 7 7 7 7 7 4 7 7 8 6 7 a 8 S 8

8 8 8

9 7 1 a 8 • ' 4 8 7

,7 8 8 g a 9 • 8 8

4 . • 4 $ S 3 4 S

7 6 7 8 9 8 3 7 e  

 6 8 8 7 S 4 6 S 9 13 t3 12 9 g 5 12 11 9 9 a 9 9 9 g

10 13 11 .. 8 S

7 7 7 7 7 7 e • 7 7

 

8 6 6 e 6 6 4 8 6 3 3 3 3 3 3 4

12 11 8 12 4 10 9

3

, g 6 6 6 6 a 6 6 3 S e

 

16 .g 1

ISO 2 086

,2 3  

230 1 046 2

133 o 47, 1 80 1 2 086 1

133 1 1 0 47 1 38 t 048 1 130 1 O.&e 1, ,, 922 10 9 262 9 40 10 4 120 10 44 10 12 306 1030 9 8 1 986 13 6 2 70 867 9 8 221 1050 11 6 220 980 204 10 8

1180 , 10 262

10 30 2 24, 1120 12 7 209

1067 12 10 294 850 9 4 120 880 4 120 9 390 • 1 0301070 ,

1 7 6 3

1050 10 248, 1070 12 7 210  

640 9 4 120 370 4 1 0 40 3 70 4 2 08• 130 1 1 0 46 140 1 1 0 49 140 1 2 066 130 1 1 0<8

130 , 1 046

433 • 3 082 480 5 2 O&O !>78 S 2 083

780 9 e o 

4 80 , 480 S 2 oseo 6 2 080 890 10 7 202 S 70 S 2 084 !J 70 5 2 064 544 S 2 058410 • 1()4

790 8 070 3 4 00 4 o 000 $90 S 1 030 870 1 l 090 $&0 2 oee 8 390 1 030 • 490 ' : 0 34

370 4 2 084 seo 6 2 066 730 8 6 , 79 620 3 ove 7

3 80

.380 4 1 0 49

770 8 3 090

2 080 625 2 73 6$80 ,7 199

9

3il0 • 0 03480 5 3 o 75 S60 6 0802

480 040S 490 ' 3 070

, 490 S 3 083

030 710 8 145 460 2 066 S 380 2 060 • 390 • 1 030

380 4 1 0 49 610 7 4 1 22 860 7 3 092 760 8 3 092 710 7 S n o

767 8 S .., 87 7 e 56 e 1 57

0 10 \85 7

570 1 27' 4 4 n 4 063' 880 7 2 066

1033 ¡¡ 8 '5 833 9 4 1:!5 880 7 3 092 $50 6 156 S 1138 10 o 138 S 70 e 2 084 313 3 1 033 570 8 3 090 956 12 8 236

57

, e 67 6 3 10$ 0 046 e 1

10 78 2 083•se 820 8 6

860 3 092 1

 9

 '0 a

 10 10 9 5 4   a

1 1 o e 8 8 5 • 5 4

93 3 ' •22 3 5

7 7 7 7 7 7 6 $ 7 e93 3 10 41 2 3

 6 • 7 e 6 2 6 7

 10 g 12 12

  2 1 1 1 1 2 1 2 130 13 438 3 9 10 9 1000

93 1 14 435 3 11 93 15 438 3

10 13 11 1 10 10 3

 93

 1  

19  

41 4  

3

13 12  

0 13 9 9 7 10  

9 6 8 87  

9 10

l'oo<way

, ,

 2 34

S S 5   040

 12 , 11

 12 12 12 10 8 10

 

     9 12

• • • 8 8

.

, 43

7 6

4 4 3 • 4 4 4 2 • 4

4 •

-

•3

 Table 2.1. SOUTHERN AREA. MACKEREL lrom Olv. Vlllc •IXa an<llrom Fbruary • AprU an<l Sept mr.

lite<\ S<a!o...S Eog..nd Sc>( ) W..al'd So( Pott •• Defi'N!.rX Rln•·a IEO v92 , 2 1 2

 

    

1 046 2 093

 

     

12

6

3 9 12 13 12

  11

,   

    

93 246 3

9 9 g 9 S 6 9 9 e 9

• 3 44 4

2 1

• 2 4 4 •

 

 1 91 1266 & 19 1$ 6

93 1 3' 1 3

 ..

 9

 9 11 7 10 10 4 & 10 ;

8 5 $ 5 5 & e 4 6 8

4 4 390 0304 4 4 4 4 3 6 8 8 8 8 9 & 8• • • • 4

, • • 4 4

  

4 4 4 • • • 2 • 380 2 08 • 0

 8 8 7 8 8 87 7 S 7 8 6 4 7 6 8 5 e & 8 8

8 8 2 8

93 387 6 8 5 :1 2

4 4 3 • 4 o 3 4 3

   

113 2 38 238

93 2 :l 2 S 5 S 490

9

.., eS

5 5

5 4 S e e 3 5 5

5 4 5 6 S 3 5 5

5 5

S S $ 5 5 5 4 8 7 7 6 8 7 7 3 8 a

S ,

• • 4 4 4 • • 2 4 •  

93 2 401 2 4& 6 e S 1 7 1 e 3 1 7

 6

 113 3 3

  

S 75 083 6 3

Page 21: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

3 3AG 3 3 27 41 1 3 3 430 3 3 2G 4.. 3 3 30 44 1 3 3 31 396 3 3 32 415 3 3 33 37' 3 3 3A 345 3 3 35 41 3 3 3 :le 401 3 3 37 43 4 3 3 38 391 3 3 396 338

3 409 3., 3 40 345 3  

3 42 423 3 3 445 3 43 3 •• 400 3

• 4 3 41 0 4 4 4 360 4 4 5 360 4 4 e 360 • 4 8 370 4 4 9 37.0 4 4 10 370 4 4 11 420 • 4 13 360 • 4 15 330 4 4 16 390 4 4 17 360 4

18 41 0 4 4 IV 330 • 4 21 350 • 4 22 350 4 4 2'3 36 0 4 4 ?4 400 4 4 :t5 370 4 4 28 350 4 4 27 34 0 4 4 28 310 4 4 29 360 4 4 30 34 0 4 4 31 24 0 4 4 32 390 4

