feeding horses student version 2013
TRANSCRIPT
-
7/28/2019 Feeding Horses Student Version 2013
1/14
26/04/20
227.208 Equine Nutrition
Erica Gee BVSc, PhD, Dip ACTRoom 1.21
[email protected] 2013
Acknowledgement: Dr Simone Hoskins
Equine Nutrition Lectures
Monday 29th April 11am SSLB6
Thursday 2nd May 9am SSLB6
Monday 6th May 11am SSLB6
Powerpoints on Stream, plus required reading
Who am I?
Senior lecturer in Equine Science
Equine nutrition and reproduction
Learning objectives
Evaluate the diet of a horse
Explain the main principles of feeding horses
Discuss the benefits and potential problems of
pasture based diets for horses
Discuss the effects of nutrition on health and
well-being of horses
Lecture outline
The basics of feeding horses (and why the horse isnot a cow!)
The role of fibre
Assessing fatness and adequacy of diet
Pasture as a nutrient source
Where problems occur & how to prevent them by
feeding practice
Why feeding is both art & science
Prior knowledge assumptions
Equine GI tract anatomy
Equine GI tract physiology
Nutrients & nutrient digestion
Protein & amino acids
Lipids
Carbohydrates & fibre
Vitamins & minerals
Some knowledge of equine management, behaviour,breeds etc
mailto:[email protected]:[email protected]:[email protected]:[email protected] -
7/28/2019 Feeding Horses Student Version 2013
2/14
26/04/20
The digestive system
Foregut
Mouth
Oesophagus
Stomach
Small intestine
What is happening in the small
intestine?
Enzymes, bile, other digestive juices
Feed passes through in 30-90 minutes
Digestion
Soluble carbohydrates (sugars and starch)
glucose
What enzyme is rate limiting for starch digestion?
Most protein from grains
Oils and fats
Calcium, phosphorus, B vitamins, vitamins A, D, E, K
The equine digestive system
The hindgut
Caecum
Large colon
Small colon
Rectum
What happens in the hindgut?
Giant fermentation vat
Bacteria break down fibre volatile fatty
acids (VFA)
B vitamins
Fermentation takes days
Quality
Water reservoir
Features of GI tract Lips, tongue, teeth
High degree of selectivity
Fibre grinders
Stomach
Small capacity, fast passage
Cardiac sphincter = one way valve
Normally rarely empty & highly buffered
Small intestine
Relatively fast passage
Urea secretion terminal ileum
Gall bladder
Large intestine & caecum
Big capacity
Slow passage
Mucus & motility
Bends & flexures
Microbial fermentation
Absorption VFAs, H2O, NH3, vits
Acetic, butyric & proprionic
Designed for high fibre diet low in soluble
carbohydrate & no sudden changes in diet
-
7/28/2019 Feeding Horses Student Version 2013
3/14
26/04/20
Natural feeding behaviour
14-18 hours per day grazing
Covering 10-15 km per day Grasses preferred with some legume/ browse
The domesticated horse
The equine athlete Sources of energy
Horse at pasture
75 to 80% of energy from VFAs (ie hindgut
digestion
Horse in heavy work (fed lots of grain)
obtain 70 to 75% of their energy from starch and
oil digested in the small intestine
How do we know a diet is adequate?
The horse can
Perform as desired
Maintain appropriate body condition and weight
Excess dietary energy not desirable
Assessing fatness/adequacy of diet
Humans
Weight Skinfold thickness
BMI
Waist circumference
Bioelectric impedance analysis, MRI and CT
And the horse?
Weight
Condition score
-
7/28/2019 Feeding Horses Student Version 2013
4/14
26/04/20
Body condition scoring
Condition Scores: Texas A&M
1 POOR
Horse is extremely emaciated. The backbone, ribs, hipbones and tailhead project prominently. Bone
structure of the withers, shoulders and neck easily noticeable. No fatty tissues can be felt.
