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  • 7/28/2019 Feeding Horses Student Version 2013

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    227.208 Equine Nutrition

    Erica Gee BVSc, PhD, Dip ACTRoom 1.21

    [email protected] 2013

    Acknowledgement: Dr Simone Hoskins

    Equine Nutrition Lectures

    Monday 29th April 11am SSLB6

    Thursday 2nd May 9am SSLB6

    Monday 6th May 11am SSLB6

    Powerpoints on Stream, plus required reading

    Who am I?

    Senior lecturer in Equine Science

    Equine nutrition and reproduction

    [email protected]

    Learning objectives

    Evaluate the diet of a horse

    Explain the main principles of feeding horses

    Discuss the benefits and potential problems of

    pasture based diets for horses

    Discuss the effects of nutrition on health and

    well-being of horses

    Lecture outline

    The basics of feeding horses (and why the horse isnot a cow!)

    The role of fibre

    Assessing fatness and adequacy of diet

    Pasture as a nutrient source

    Where problems occur & how to prevent them by

    feeding practice

    Why feeding is both art & science

    Prior knowledge assumptions

    Equine GI tract anatomy

    Equine GI tract physiology

    Nutrients & nutrient digestion

    Protein & amino acids

    Lipids

    Carbohydrates & fibre

    Vitamins & minerals

    Some knowledge of equine management, behaviour,breeds etc

    mailto:[email protected]:[email protected]:[email protected]:[email protected]
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    The digestive system

    Foregut

    Mouth

    Oesophagus

    Stomach

    Small intestine

    What is happening in the small

    intestine?

    Enzymes, bile, other digestive juices

    Feed passes through in 30-90 minutes

    Digestion

    Soluble carbohydrates (sugars and starch)

    glucose

    What enzyme is rate limiting for starch digestion?

    Most protein from grains

    Oils and fats

    Calcium, phosphorus, B vitamins, vitamins A, D, E, K

    The equine digestive system

    The hindgut

    Caecum

    Large colon

    Small colon

    Rectum

    What happens in the hindgut?

    Giant fermentation vat

    Bacteria break down fibre volatile fatty

    acids (VFA)

    B vitamins

    Fermentation takes days

    Quality

    Water reservoir

    Features of GI tract Lips, tongue, teeth

    High degree of selectivity

    Fibre grinders

    Stomach

    Small capacity, fast passage

    Cardiac sphincter = one way valve

    Normally rarely empty & highly buffered

    Small intestine

    Relatively fast passage

    Urea secretion terminal ileum

    Gall bladder

    Large intestine & caecum

    Big capacity

    Slow passage

    Mucus & motility

    Bends & flexures

    Microbial fermentation

    Absorption VFAs, H2O, NH3, vits

    Acetic, butyric & proprionic

    Designed for high fibre diet low in soluble

    carbohydrate & no sudden changes in diet

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    Natural feeding behaviour

    14-18 hours per day grazing

    Covering 10-15 km per day Grasses preferred with some legume/ browse

    The domesticated horse

    The equine athlete Sources of energy

    Horse at pasture

    75 to 80% of energy from VFAs (ie hindgut

    digestion

    Horse in heavy work (fed lots of grain)

    obtain 70 to 75% of their energy from starch and

    oil digested in the small intestine

    How do we know a diet is adequate?

    The horse can

    Perform as desired

    Maintain appropriate body condition and weight

    Excess dietary energy not desirable

    Assessing fatness/adequacy of diet

    Humans

    Weight Skinfold thickness

    BMI

    Waist circumference

    Bioelectric impedance analysis, MRI and CT

    And the horse?

    Weight

    Condition score

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    Body condition scoring

    Condition Scores: Texas A&M

    1 POOR

    Horse is extremely emaciated. The backbone, ribs, hipbones and tailhead project prominently. Bone

    structure of the withers, shoulders and neck easily noticeable. No fatty tissues can be felt.