• 33 390 4

4 36 36 0 4 4 37 34 0 4 4 36 340 4

4 39 390 4 4 350 40

C1

. 330

• 4 350 •

. 4 4

340

4 4 <5 5 '!.80 Q

5 2 180 ll 5 3 180 V S 4 Q 18 0 5 5 le O 9 S 6 190 9 S 7 190 V S 8 190 Q 5 g 190 Q

S 190 Q 10

5 .. 200 Q

5 ll 200 p

 S 1S 20 0 S 21 2'30 9 S 22 2'30 g S 23 23.0 9 S 24 230 9 5 30 9

.<,., 27 2.4,.0, 25 2

.Q,.,' .,.,

4 5 • 4 4 • • 3 • 4

10 g g 10 10 9 8 S 10 8

o e 6 e e 6 3 5 7 6 a a 7 7 g a 6 3 a 7 3 3 3 3 4 3 3 34 4 4 4 • 4 4 2 • 4

e 6 7 5 7 6 5 • 7 5

6 4 6 6 5 4 6 6 5 7 6 6 8 e 8 4 6 e 6

• 3 2 3 4 3 3 2 4 3

15 15 10 15 12 11 • 10 ,8 10 7 8 10 8 1! 4 7 8  

4 • 3 4 4 • 4 3 • 4

4 3 4 4 4 4 3 4 4 9 9 9 9 9 9 9 9 9 6 6 6 6 e 8 5 3 6 e 8 5 6 • • 5 7 6 • 5 6

7 e 6 6 7

o o 5 e e e 4 6 e e 5 4 4 5 5 5 • 3 4 5

8 11 5 11 9 9 4 7 8 7 6 5 7 7 6 7 4 7 7 e 7 3 6 6e 6 5 6 e e 6 • e e

5 6 6 6 8

1 7 7 e 7 e 4 7 7 8 7 9 7 6 7 e S 4 8 S 7 9 Q 9 V V 8 9 Q

5 S 4 ' 4 4 • 3 • • 4 4 3 • 4 • 3 4 4

4

e S S "e . • 4 4 4 4 4 4 3 4 4

7 7 • 6 e 8 5 3 7

3 2 • 4 4 5 4 3 • 4

3 1 1 3 2 1 1

e 1 e 8 e e 5 4 e 6 a 7 9 7 8 9 7

3 " 2 6 e 8 e 5 4 3 • • • 3 4 • 4

e '3 e

• 4 • 4 • 4 2 • 6 e e 6 e e

e . . 7 7 7 e ¡ e 5

 7 6 5 8 a e 5 3 S 5 5 4 3 4 3 3 4 4 4 4 4 4 4 • • 4 4 2 4 4

o o o o oo o o 1 o o o o o o o o o , o o o o o o o o

o o '1 o 1 o o o o o o o o o o o o o 1 o o o o o o o o o 1 o o o o o o r o o 1 o o o o o o o o o o o o o o o o o 1

o o 1 n n " n o n o , " o o o o o o o o o o o o o o o 1 o

o o o o o o o '1 o 1 o o

o o o o o o o 1 o o o o o o o o o 1 o o o o o o o o o 1 o o o o o o o o o ' o

63

•oo 4 045 2

880 •5 1 47

10 8

1250 ,. 7 •• 1

570 &10 215

6 • 100

313 3 1 033 380 4 0 2 085 11 5 • 120

933 7 211 12 540 e 080 2 830 6 • 110

7 00 7 • 110

11 44 11 330 15590 e 1 030 380 4 1 0 40 a89 g 1 031 380 4 0 40 1 900 9 o 000 560 6 3 092 e13 2 080 e 513 • 3 105

570 6 2 064 •40 5 2 086 5:1$ 2 066 5 758 e 7 2 41 630 7 3 100570 e • 100

e80 7 • 10<

8 6' g 2 067 410 4 2 054 580 8 2 080 380 4 1 0 40 460 S 2 oso 500 S e 224 514 5 2 084 300 4 1 030 300 4 1 030 233 2 2 067 S 67 7 4 133 3 75 4 1 0 43 1 89 2 087 1 4 00 • 2 0 50

560 e 3 o 75 a::s 7 4 130 483 4 167 6 390 • 2 054

57e 43 2 073

0 6 3 090 7

380 • 2 080 470 3 078 5 570 e 5 1 27 380 2 060 4

380 • 2 080

011 o 031 1 0 00 o o 000 020 1 040 o 020 o 1 0 40 010 o 1 030 o 10 o 1 030 o 10 o 1 030 o 10 o 1 030 o 10 o 1 030 ....,.., ' .... n 020 1 040 o 010 o 1 030 010 1 030 o 020 o 1 040 0 11 o 1 031 010 1 030 o 010 o 1 030 o 10 1 0 30 o 010 o 1 030

otartad TOla1 NOTre•ó

Em.o.!yf'OW'fo A9reement abovt aver•ot'?

158 HO 1... 188 ""' 11' la> 180 o 18 1 :t5 S o 17 3 8 8 ,¡ 12 12 12 ,: 12 12 17 17

y y N y y y -':' N N y

 

 a3 93 n n n 93

 93 113 93 113 113 n 93 93 93 113 113 113

 7 9 7

,. ,. 13 13 15 13 12 5 ,. 12

e 8 ll 8 5 8

  5 S 5 5 6 S 2 6 o 12 11 9 12 9 1(; 5 8 8

 8 8 7 7 8 7 • • 7 7

e e 6 6 8 8 6 5 6 8

 

   710 a 8 158

 sao 6 3 098  

 310 3 2 070

780 8 8 180

113 93 93

n 93 113

1 40 0 4 g 2 360 4

ll ll ll 9 9 a g g

93 113 93 113 93 a3 113

4 7 370 4 5 5 4 5 5 5 5 3 5 S 470 5 2 084

 8 5 4 5 5 6 8 S

4 12 400 4

113 93 93 93 93 93 113 93 113 93

 113

4 14 370 4

 

  4 20 34 0 4 e 8 e 5 6 8 6 4 8 6

5 S 5 5 S S S 3 5 5

570 6 2 084

 64() 7 S , 4.l

113 Q 5 S 5 5• • • 4 4 4 • 3 4 4

3 S

113 93 113 93 93 113 113 93 113 113 93 113

     

34 350 4 4 35 40 0 4

4 2 2 2 3 2 2 2 2

3 • 4 4 4 • 3 •  4 5 5 5 5 S 4 2 5 5 4 50 5 3 092

93 113 113 93

4 4 • 3 4 4 • • 4 370 • 2 0114

93 113 113 4 93 113 113 113 93 113 113 93 93 93 113 113

 

93

2 360 43 .,o

5 5 4 5 5 5 4 3 S 6

 o e o o o o e o : o 010 o 1 030

113  

93 93 93 93 113 113 113 '.),.?,

5 13 200 Q o o o o o o o 1 o a

 

  

o o o o o o o o ' o

" 2 77

Page 22: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

hble 2.2. SOUTHERN AREA. CumulatJVe¡¡9e compositlon byre¡¡der tn percent¡¡ge. Cumulative age composition of all and beslreaders obtained from the modes.