2 VERY THIN
Horse is emaciated. Slight fat covering over vertebrae. Backbone, ribs, tailhead and hipbones are
prominent. Withers, shoulders and neck structures are discernible.
3 THIN
Fat built up about halfway on vertebrae. Slight fat layer can be felt over ribs, but ribs easily discernible. The
tailhead is evident, but individual vertebrae cannot be seen. The hipbones cannot be seen, but withers,
shoulder and neck are emphasized.
4 MODERATELY THIN
Negative crease along back. Faint outline of ribs can be seen. Fat can be felt along tailhead. Hip bones
cannot be seen. Withers, neck and shoulders not obviously thin.
Condition Scores: Texas A&M
5 MODERATE Back is level. Ribs can be felt but not easily seen. Fat around tailhead beginning
to feel spongy. Withers are rounded and shoulders and neck blend smoothly into the body.
6 MODERATELY FLESHY May have a slight crease down the back. Fat on the tailhead feels soft.
Fat over the ribs feels spongy. Fat beginning to be deposited along the sides of the withers, behind
the sholders and along the neck.
7 FLESHY A crease is seen down the back. Individual ribs can be felt, but noticeable filling
between ribs with fat. Fat around tailhead is soft. Noticeable fat deposited along the withers,
behind the shoulders and along the neck.
8 FAT Crease down back is prominent. Ribs difficult to feel due to fat in between. Fat around
tailhead very soft. Area along withers filled with fat. Area behind shoulders filled in flush with the
barrel of the body. Noticeable thickening of neck. Fat deposited along the inner buttocks.
9 EXTREMELY FAT
Obvious crease down back. Fat is in patches over rib area, with bulging fat over tailhead, withers,
neck and behind shoulders. Fat along inner buttocks may rub together. Flank is filled in flush withthe barrel of the body.
Optimal body condition
Varies with
discipline
Individual variation
Evidence for
optimal body
condition
Racehorses
Endurance horses
Overall weight and
body condition score
What is normal?
Depends.
Age
Sex
Purpose
Season
Breed
Etc etc etc
-
7/28/2019 Feeding Horses Student Version 2013
5/14
26/04/20
Body weight
In humans excessive body weight may be
associated with
Cardiovascular disease
Type 2 diabetes
Sleep apnea
Osteoarthritis
Visceral fat is important.
Fat animals?
Not desirable in meat producing
animals
Overfat heifers have decreased lifetime
milk production
Cats and dogs associated with
increased risk for diabetes
And a fat horse?
Excessive fatness and obesity is a growing problem
Predisposed to laminitis, metabolic disease
Risk factor for hyerlipidemia
Athletic performance
Heat tolerance
Exacerbate osteoarthritis
Reproductive performance???
Developmental orthopaedic disease???
Condition scores for athletes
BCS for broodmares
Start of breeding season:
-
7/28/2019 Feeding Horses Student Version 2013
6/14
26/04/20
Typical body condition scores
Dressage 2.5-3/5
Endurance 1.5-2/5
Eventing ~2/5
Show horses 2.5-3/5
Race horses 1.5-2.5/5
1.5 trim, poor-moderate
2.5 fleshy, mod-good
3.5 heavy, good-fat
What horses evolved to eat
Evolved on prairie/ rangeland
10-20 km per day travel
Native grasslands
Big variety in mostly perennial species
Predominantly grasses, but also herbs & legumes
Unselected for digestibility or performance
Diet
Highly fibrous, high in minerals, low protein
Energy mainly from VFAs not glucose
Pasture too good?
But horses evolved as
pasture grazers didnt
they??
Modern pasture ok for modern horses?
No, modern pastures designed for ruminants
High sugar, low fibre
Not great for pasture ornaments
Yes, modern pastures are great for some
Rapid growly young horses, pregnancy, lactation,
horses in hard work
Yes, poor/older pastures are great for some
Pasture chemical composition
% DM Leafy RG Seeding RG
Soluble sugars 14 10
Hemicellulose 16 26
Cellulose 21 27
Lignin 3 7
Protein 15 6
Lipid 9 5
Ash 8 6
Waghorn et al, (2007) NZSAP Occ.Publ. 14.