    2 VERY THIN

    Horse is emaciated. Slight fat covering over vertebrae. Backbone, ribs, tailhead and hipbones are

    prominent. Withers, shoulders and neck structures are discernible.

    3 THIN

    Fat built up about halfway on vertebrae. Slight fat layer can be felt over ribs, but ribs easily discernible. The

    tailhead is evident, but individual vertebrae cannot be seen. The hipbones cannot be seen, but withers,

    shoulder and neck are emphasized.

    4 MODERATELY THIN

    Negative crease along back. Faint outline of ribs can be seen. Fat can be felt along tailhead. Hip bones

    cannot be seen. Withers, neck and shoulders not obviously thin.

    Condition Scores: Texas A&M

    5 MODERATE Back is level. Ribs can be felt but not easily seen. Fat around tailhead beginning

    to feel spongy. Withers are rounded and shoulders and neck blend smoothly into the body.

    6 MODERATELY FLESHY May have a slight crease down the back. Fat on the tailhead feels soft.

    Fat over the ribs feels spongy. Fat beginning to be deposited along the sides of the withers, behind

    the sholders and along the neck.

    7 FLESHY A crease is seen down the back. Individual ribs can be felt, but noticeable filling

    between ribs with fat. Fat around tailhead is soft. Noticeable fat deposited along the withers,

    behind the shoulders and along the neck.

    8 FAT Crease down back is prominent. Ribs difficult to feel due to fat in between. Fat around

    tailhead very soft. Area along withers filled with fat. Area behind shoulders filled in flush with the

    barrel of the body. Noticeable thickening of neck. Fat deposited along the inner buttocks.

    9 EXTREMELY FAT

    Obvious crease down back. Fat is in patches over rib area, with bulging fat over tailhead, withers,

    neck and behind shoulders. Fat along inner buttocks may rub together. Flank is filled in flush withthe barrel of the body.

    Optimal body condition

    Varies with

    discipline

    Individual variation

    Evidence for

    optimal body

    condition

    Racehorses

    Endurance horses

    Overall weight and

    body condition score

    What is normal?

    Depends.

    Age

    Sex

    Purpose

    Season

    Breed

    Etc etc etc

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    Body weight

    In humans excessive body weight may be

    associated with

    Cardiovascular disease

    Type 2 diabetes

    Sleep apnea

    Osteoarthritis

    Visceral fat is important.

    Fat animals?

    Not desirable in meat producing

    animals

    Overfat heifers have decreased lifetime

    milk production

    Cats and dogs associated with

    increased risk for diabetes

    And a fat horse?

    Excessive fatness and obesity is a growing problem

    Predisposed to laminitis, metabolic disease

    Risk factor for hyerlipidemia

    Athletic performance

    Heat tolerance

    Exacerbate osteoarthritis

    Reproductive performance???

    Developmental orthopaedic disease???

    Condition scores for athletes

    BCS for broodmares

    Start of breeding season:

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    Typical body condition scores

    Dressage 2.5-3/5

    Endurance 1.5-2/5

    Eventing ~2/5

    Show horses 2.5-3/5

    Race horses 1.5-2.5/5

    1.5 trim, poor-moderate

    2.5 fleshy, mod-good

    3.5 heavy, good-fat

    What horses evolved to eat

    Evolved on prairie/ rangeland

    10-20 km per day travel

    Native grasslands

    Big variety in mostly perennial species

    Predominantly grasses, but also herbs & legumes

    Unselected for digestibility or performance

    Diet

    Highly fibrous, high in minerals, low protein

    Energy mainly from VFAs not glucose

    Pasture too good?

    But horses evolved as

    pasture grazers didnt

    they??

    Modern pasture ok for modern horses?

    No, modern pastures designed for ruminants

    High sugar, low fibre

    Not great for pasture ornaments

    Yes, modern pastures are great for some

    Rapid growly young horses, pregnancy, lactation,

    horses in hard work

    Yes, poor/older pastures are great for some

    Pasture chemical composition

    % DM Leafy RG Seeding RG

    Soluble sugars 14 10

    Hemicellulose 16 26

    Cellulose 21 27

    Lignin 3 7

    Protein 15 6

    Lipid 9 5

    Ash 8 6

    Waghorn et al, (2007) NZSAP Occ.Publ. 14.