 

,,

- - -

-¡m- ''• """ S<o<loMS E.""'and "<xwoy --sp¡!EOJ kt aOd S¡>("Zll) """"l¡o ' 'lus ALL BEST

ft :1,., 11% 1:1% 11% ""' ""' m. 11% 11'1 11..

o Re-,_,1 Reo2 q;uce3 Reade-; RMdet 5 R& Re7 :te8 Ruder 9 RtaderiC re.a<Sers reade

3

-

1 ,,. 16'11. -18%

 

3&'10

 

3&..

1 .. ,,  

- 4

 74.. 73!1. 80'!<. 711%

e

38% 38% 3Q'I 3·

&%

 

70'11> 72% 74'tt 100!0 73% 74!1. 74'1. 7214.

8

  

 

7ft 76'11. 84% 85!1. 77% 77'11. 8214. 100% 80!0 84'11.

78..

.,..,. 20'10

- 18'

16'l<o 18'1 15%

- 88'1.

·  

Go% 83%

- SI% 4ft 47%

91'1 100!0 85'1. 9111. BN

2 18% 18'10 111% 2:1% 20% 18% 21% 28% 17'10 17'10 1ft

21'11 20'10 21'11. 25'0 20'10 20'10 14 Sé'IO 19% 23% 21.. 21'1 3&'10

3&'10 5

54% 56'11.

 

Page 23: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

11 94'10 -93' 97!1. 96, 111!1.

100'1. 90% 93'!1. 91'!1.

96!1. 100!0 95'1> 98'!; 95'1> 94!1.

-e es-.. 7

7Ci'!E> 72'11> 8:1% 70% e.-.. e7% 8814. 8514.

100!0 iN -

100% 1 100!0

- 100'11>

- - 1 1- 1-00'11>

ea-.

•-om. 100% - - 9

9111. """

10

 1,.3 96%

9 94% 87'!1. 91'!1.

 100!0 8 100'11>

100!0 100!0 96..

12 97'11. 95!> se'

· 95%

100!0 98!1. 100!0 98'1> 97-..

100!0 15

l()i)lo 100'11> 100% 100% 100'11> 100'11> 100'11> 100%

18 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100% 100!1. 100!1. 100'11> 10()% 17 100!0 100% 100% 100% 18 100'11> 100'11> 100!0 100!0

100'11> 100!0

100'11> 100!0 100!0 100!0 lOO.. 100!0 100!0 100!0 100% 10()!0 100!1. 100'11> 100% 100% 19 100'11> 100.. 100!0 100!0 10()!0 100'11> 100.. "00'11> 100!0 100'11> 100'1. 100'11> 20 100'1. 100'11> 100'1. 100!1. 100'1. 100'11> 100'1. 100% 100'11> 100'11> 100'11> lOO% 21 100'1. 100!1. 100% 100'16 100'11> 100!1. 100'11> 100'11> 100!0 100'16 100% 100'11> 22 100'11> 100'11> 100'11> 100!0 100'11> 100!0 100'11> 100!0 100'11> 100!0 100!0 100'11> 23 100'11> 100'11> 100!1. 100'11> 100ll 100'11> 100'11> 100'11> 100!0 100'11> 100'11> 100'11> 24 •oc¡,., 100!1. 100!0 100'!1. 100'11> 100'11> '00!0 100'11> I()Oll 100!1. 100% 100!0 25 100'\ 1 10()!0 1 100!0 100'0 100'\ 100'11> 100'1. 100'0 100!0 100'0 20 100% 100'0 100'11> 100'11> 100'11> 100!0 1()()% 100!0 100'11> 100!0 1()()% 100!0 27 100!0 100'11 100'11 10()% 100'11> 100'11> 100'11> 100% 100'11> 100'11> 100% 100'11> 28 100'11> 100'!1. 100% 100'11> 100'11> ·om. l X'l< 100'11> 100'11> 100'1. 100'11> 100'11> 29 100'!1. 100'11> 100'11> 100!1. 100!1. 100'11> 100'!1. 100'1> 1()0% 100'11> 100!1. 100'11> 30 100'11> 100'11> 100'11> 100!0 100'11> 100'11> 100'11> 100!0 100!1. 100'11> 100'11> 100'11> 31 100!1. 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100!1. 32 100'11> 100'11> 100!1. 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 100'11> 33 100'0> 100'11 100!1. 100'11> 100'0> lOO% 100"< 100!1. 100'1> 100!0 100% 100'11> 34 100'!1. 100!0 100!0 100'11> 100'0 100'!1. 10()!0 ICIO!ó lOO% lOO% 100'!1. 100'11> 35 1000. ·ooo. 100% lOO% 100'0 100'10 1000. lOO% 100'11> 100'11> lOO% 100'11> 38 10Cl!ó 100'11> 100'11> 100% 100'11> lOO% 100% 100!1. 100'11> 100'11> 100% 100% 37 100'11> 100'11> 100'16 100'11 100% 100'11> 100'11> 100!0 100!0 100!0 100% 10011.

-

38 100'11> 100'11> 100'11> 100'11> 100!0 100!0 10()!0 1Ci0'<, 100'11> 100!0 100'11> 1000. 39 100'1> 100'11> 100'11> 100% 100'11> 100'11> !()()lo, 100'11> 100!0 100!0 100'11> 1000. 40 100'11> 100!0 1 100% 100% 1000. ICIO'JO 10C'!O 10Q!I. 100'11> 100'11> 100'1.