Whats wrong with high soluble
carbs/ low fibre?
MAY result in:
Digestive disturbance, rapid fermentation
Hindgut acidosis, risk factor for
Laminitis
Colic
Azoturia, Exertional Rhabdomyolysis etc
Behavioural problems
Metabolic disorders, insulin resistance
Developmental Orthopeedic Disease?
-
7/28/2019 Feeding Horses Student Version 2013
7/14
26/04/20
Nutrition and laminitis
Pasture-associated
laminitis Fructans
Soluble sugars
Grain overload
Undigested grain in
hindgut, rapid
fermentation
Can pasture be ok as sole diet?
It can be too good
Pasture can provide enough energy & protein
Similar growth of TB youngstock on NZperennial ryegrass pasture as pasture + grain
overseas
However, macro & micro minerals?
But can be difficult to assess quantity/ quality
Pasture vs requirementsPasture/ hay MJ DE/kg DM g CP/kg DM
PRG/WC NZ
Leafy spr ing 12 230
Stalky summer 8 100
USA
Bermudagrass 10 125
Kentucky BG 9 175
Native Meadow (h) 7 87
Native Prairie (h) 7 64
Requirements* MJ DE/kg DM g CP/kg DM
Maintenance 8 80
Growth 4-6mo 12 145
*NRC (1989) Nutrient requirements of horses (USA)
Pasture problems
Too little
Too much
Damage to pastureand soil
Overgrazing
Encouraged growth ofweeds eg docks
Horse sick paddock:faecal rejection
Pasture
Not just for exercising
Nutrient content
variable
Faecal rejection and
overgrazing
Horse sick pastures
Infective parasite
larvae
But all horses need forages!
Forage basis of diet: min 1.5% BW as DM
Essential for hindgut fermentation
Pasture, hay, preserved forages
Grass, alfalfa/lucerne, clover
Quality importance
Smell, colour, dust, stem:leaf
Not energy dense
Variable energy, digestibility and palatability withmaturity
Variable nutrient
-
7/28/2019 Feeding Horses Student Version 2013
8/14
26/04/20
Conserved forages for horses Hay (soaking vs not soaking)
Grass vs legume mix:
Lucerne, clover, pea-vine, herb high protein, Ca
leaf:stem, smell, dust, mould, damp, weeds etc
Chaff (chopped legume hay or oat straw)
Commonly lucerne, red clover, oaten, pea-vine
Hage, baleage, silage
Lucerne and chopped oat hage custom made for horse
Conventional pit pasture silage not recommended (bewaresecondary fermentation, clostridium)
Baleage/ haylage lower risk
Pasture and macrominerals
Ca:P min ratio 1.5:1 vital
Forages tend to be high Ca, grains high P
K - reqts easily met by forages
Na - specific reqts not known, adaption to Na
restriction, pasture likely to be limiting?
Cl - specific reqts unknown, adequate when Na reqts
met with NaCl
Pasture Ca P Cu Se
PRG/WC 3.3-4.2 3.0-3.8 7.5 0.02
Tall Fescue 3.0 3.1 6.3 0.02
Reqts.* Ca P Cu Se
Maint. 2.4 1.7 10 0.1
Growth 4-6
months
5.6-
6.8
2.2-3.4 10 0.1
Pasture vs requirements mins
*NRC (1989), Hoskin & Gee (2004)
g/kg DM mg/kg DM
Nutritional-secondary
hyperparathyroidism NSHP Millers, Bran or Big Head Disease
From diet with low Ca:P, low [Ca], oxalates (kikuyu
grass)
High P levels or oxalates can impair utilization of Ca
which result in low blood Ca levels.