    Whats wrong with high soluble

    carbs/ low fibre?

    MAY result in:

    Digestive disturbance, rapid fermentation

    Hindgut acidosis, risk factor for

    Laminitis

    Colic

    Azoturia, Exertional Rhabdomyolysis etc

    Behavioural problems

    Metabolic disorders, insulin resistance

    Developmental Orthopeedic Disease?

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    Nutrition and laminitis

    Pasture-associated

    laminitis Fructans

    Soluble sugars

    Grain overload

    Undigested grain in

    hindgut, rapid

    fermentation

    Can pasture be ok as sole diet?

    It can be too good

    Pasture can provide enough energy & protein

    Similar growth of TB youngstock on NZperennial ryegrass pasture as pasture + grain

    overseas

    However, macro & micro minerals?

    But can be difficult to assess quantity/ quality

    Pasture vs requirementsPasture/ hay MJ DE/kg DM g CP/kg DM

    PRG/WC NZ

    Leafy spr ing 12 230

    Stalky summer 8 100

    USA

    Bermudagrass 10 125

    Kentucky BG 9 175

    Native Meadow (h) 7 87

    Native Prairie (h) 7 64

    Requirements* MJ DE/kg DM g CP/kg DM

    Maintenance 8 80

    Growth 4-6mo 12 145

    *NRC (1989) Nutrient requirements of horses (USA)

    Pasture problems

    Too little

    Too much

    Damage to pastureand soil

    Overgrazing

    Encouraged growth ofweeds eg docks

    Horse sick paddock:faecal rejection

    Pasture

    Not just for exercising

    Nutrient content

    variable

    Faecal rejection and

    overgrazing

    Horse sick pastures

    Infective parasite

    larvae

    But all horses need forages!

    Forage basis of diet: min 1.5% BW as DM

    Essential for hindgut fermentation

    Pasture, hay, preserved forages

    Grass, alfalfa/lucerne, clover

    Quality importance

    Smell, colour, dust, stem:leaf

    Not energy dense

    Variable energy, digestibility and palatability withmaturity

    Variable nutrient

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    Conserved forages for horses Hay (soaking vs not soaking)

    Grass vs legume mix:

    Lucerne, clover, pea-vine, herb high protein, Ca

    leaf:stem, smell, dust, mould, damp, weeds etc

    Chaff (chopped legume hay or oat straw)

    Commonly lucerne, red clover, oaten, pea-vine

    Hage, baleage, silage

    Lucerne and chopped oat hage custom made for horse

    Conventional pit pasture silage not recommended (bewaresecondary fermentation, clostridium)

    Baleage/ haylage lower risk

    Pasture and macrominerals

    Ca:P min ratio 1.5:1 vital

    Forages tend to be high Ca, grains high P

    K - reqts easily met by forages

    Na - specific reqts not known, adaption to Na

    restriction, pasture likely to be limiting?

    Cl - specific reqts unknown, adequate when Na reqts

    met with NaCl

    Pasture Ca P Cu Se

    PRG/WC 3.3-4.2 3.0-3.8 7.5 0.02

    Tall Fescue 3.0 3.1 6.3 0.02

    Reqts.* Ca P Cu Se

    Maint. 2.4 1.7 10 0.1

    Growth 4-6

    months

    5.6-

    6.8

    2.2-3.4 10 0.1

    Pasture vs requirements mins

    *NRC (1989), Hoskin & Gee (2004)

    g/kg DM mg/kg DM

    Nutritional-secondary

    hyperparathyroidism NSHP Millers, Bran or Big Head Disease

    From diet with low Ca:P, low [Ca], oxalates (kikuyu

    grass)

    High P levels or oxalates can impair utilization of Ca

    which result in low blood Ca levels.