 

Tabel 2. 3. SOVTHERN AREA. Ag,.composltion by age re.ockr (on numbor5 of otolltha re>od Age composition ol all and bes!ruders obt.inood from !he modu

.  

NO!herl Scotlo•d !;•glano NO<WOy Sp(IEO¡ ,.,...,., SP{AZTI¡ 0 1\ gel Denmark ll ST

Ruwo Arm- RuMt 1 Quder 2 R:udet 3 Ruder 4 "-•S R.e6 R7 Qt.ac.r 8 Ae9 Re..,.,·c , 9E 

.. ...

 

Page 24: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

,.

3

•2 13 2 3 3 ,. 5 17 12 7

• •

11 10 9 ·e 13 11 6 11 23 16 3 13 13 15

,7 17 16 18 11 15 6 16 13 18 12

o ·e 16 20 20 20 -.o- 20 "9 20 20 20

2

1 • 9 13 13 •3 12 12 20 g g

7 4 15 5 1 7 52 12 4

• 31 3' 36 29 !13 2& 41 34 211 33 32

.5 e 32 2• 18 29 28 38 20 3 31 30 31 SS

111 19 27 18 16 15 2& 20 18 23 20 16

11 6 S 9 10 12 a 13 17 10

10 7 5 • 8 9 8 10 3 8 9 11 5 11 8 2 7 S 8 9 7 7 12 5 3 5 7 6 5 13 2 2 3 3 14 2 1 15 2 2 •6 17 18 111 20 2" 22 23 2• 25 211 27 28 29 30 31 32 33 34 35 3& 37 38

40 3Q

'&' .,o  _ ló >U ""

 

Page 25: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

f.2 f. 1 1es 24 2 24 1 20 5

Tab2l.4. SOUTHERN AREA. 1\!Nn n¡¡lh atby otolilh rtadet (wilhout C()(lfion 101 meaaured below).

N•:t>er ,..,..,.., tn;ond Nc>v..

f. f. f.

. .y Sp(IEOJ 11e,anó P("", O!'!IJ9•1 l..l':nmatk ' "'... •Ll

rm- Réad•l 1 Rto der 2 Ruder 3 Reader .t Re5 Rudef & Re;,:S.r 7 e:r 8 Reader 9 Readet 10 feJÓf'f'S .. o 203 202 202 202

2 242 266 353 289 2Q2 20 28 . 357 253 31 o 31o 26• 3 339 376 35 o 367 318 316 382 37 3 33 7 4 362 358 365 355 357 358 382 3$7 358

332 37 . 374 359 358 357

S 357 363 383 369 386 386 37 42 3 373 7 S 370 86 8

7 400 400 39 e 3i 1 398 385 ,.,. 438 399

391 394 6 388 38& 391 391 382 389 3$2 433 38 4 386 3$1 3&8

... .. 391

8 3e • 380 .,3 40 4 399 401 406 403 407 403 396

10 434 423 42. 416 41 o 438 , 426 413

9 41' 41 o 415 415 41 3 41. 420 41 o 42 o 413 41 1

, 431 4Z4. 443 434 · S 429 432 ..., 431 429 429

.42

.9

13 ••o 436 43 3 448 435 ••1 445 .... 435

12 32 430 43 8 433 43 3 42 7 431 2 14 435 44 8 4.<8 43 4 445 44 1 441 , 435 uo 433 43

15 43 5 42 3 44 9 435 435 16 17 18 19 20 21 22 23 24 2S 26 27 28 2Q 30 31 32 35 34 3S 38 37 38 39 40

                                  

 

Page 26: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

Page 27: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

0

=6 'le-IC

Rué01! .....

33'1; 25'1; 25'1; '"' ........

Reoder 1 10% 17'*> 6% 10'!. 8% 1.6.% 64'!1. ·, ""

.. ,

=y· ==:rA deviation

o,.

,,.-;jseR.e8"'!'..r 4 6 O liO 02& 4 3 0.5'1 051

 Table 2.6. UTHERN AREA.Agreement {'!.) ool tolith readSO

'!·-··

=··::!E.,

ings.

7'JI,

 

......,." ·"" r'and Reader 1 R adef 1 Ruder 3 ;:::·::: · _R.,...

47% <511. !S< 46 11'!1. 61... 57'!1. lmó

Re-¡arz $-4'1; 7:1% &N Re er l .. "l!% '!! 1811. 43% 1'!1

66 >4'\ 18'1; 55 !)4'1;

Rtao.r 41 81% 73 6 2:1% Sol'!; 78'1; Reader 52'!!> IV.. es" S< 58R..o.r! 1211. > 6211.

% N

R-7 22'iO 400r. 154'1; 21'1; s -. Reade·9

Reader 10

  

Table 2.6. SOUTHERN AREA. Oiugreement ol one year {%) be!Ween otollth readlngs. Thls eoukl be due to mlslnt•rpre ll on olthe hyalinelopaque edgo of thc> otomn¡.

 

Nethert tland Engrono_ NO!WOy _So(IEO) lle!and C>(Allli Potb,gal nma:r ..RUstla k Reader 1 R•r 2 Reaoer ) Rtae.r R•aGt r S Rea.tl..r e Ruder 7 Re.adef 8 Rea.ser a RHriO

R...-,.,1 28'1; 38'!1. 2ft 2<'1; 3<'1; '1; 26'1; 30'1;

25

..2

Rea 33% 2N 311% . 31 R:eader • 25% :l9'lL 1'1; 30'll. 1$% Ruder5 27'1; 24,. 18'1; :l9'lL 18% RN<I tt 313% 26'1; 36% 2v.. Reo<ler 22'iO .&5% 23'!1. RO..SOI8 3-111 18% R. de,g 38'!.

Reade1  

  

T; ble 2.7. SOUTHERN AREA. Oi sagreement of IWo or more years (%) be!Ween otoilth readings.

>:e:nen :.and 5n;1and Nolway P{IEO) trela nd ?<'•AZTil ."'O<WQ• uenmarl< _ R\KSti

'a er 7 1

N 9'11> .,.,. $3.... lO% Reader 1 Rtade• 2 Ru•db! 3 Reao«4 Rf' er 5 Reade16 Rf Reader 6 Read&t 9 Re•der 0

'""•N 12'!1. R.e•de·

Re-3 IV.. 13'11. RUt'et N ..... V.. 57'!1. .,.,.