Stimulation of parathyroid gland which results in Ca
removal from bones
Often caused by high feeding levels of wheat bran
(Ca:P ratio, 1:12), tropical grasses like kikuyu (oxalates)
Microminerals
Mg - deficient pastures causing Mg tetany in
ruminants ok for horses?
Behavioural problems w spring pasture, low Mg?
Lots of supplements sold
Research required
Microminerals
Co*, pastures inadequate for ruminants ok for horses
Cu, important for growth, developmental orthopaedic
disease prevention, potential interaction with other
minerals, NZ soils deficient
I*, deficiency = goiter, abnormal oestrus
Mn*, Mo* not deficient in NZ pastures?
Se, NZ soils deficient = white muscle disease,
nutritional myopathy, toxicity w overdose supplements
Fl* Zn*, Fe*?
* Not specifically determined for horses
-
7/28/2019 Feeding Horses Student Version 2013
9/14
26/04/20
Vitamins Fat & water soluble vits high in fresh forage
Hay storage reduces vit content
Soaking hay leaches vits
B & K synthesised by hindgut bacteria
But absorption may vary
High fibre diet needed
Deficiencies may occur w low fibre pasture, acidosis, concentratefeeding
D, need ultraviolet light (stables & covers)
Vit C synthesised in hepatic tissues
Biotin supplementation above adequate enhanced hoofgrowth in some horses
But
We usually dont know the nutrient content of
pastures or hays for horses!
Review lecture 2
Pasture is a good, cheap source of nutrition for horses,especially young horses, broodmares
PRG good energy and protein, marginal Ca? NB usually dont know the nutrient content
Energy content can be too high
NSH oxalates bind Ca, altered Ca:P BigHead
PRG may have low fibre and high sugar during rapidgrowth that not all horses do well on GI disturbances, behaviour
Overgrazing, lawns and troughs, faecal contaminationand parasites
Ryegrass staggers
Lolitrem B in high endophyte grasses Yes, affects horses too
Over-react to external stimuli Muscles tremors, uncoordinated gait
Usually late summer after drought when grazingvery low
Remove from pasture, hay Low or no endophyte pasture? Horse friendly
pasture?
Blood tests Mycotoxin binders
Feeding the equine athleteWhy not pasture alone?
As energy requirements increase
the ration needs to become moreenergy dense
Maintenance: eats about 2% BWper day as DM
Can consume up to max 3% BW perday as DM
Pasture has low energy density
BUT must consume at least 1.5%BW as forage each day for GI health
-
7/28/2019 Feeding Horses Student Version 2013
10/14
26/04/20
What other energy sources are there?
Grains (oats, barley, maize)
Premixed feeds (grains, chaff, minerals and
vitamins often)
Fats and oils
Superfibres
(Protein)
Meeting energy requirements:
carbohydrates
Oats
Barley Maize
Palatable
Energy varies
Starch digested in small
intestine
-amylase limiting: no more
than 2.5kg per meal!
GI disturbances common
Meeting energy requirements:
carbohydrates
Processed grains
Loose
Pelleted
Extruded
Sweetfeeds
Added sugar
Complete and premixed
feeds
Must be fed as directed
Same for balancers
Are grains bad?
No!
Energy dense, variable protein, often low in
minerals and vitamins
Quality variable
Affordable
Problems with too much grain
Rapid fermentation of undigested grain in hindgut Behaviour
Problems with grain: grain overload
Undigested soluble CHO in hindgut
Proliferation of lactic acid producing bacteria
Change pH
Death of gram negative bacteria
Endotoxin in cell wall
Any disruption in mucosa allows endotoxin to be absorbed
Laminitis and endotoxaemia
How much grain to feed?
Limit of 2.5kg per
meal/500kg
-
7/28/2019 Feeding Horses Student Version 2013
11/14
26/04/20
Grain meals
Sugar high after meal
Linked with
Hot and excitable behaviour
Stereotypies
Muscle problems in susceptible horses
Development orthopaedic disease in
youngsters
What if grains make the horse too hot?