    Stimulation of parathyroid gland which results in Ca

    removal from bones

    Often caused by high feeding levels of wheat bran

    (Ca:P ratio, 1:12), tropical grasses like kikuyu (oxalates)

    Microminerals

    Mg - deficient pastures causing Mg tetany in

    ruminants ok for horses?

    Behavioural problems w spring pasture, low Mg?

    Lots of supplements sold

    Research required

    Microminerals

    Co*, pastures inadequate for ruminants ok for horses

    Cu, important for growth, developmental orthopaedic

    disease prevention, potential interaction with other

    minerals, NZ soils deficient

    I*, deficiency = goiter, abnormal oestrus

    Mn*, Mo* not deficient in NZ pastures?

    Se, NZ soils deficient = white muscle disease,

    nutritional myopathy, toxicity w overdose supplements

    Fl* Zn*, Fe*?

    * Not specifically determined for horses

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    Vitamins Fat & water soluble vits high in fresh forage

    Hay storage reduces vit content

    Soaking hay leaches vits

    B & K synthesised by hindgut bacteria

    But absorption may vary

    High fibre diet needed

    Deficiencies may occur w low fibre pasture, acidosis, concentratefeeding

    D, need ultraviolet light (stables & covers)

    Vit C synthesised in hepatic tissues

    Biotin supplementation above adequate enhanced hoofgrowth in some horses

    But

    We usually dont know the nutrient content of

    pastures or hays for horses!

    Review lecture 2

    Pasture is a good, cheap source of nutrition for horses,especially young horses, broodmares

    PRG good energy and protein, marginal Ca? NB usually dont know the nutrient content

    Energy content can be too high

    NSH oxalates bind Ca, altered Ca:P BigHead

    PRG may have low fibre and high sugar during rapidgrowth that not all horses do well on GI disturbances, behaviour

    Overgrazing, lawns and troughs, faecal contaminationand parasites

    Ryegrass staggers

    Lolitrem B in high endophyte grasses Yes, affects horses too

    Over-react to external stimuli Muscles tremors, uncoordinated gait

    Usually late summer after drought when grazingvery low

    Remove from pasture, hay Low or no endophyte pasture? Horse friendly

    pasture?

    Blood tests Mycotoxin binders

    Feeding the equine athleteWhy not pasture alone?

    As energy requirements increase

    the ration needs to become moreenergy dense

    Maintenance: eats about 2% BWper day as DM

    Can consume up to max 3% BW perday as DM

    Pasture has low energy density

    BUT must consume at least 1.5%BW as forage each day for GI health

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    What other energy sources are there?

    Grains (oats, barley, maize)

    Premixed feeds (grains, chaff, minerals and

    vitamins often)

    Fats and oils

    Superfibres

    (Protein)

    Meeting energy requirements:

    carbohydrates

    Oats

    Barley Maize

    Palatable

    Energy varies

    Starch digested in small

    intestine

    -amylase limiting: no more

    than 2.5kg per meal!

    GI disturbances common

    Meeting energy requirements:

    carbohydrates

    Processed grains

    Loose

    Pelleted

    Extruded

    Sweetfeeds

    Added sugar

    Complete and premixed

    feeds

    Must be fed as directed

    Same for balancers

    Are grains bad?

    No!

    Energy dense, variable protein, often low in

    minerals and vitamins

    Quality variable

    Affordable

    Problems with too much grain

    Rapid fermentation of undigested grain in hindgut Behaviour

    Problems with grain: grain overload

    Undigested soluble CHO in hindgut

    Proliferation of lactic acid producing bacteria

    Change pH

    Death of gram negative bacteria

    Endotoxin in cell wall

    Any disruption in mucosa allows endotoxin to be absorbed

    Laminitis and endotoxaemia

    How much grain to feed?

    Limit of 2.5kg per

    meal/500kg

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    Grain meals

    Sugar high after meal

    Linked with

    Hot and excitable behaviour

    Stereotypies

    Muscle problems in susceptible horses

    Development orthopaedic disease in

    youngsters

    What if grains make the horse too hot?