... Ro>der!

18'1< $3'!. 13% 17.'1.; RH6et6 11% 81'!1. 7% V.. RNCI-•1 7 Reac:sere 15%

52'!!> 61.. Ru<*l V..

Rooá••1C

  

Tabl2.8. SOUTHERN A.REA.Slope lrom funetlonalllnear regr ssions ol age determinatíons (shooldbe elose to 1). lrom 1indieates that eertain age groups are aged older or younger.

v..,. _.,ot1 S<cGind En¡ono ke _ ....-.. X-UJS Reader 1 Ruder 2 eader 3 Rt-aclet 8 Ro or 10 od

RNd 11 I()Q o l OS 1 12 097 Oll3 o • o 94o 098 103 Oal 086 043 o 87 086

Rudet 092 090 Aeader 10 7 Ruo.,• o 1()2 o 1 • 095 09• 9 092 093

4 1 f2 l OS 11 101 OP7 050 099 097

0 •Rooo.,5 089 097 086 0 98 086 Ca3 o <2 O&o< 084 R.-ra 103 112 099 lOS 1 18 095 046 0 98 097 U!Mf7 1 07 , •e 103 1 10 120 OSI 100 104 lo:'

Reaaer & 213 232 202 203 2 37 207 203 8 196 19RtaotrO 105 1 18 , 01 1 08 1 19 102 cge O<IQ o

Re.tder 10 l()e , 16 103 107 1 19 103 1()() 050 101

  

Table 2.9. SOUTHERN AREA.lnt rcept lrom fur.,tional linear regressions olage determinations (st¡ould be elose to 0). A c:levlatlon lrom O lndlcatca consiSten! mlsreac:l1"1j ol1 01 more years or that cer taln / age groups are aged okler 01' ycuoger.

 v.an .";f$,...., Soocand Engaw No<w2y SO(>EOJ "eoanc SI>(Ar.Jl ,. ..,,. - R• u •

X-ax¡a C(uOer 1 -R 6er 2 qeadei' 3 Reade'4 R...S• S Rl!a ·6 Reader i Re.ao•r 8 Re..adeo·9 Rea · de"

uoer1 .0 43 ........... . 1 .090 .0 .o 13 0:;.4 0 <3 .o 02 () 4

Rnder 9 .o 12 .059 .QOG 050 020 OSI 080 039 03Rr 3 o 42 013 -<U8 .005 054 0 7 04Q CS1 045

08 81 Ru S 7 OOQ 004 -0 54 03' o C al! o c.:

8o 4

Q-.é .o 14 .056 .Q$3 -087 .C5 .026 028 029 11 .0RNdt•7 ..0 2A .028 .0 63 .037 0 27 030 054 o 15 014 R•ec:s.-' a .on ., 17 .0 -103 127 .058 .o 70 -028 .060 R••d•t9 .o 45 .0 2 .083 ·1 20 ·1 02 ·030 .0 OIC .04()

RooM•10 002 .e •s .0 47 .0 66 .o 53 011 .o 15 030 04()

.001

.081 .06.o 16

1

 

Page 28: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

s.ze "'•"'" 050 0.2 9-11

 1 25 100 1 7 150 225 200 2 75 250 325 300 3 7S 350 4 25 4 00 4 75 4 50 525 500 5 75 5 50

1 1 1 o g

6 g o • 1S 6 o ()

:¡ 30 3 o o

o 1 1 8 • o o , o o o 1

o o o o o o o o o o o o o o o o o o o o o o o o

 

36 5 57 28 8 2

S4:t tte"\\'a

050AGE·; AGE · e ().2 4 1!>•

125 1 00 175 1SO 225 2 00 275 2 50 325 300 3 75 350 4 25 •oc 475 450 525 6 00 5 7S 550

16 28 24 7 1  

S 1 12 10 2 o o o 6 S 2 o o o o • 2 1 o o 1 o o 1 o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o o

 

36 52 58 31 9 188 1

·Table 2.10. SOUTHERN AREA. Frequency distribution

ol STDEV lrom al!readera.  

•Gc ';: AG• 4

Table 2.11. SOUTHERN AREA. Frequency diatrlbution o!STDEV from t: .e readera with an agre-ement bove average.

:  :19

 22  

 1 o  

 1S 23

 1S

 S

 2

o o  

8 •  

1  

1        3 oo o o 4        

 

Page 29: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

¡

20 " ' 33 20 ,.2 \8 10 13 a 1 7 o ' 1 1 2

!,,

. • •• • "· t

..

  

  

MACKEREL (Div.VIllc and IXa) EXCHANGE SA. .PLE Fit readings. MACKEREL (Div.VIllc and IXa) EXCHANGE SA. .PLE Fit readings.   

T;obsa1...,1:Sy modal age ·prese-nled ltl t age racorded, the stdev and number af T;obsa1...,1:Sy modal age ·prese-nled ltl t age racorded, the stdev and number af t otwugagor inga by readef ancs «al! rert..

t otwugagor inga by readef ancs «al! rert..

  REAOER 1REAOER 1

fr 1 AG« -orded o,2t> 2.lod:ll · O 1 1

01

2't o2 o;e: ld

...... 1 o , S , , 2 A¡jt l'ttcrdtd 0,00 ' ta TóO lOO .00 U3 ' 1 .1! . 14.00 J 1•.00

 

1 • ! • 1 10 1 H 1 12 u 14 15

1 7'llt<lw

1 ' 0 1 1 ·" U2

000 o,at 4.00.J, o.e2 1.2( 1 2.14 Xca l2 • ;: 10,&& 2H 3,20 ! ua  

OO n

1 11 1 11 1 ' .2•

'2 11 26 ..JO. _ e 1 12 1 ,  _!_ 2.8'3

o :¡--¡21 S O O

 Modlll a;• r·-e-·--r· ' -t  

' 'l ¡ • • 7

• 1

e +-...1L 11 1 12 14 1.14 ' 2,50 1 2,75 • 3,70 .SS 5.47 ' O,:la 7,40 e.oo a.oo ·an· 10.50 7oo 1300 • 1200

  '

 

15-.