Try other energy sources
Fermented forages
Other sources of highly digestible fibre
Sugar beet pulp
Soya hulls
Fat-fortified feeds and fat supplements
Vegetable oil
Stabilised rice bran
Alternative energy sourcesFats and Oils
Incorporated in many prepared feeds
Up to 10% DE (even up to 25% of total MJ)
Energy dense so smaller volume required
Vegetable oils more palatable, introduceslowly
Performance benefits?
Calmer
Improved fat oxidation; glycogen sparing
Fats and Oils
Incorporated in many prepared feeds
Up to 10% DE (even up to 25% of total MJ)
Energy dense so smaller volume required
Vegetable oils more palatable, introduceslowly
Performance benefits?
Calmer
Improved fat oxidation; glycogen sparing
Protein Supplements
Often overdone!
Usually as the energy content increases the
protein also increases
Most protein supplements 20-25%CP
Quality of protein in important
lysine for young horses
Usually expensive
-
7/28/2019 Feeding Horses Student Version 2013
12/14
26/04/20
Minerals and vitamins
Often overdone!
Complete, premixed and balancer feeds
Ca:P ratio (NB grains)
Iron not required
Selenium important
Toxicity: acute and chronic
Other supplements
Often not much information
Overdone?
Creatine, carnitine, branch chain amino acids
Antioxidants
Evaluating the diet of a horse
Diet evaluation
Check energy content
Usually too much (fat, fizzy horse), or too little
(thin, lethargic horse)
Check forage content and quality of feedstuffs
At least 1.5% BW as forage DM each day
Check protein content
Minerals and vitamins
Basic rules
50 to 100% diet as forage
Horses eat between 1.5 and 3% of their bodyweight as drymatter each day
Small stomach
Important hindgut bacteria
No more than 2.5kg grain at any time
Small feeds often
Feed by weight not volume
Change feeds slowly (hindgut bacteria)
Feed as individuals
Optimal nutrition
May optimise performance
Wont improve genetic potential
Wont make up for poor training
-
7/28/2019 Feeding Horses Student Version 2013
13/14
26/04/20
Dietary problems in the equineathlete
Feeding behaviour issues
Horses are not nutritionally wise!
Feed shortage = abnormal behaviour
High concentrate:roughage / low roughage availability
= abnormal behaviour Stabling w. ad-lib hay = similar behaviour to grazing
pasture
Feeding time budget constant despite herbageavailability
Herbage shortage, horse needs to extend feedingbudget but doesnt = undernutrition
Herbage excess, horse needs to restrict feedingbudget but doesnt = overnutrition
Effect of feed type on time spent behaving
abnormally in horses(Cuddeford,1996)
nFeed Feed
level% of time
spent
feeding
% time inabnormal
activities
Pasture 6 Grass Ad-lib. 58 0
Stable 5 Hay Ad-lib. 55 2
Tie-stall 5 Hay Ad-lib. 64
-
7/28/2019 Feeding Horses Student Version 2013
14/14
26/04/20
Typical diets
12 year old pony 450kg (430-480. Prone to beingoverweight when not in work.)
Currently: hunts twice weekly. 3kg red clover/ryegrass hay
Balancer pellets for mineralsand vitamins
Sugarbeet if pasturequality/quantity decreases
Oats if reqd
Summer: Jenny Craigpaddock Balancer pellets for minerals
and vitamins
Grass hay (stalky)
Typical diets
5yo Standardbred gelding, full training 480kg
Paddock trained
Approx 6kg premixed feed
1-2kg fermented forage
1-1.5kg lucerne/oaten chaff
Electrolytes
Mineral mix
Apple cider vinegar
Garlic
Summary
Optimal nutrition for optimal performance
Each horse is an individual and needs to be
fed as such
Forage is more than a filler
Grains arent always bad but there are
alternatives
Aim for a balanced dietthere can be toomuch of a good thing
Questions?