    Try other energy sources

    Fermented forages

    Other sources of highly digestible fibre

    Sugar beet pulp

    Soya hulls

    Fat-fortified feeds and fat supplements

    Vegetable oil

    Stabilised rice bran

    Alternative energy sourcesFats and Oils

    Incorporated in many prepared feeds

    Up to 10% DE (even up to 25% of total MJ)

    Energy dense so smaller volume required

    Vegetable oils more palatable, introduceslowly

    Performance benefits?

    Calmer

    Improved fat oxidation; glycogen sparing

    Fats and Oils

    Incorporated in many prepared feeds

    Up to 10% DE (even up to 25% of total MJ)

    Energy dense so smaller volume required

    Vegetable oils more palatable, introduceslowly

    Performance benefits?

    Calmer

    Improved fat oxidation; glycogen sparing

    Protein Supplements

    Often overdone!

    Usually as the energy content increases the

    protein also increases

    Most protein supplements 20-25%CP

    Quality of protein in important

    lysine for young horses

    Usually expensive

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    Minerals and vitamins

    Often overdone!

    Complete, premixed and balancer feeds

    Ca:P ratio (NB grains)

    Iron not required

    Selenium important

    Toxicity: acute and chronic

    Other supplements

    Often not much information

    Overdone?

    Creatine, carnitine, branch chain amino acids

    Antioxidants

    Evaluating the diet of a horse

    Diet evaluation

    Check energy content

    Usually too much (fat, fizzy horse), or too little

    (thin, lethargic horse)

    Check forage content and quality of feedstuffs

    At least 1.5% BW as forage DM each day

    Check protein content

    Minerals and vitamins

    Basic rules

    50 to 100% diet as forage

    Horses eat between 1.5 and 3% of their bodyweight as drymatter each day

    Small stomach

    Important hindgut bacteria

    No more than 2.5kg grain at any time

    Small feeds often

    Feed by weight not volume

    Change feeds slowly (hindgut bacteria)

    Feed as individuals

    Optimal nutrition

    May optimise performance

    Wont improve genetic potential

    Wont make up for poor training

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    Dietary problems in the equineathlete

    Feeding behaviour issues

    Horses are not nutritionally wise!

    Feed shortage = abnormal behaviour

    High concentrate:roughage / low roughage availability

    = abnormal behaviour Stabling w. ad-lib hay = similar behaviour to grazing

    pasture

    Feeding time budget constant despite herbageavailability

    Herbage shortage, horse needs to extend feedingbudget but doesnt = undernutrition

    Herbage excess, horse needs to restrict feedingbudget but doesnt = overnutrition

    Effect of feed type on time spent behaving

    abnormally in horses(Cuddeford,1996)

    nFeed Feed

    level% of time

    spent

    feeding

    % time inabnormal

    activities

    Pasture 6 Grass Ad-lib. 58 0

    Stable 5 Hay Ad-lib. 55 2

    Tie-stall 5 Hay Ad-lib. 64

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    Typical diets

    12 year old pony 450kg (430-480. Prone to beingoverweight when not in work.)

    Currently: hunts twice weekly. 3kg red clover/ryegrass hay

    Balancer pellets for mineralsand vitamins

    Sugarbeet if pasturequality/quantity decreases

    Oats if reqd

    Summer: Jenny Craigpaddock Balancer pellets for minerals

    and vitamins

    Grass hay (stalky)

    Typical diets

    5yo Standardbred gelding, full training 480kg

    Paddock trained

    Approx 6kg premixed feed

    1-2kg fermented forage

    1-1.5kg lucerne/oaten chaff

    Electrolytes

    Mineral mix

    Apple cider vinegar

    Garlic

    Summary

    Optimal nutrition for optimal performance

    Each horse is an individual and needs to be

    fed as such

    Forage is more than a filler

    Grains arent always bad but there are

    alternatives

    Aim for a balanced dietthere can be toomuch of a good thing

    Questions?