A;• ,.conltcl : .'..O.. !_ 1,41 1,00 O.t 101 1,40 2. · S,:\8 ) 0 •&a l J ' 5 88 • 2'1Y 1,40 ,7 ol!.

n :10 1.4 2 20 10 _ _.10 1 1 a 7 e '1 1 1 1 2

 

110<1' >ge 1 e 1 1 -·¡. 4 l S 1 7 • g 1 10 11 12 13

2 -;"-1

91'ft(>'Cf tcl 0.00 107 i 3.oo 1 4.oo toe 1 o.oo 1 1.00 &.00 ; $,:!0 10.00 ! 11,80 12.001 13"00

 16 J.

oo  

o.sa 0. 0.00.1 052 _· O (l.J O1. 00 1 !!)...!... ' .00 1 " '7 14 , 4 31 H 27 1& e ' 10 r s 1 s l 1 o 1 o

1S1 ..1J S!!.. 1 0 00 _1 . .

  REAOER i

MH:JI If t 1 1 2 S 1 4 -.IJT1-

7,),0

 

. - a· - · tT'"1' Q-·--¡-;-r 12 !J 1 14 11

.t.Q•=omed 0.00 1 IOTI 200 2 2& e. 13 8,40 DS5 j tt .2' 11,71 12,00 18.00 1600 16.00 3,eo_l 4.1c 0, .0v 0 o 53 1 0. 00 -;-:¡r- 1,08 1 50 . 1 40 z n 4

• ,,...

'

••

1 111 31 21 1, . - -

lh4o1•t• o 4 . -- -

, READERi

n 14 2 · 2 1& ·!--. ....!2._ _E_,_---_7_1 0 1 1 2  

2

 

• 7 • , .. 10 · 11 12 u j 14 \&

Allt OMIM 000 1 207 2,!0 BO 545 t 10 ,00 l.lO t,OO 10))() " ·" 11 .00noo 12.00 4,08 0,00 O,!l: '·" 1.1 o.n 1.1!1 \,17 I ,H ' 1,\4 000 2.14 l.ll :Z.It 1 000

2"ttdw

" """'--'--

 Modal •

Ao• ...:>,f>.N,i.t6a.0.00 2.00

 

o 1 1

6 • 1 7 1 • 10 ., 1 12 '  u te 10

1,14 D.QO o ,, 00 1-1 4 1,\1 \,78 . \.83 1, 1.32 1 1 2 2 4,24 .

' 4 ,00 4,10 1 5,&.3 ; 1.00 l,lO l .Y j 12S U' j iO I' 10 00 12.00 1:?50

0 0 ,77 ,43 1 ,3ol n ,, 1 2 l 1 )\ 20 2t 10 \:1 • ' 1 • 1 1 2 J

REAOER 8

M o,d. l· 1 1 1

'¡ o 1 · o 1 oo 1 200 2.50 1 2.

uo_

S ! 4 • • J J • Q _1 10 H u --" " = "":

10•.77 .17

ue- '\ C': 1 f", :o 1 í , l ó ¡ u-,w.w. l i'4 1,:12

. 4,50 1 4.28 4.00 1 4.!7 f.""' = •.loi'W .

Mt t-I'Ct41 U7'1 1 !tll ,54 1to 1,41 2.0'

n )O H 2 • ! :. 30 1 H 10 ! 10 7 6 o 1 1 \ 1  

RE.ADE.R 11  lf od:ll :oge D 1 1 2 j 1 ' f 7 2 - !..!.L.!

• • 10 H 1 !_1

1 ,00 1116t 2C.0r0ll6d 2.00 1 3,25 •.og l. •GS e 7 ' ¡¡ s.& 1 ·u• 10.07 10.00 '-'

1) ,t30.1 000 1.00 o i .1t ou ISI e• . ,,,8 ; 2 127

9 ao o

 7'r

.

11_1!! 17 0 1 2 1.41

12 _.l_LL4 -J3;t_ 20 31 0 .l 13 6 7 e 1 1 -- 2- t e 1

¡ ·.1 4 1 -,-l-,-

,,- READSR10 -,--¡r 11

MO 1 4 00 !>,Ofl; 0,00 a6o 0.00 1"14 1 0.83 1.15 1 1.48 OG4

\ 1 1 10 o ' :l2

 AI.LREADERS

Md l oa o S a ' 1 ? 1 t J 10 H 12 'U J 0.12 1,4l ,l, a. AQ•,.,...,rdod

14 ua l.U 1 6.T. 861 '- •$4 .1 IO,:IA 10.39 G.88 12 1 2]

.. 0,88 \,I D uo 1,01 1.2.(: 1.n 2 04 ' 2.51 2.84 2 1

5,04 S.t\0 U,,71 Z'tldt 4,72 •.72 45

84 67 ) 10 n lt? 1:!0 1t le w ltC 1 2tl 163

 

Page 30: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

             

Table 2.13. Results of Wilcoxson's test of the sample of Divisions VIIIc and IXa.

  

- ** * -

READERS 2 3 4  

- ** ** - ** 1  

  ** - - - ** ** - ** 2  

    ** ** ** - ** ** ** 3  

      ** - ** ** ** ** 4  

        ** ** ** - ** 5  

          ** ** - ** 6  

            ** ** ** 7  

              ** ** 8  

                ** 9  

       10

5 6 7 8 9 10

   

A* : Possibility of bias

bsence of bias

** Certainty of bias

 

Page 31: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

o

  

        

1... 1...

  

F -Ru.der 1 -<>- Rc.ad<r 2 --R<.odor 3 --R<.odor 4 --!Woder S --Ru.der 6 kader 7 -> R<.odor a --Rc.ad<F -Ru.der 1 -<>- Rc.ad<r 2 --R<.odor 3 --R<.odor 4 --!Woder S --Ru.der 6 kader 7 -> R<.odor a --Rc.ad<r 9 -·-Rc.ad<r 10 -·-

 

/ .-•--•--x--x--x--x==v   

"  

                

x-----x,----+----- ----- ------------ ---- -------------------------- ------r----- ---- 10 " 12 13 ,. 15

AClE

 Figure 1.1. WI!STfRH AREA.CumiN11n• compoaltlon by age .eld.r In percenlage.

 

 --BEST READERS

 110

  

 so

          

Q

 

.X

o• •

•o o o

10 o

 X

 X

  

AClE

 

       

10 " 12 13

         

 14 15

 f12ure 1.2. W!:STfRN AAEA .Aoe composlllonInnumbers of olclllhs .eldby otclllh reld.r. Aoe ccmposiUon trom

"IM>st relde.s" 1& oblalned from leldel$ above av"*.l• agr-..en1($M Tat>r.5).

 

Page 32: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 E .---_..:R<.ode.::::;;.:;.'.:.

  

50.0

 <S.O

 

40.0  

3S.O  

 300

 250

 

 20.0

 

 1S.O

 

 10.0

 so

10 11 12 13 1. 00-- --------- ------------------T---------- ---------- ----------------------------

 

1S 

AOE

 Figure 1.:S. WESTEftN AAEA. M&an lengl! a1age by ololll1re3<19r (wtlhoulcorrectlon lor meaaurecl beloW).

 

Page 33: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

-AGE0-2 AGE0-2

FI\EOUI!HCY FRI:OVEIICY

30

20

10

.... 0.3 O.ll 1.$ 2.1 2.7 3.3 3.ll 4.5 5.1 5.7 u u 0.3 O.ll 1..5 2.1 2.7 3.3 3.11 4..5 5.1 5.7 1.3 u

SOEV SOEV

 AGE3-5 AGE 3-5

FRI:OVEHCY I'I\EOUEHCY

30

20

10

 0.3 0.8 1..5 2.1 2.7 3.3 3.8 4.5 5.1 5.7 f.3 u 0.3 0.8 1.5 2.1 2.7 3.3 3.8 45 5.1 5.7 8.3 u

SOEV SOEV

 AGE6-8 AGE 6-8

FIIEOUEHCY FIIEOUEHCY

40 2S

30 20

20 10

10 S  

0.3 0.8 15 2.1 2.7 3.3 3.8 4.5 5.1 5.7 13 u 0.3 O.ll 15 2.1 2.7 3.3 3.11 45 5.1 5.7 6.3 u

SOEV SOEV

 AGE9-11 AGE 9-11

FRI:OVEHCY FIIEOUEHCY

1 ..111.111....,

15

10

 0.3 0.8 15 2.1 2.7 3.3 3.ll 45 5.1 5.7 1.3 u 0.3 0.11 15 2.1 27 3.3 3.11 45 5.1 5.7 u u

SOEV SOEV

 

1 AGE 12-14 AGE 12-14

FIIEOVEHCY FIIEOUEHCY

10 8

 

· · .. .

-- ..

 

'!! •!. 

. 7 -- ..  4

2o

o. M

 

.. -· !.1 .3 3.8 <5 : 7 a.:: :.ll

SOEV SOEV

 AGE 15+ AGE 15+

FIIEOUENCY FIIEOVEHCY

   

o 0.3 0.8 15 2.1 2.7 3.3 3.ll 4.5 5.1 5.7 1.3 u 0.3 0.11 1 5 2.1 2.7 3.3 3.11 •.S 5.1 5.7 6 3 u

SOEV

  Agure1. 4. Agure1. 4.

  W!:STERN AAEA .Fr-.queny dls1rlbutl0ft W!:STERN AAEA .Fr-.queny dls1rlbutl0ft

  

Flgu15. . Flgu15. .   

WESTCRN AAEA.Frequey dl o.-lbuUon of STDEV WESTCRN AAEA.Frequey dl o.-lbuUon of STDEV

  of STDEV!10m all !10m lile readers wt1l'l an ag-.nent above av•·

 

Page 34: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

MACKEREL (Div. Vla and Vllbc) EXCHANGE SAMPLE Fi rst readings.

f 1 20 -

s

i 1

s 1 -l- t 1 1

u-'2 10 i 1l

-1 ' T T w--r-1 - 1 '2 10 1-.. . -I

S { 1 S -t-I---t--I Ag.: Age

o J 2 3 4 6 7 8 9 10 J J u 13 J4 15 o o J 2 - ; ;- 6 7 8 9 10 11 12 13

.. 20

1S "' -.

+'2 10 "oE S

o

   

r:t 5 6 7 8 9 JO 11 J2 13 14 15

 

  

 

 

: yrrl! . . Age 1

o 1 2 3 4 5 6 7 8 9 19 11 12 J3 14

     

Q\

  

 O 1 l 3 4 6 7 8 9 JO 11 12 J3 14 15

..

r:1 -

    

.. -s 20

 Age

 O 1 2 3 4 5 6 7 8 9 JO 11 12 13 14 15

 {;¡J

 

!

¡:: j ?t::' 1

- o 1 2 3 4 5 6 7 8 9 10 J 1 ll 13 14 15

"' lS

+ o '2 JO

!S

: -- -t--i ,r-t-+--+-o+--+e -+--t--1--t--1- -l - +-+-+-+-+-Age Q< O J 2 3 4 S 6 7 8 9 10 11 12 13 14 15 < o 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15

--- --------------------------- ------------------------------------ Figure Í - .- In above age bai s pol ts average age +/- 2stdev of each age reader ls plotted against modal age.

 

 

Modal age o_:_ 1 1 2 1 3 1 4 ; 5 1 o _._!.. I 1 J 9_ 10 1"112 - , 15 1

Age recon!t<l· · 2.01 2.98 .92 5.01 1 5.80 6.93 8 00 8:.6 9.48 10,37 11.38 11.96 12,22 12,433 1 2' tdev • ....._; .'13 ; 0,216 1 1 52._._!}2 _ 2.33 ' 3 .29 3,81 4,25'5,11 . 4,76 5,58 1 7 33 1

r .. -E 20 .,..

   

ál l ,.....- Ag.:

o + + .., + "' -1 -+ , -1--i

< Figure 1. .1- In above age bias plot average age +/- 2stdev of

all age readers is plotted agalnst modal age.

 

Page 35: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

 

o

¡

1

:•

.. _,

1

i

          

 

.o. <..•

"•' c. :z 1

"e• .•'. "c•. "o z

 

   

¡-

 

.....-·     1 1 ¡- i   

 i

           

 

- - -- - -

          

------------------

 

   

-      

-·-----         

-;N

--- <' ---

         

 5 '-"

 

::

 

------------------- _ .  

 

... .. N N

 ee11• ll:llSHiON

  

·<-  

  

l l.. ¡ ! 1 1

 1 1 1'---.

 +

... ... -- <.. •"•'

 

---------- ! y 1

 

 ---------------

   

J -

1 <..

'--

J 1

f-- :z 1

"e' !

•' .o. --

·---- ----:,r---------·----

"c• " oz

'   

 

'!__ ·-   --·

    

·--------- - :

1 g ...

+-r  

  

.. N

 

Page 36: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

olllf'liiJ.SJ.nos

 

Page 37: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

X-·

                            

AGE

 Figure 2.1. SOV1llfRN AREA. Cumulallve age compoalton by age reader In pen;enlage.

 

 • -1 o -2 • _, o -· • R-5 • -· • Ro.lor7 o R-$ " R-10 • , --BEST REAilERS

 

               

20 \:  

10  

 

• •

    

 o

 o

o •• g

0---- ------,------ ---- ------------ ---- ------•------ ------•------x------•------------x------ o 10

AGE

11 12 13 14 15

 Figure 2.2. SOUTl1fRH AREA. A,gcompo&ltlonIn numl>tft olotoll1hs read by ololl'll reader. AQe composl1lon trom

" tMoslrn d·ls ol>lalned flom ..adabove average ag<eemMI(see Table 5).

 

Page 38: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

J --- Rudcr 1 -<>- Rud<r 2 -+- R<.od<r 3 -<>- R<Odor 4 --Reodcr S --<> 6 -o- Rud<r 7 x-Rc.adc.r 8 -<>- R<..dcr 9 -x R<..dcr 10 -•-

               

 15.0

  100

  

5.0  

 0.0 ----- ----- ---- -----+-----+-------- ------ ----+--------------+---- ------ -

o 10 " 12 13 14 15

AGE

 Figure 2.3. SOUTHERN AREA. JManat age by 01oll1t1 reader (wt1hoU1conoectlon lof !lleUUred beloW).

 

Page 39: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

:

•.zs •. 0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 75 5.25 5.75

  

FREOUEHCY

 

AGE 0-2  AGE 0-2

    

0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.25 •.75 5.25 5.75

soev 0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.25 U5 5.25 5.75

  

FREOUEHCY

30

20

10

AGE3-5  FREOIJEHCY

AGE3-5

 0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.25 •.75 5.25 5.75

soev

 0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.zs •.75 5.25 5.75

soev

  

R

:\EOIJ

LEHCY

AGE6-8  

 F llfOUE)jCy

 15 10

AGE6-8

o ·-- ---- ----r- 5

0-- --4----- --+--+-- •.zs •.75 5.25 5.75

soev 0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75

soev   

R\EOIJEHCY

AGE 9-11 AGE 9-11

 

   

0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 U5 •.75 5.25 5.75

soev

0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.zs •.75 5.25 5.75

  

FIIEOVEHCY

AGE 12-14 AGE 12-14

 

 

o1 L · ,rn    soev

   soev

  

 FIIEOIJEHCY

 

0.05 -- ---- AGE 15+  

FIIEOVEHCY

AGE 15+

0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.zs 05 5.25 5.75

soev  

AgUr@ 2.4. SOUlliERN AAEA. Froequ.ncy dlalrlbulkln ot STDEV flom all "'l><lera.

0.25 0.75 1.25 1.75 2.25 2.75 3.25 3.75 •.zs U5 5.25 5.75

soev

 Flgu.- 2.5. SOUTHERN AREA. Freque.wy <la1rlbutlon of STDEV

trom lhe rea®rs wlth an a<oJ,.l.'!n«<l above average.

 

Page 40: Final Resul ts of the Mackerel Scomber scombrus , L ... Otolith...1 Working Document for the Workshop on Mackerel Otolith Reading. Vigo 8-14 February 1995 Final Resul ts of the MackerelScomber

¡:: j

1

t +

7

ACKEREL (Div.Vl!lc anIXa) EXCHANGE S LE First readings.

- "' lO

o s _ t.-r-t-

¡ "' o • - - -+---i----1--' . .

:/ O 1 2 3 4 6 7 8 9 lO 11 12 13

\----------------- 14 J

'E "' 15

20

+'2 10

 

o S

     

 O 1 2 3 4 S 6 7 8 9 10 11 12 13 14 I S

        

... -E 20 t "-

"'f

:'2t 150 +t

l:  

Q()  

   

O 1 2 3 4 S 6 7 8 9 10 11 12 13 14 IS

O 1 2 3 4 S 6 7 8 9 JO 11 12 13 14 IS [;iJ

r--------------------------------- -..--------------------------------4 20 -;:

¡::1 A

1 ge 1

o 1 2 3 4 S 6 7 8 9 10 IJ 12 13 14 15 O O 1 2 3 4 S 6 7 8 9 lO 11 12 13 14 15 l

-----------------------------------------------4

 

...  

20 -

IS ..!.

ll'l '2 1 1 [;IJ 1 ;) 1  

 

} -I-r

o.... ¡;¡;¡

- T

 

o ----,- -t-

-

Ag.: --+---- r tAge

1--o 1 2 3 4 s 6 7 8 9 JO 11 12 13 14 15 O 1 2 3 4 S 6 7 8 9 JO 11 12 13 14 15 1

--------------------- Fgi ure 'Z . b In above age bai s plots average age +/- 2stdev of each age reader ls plotted against modalage.

 Moclal o l 12 1 3 ; ......-. 5 e 1 1 1 11 1 !1i 3 _!_ 14-=rJ.[)

As¡e rtc<>r <le<l , 0.12 ¡ 1.Úi. 32 3 . 03 3. 89 89 _,_ !>.79 6,!>9 1 7!.>0 8.84 9, !>3 1 10,34 10,39 9.86 • !>O 1

1 1 2"stdtv1 0,86 '1,1 1 1,16 , 1,01 . 1,20 1,78 2.0.C 1 2,51 2 84 3. 45}4,72 -•7U 5, !>.80 5,87 1

 

"l1 r:o _t:_:

0

   

 

oe,--H-1

 

tlt11  . -+-;-- -'! - ,

:;: < o 1 2 3 4 5 6 7 8 9 1 11 12 13 14 15

 

Figure '2.1 In above age bias plot average age +/- 2stdev of all age readers ls plotted against modalage.