-
1988KELP INVENTORY,
JUAN DE FUCA STRAIT
prepared by
I. R. Sutherland
IEC Collaborative Marine Research
and Development Ltd.
Fisheries Development Report No.35
Aquaculture and Commercial Fisheries Branch
Ministry of Agriculture and Fisheries
Province of British Columbia
March, 1989
-
-i-
ABSTRAcr
A modification of the Kelp Inventory Method ( KIM-1 ) developed
by Fore:nan (1975) was used to estimate the total standing crop bianass
of Nereo~stis luetkeana along the Canadian shore of Juan de Fuca Strait
for August, 1988. Estimates of kelp bed area were produced for ~
luetkeana and Macro~stis int§g:J;:ifolia. Results indicated that 50,148
tonnes of ~ luetkeana were present. 'Ibtal bed surface area was
estimated to be 511 hectares, of which less than 2% was ~ int§g:J;:ifolia .
Six charts are presented which sho..l the position, extent, species, and
densityof every discernable kelp bed within the survey area. For
managenent purposes the area is divided to permanent, numbered,
kilaneter wide blocks .
-
-ii -
ACKNavr=-rS
The author wishes to thank Michael Coon, Bob Cox and Ted Pobran
who braved the ele:nents to collect kelp bianass data .Michael Coon and
Bob Cox also reviewed m:>st of the manuscript. Thanks are also due to
Bill Roland for his suggestions regarding mean bianass per plant
calculations.
-
1
IN'IRODUCI'ION
Nereo~stis luetkeana (Mertens) Pastels and Ruprecht and
Macro~stis int§gI:ifolia Bory form beds along extensive portions of the
British Columbia coast. In 1975 the Provincial Government undertook a
program to locate and quantify the standing crop of these econcmicall y
~rtant kelps using the inventory method ( KIM-1 ) developed by Forenan
(1975) .Since that time major kelp stocks have been inventoried
throughout the province (Coon, 1981. Coon et al, 1977, 1979, 1980,
1981, 1982. Field et al., 1975, 1977, 1978).
This report contains the results of a 1988 surveyof the
Vancouver Island shore of Juan de Fuca Strait by the Aquaculture and
CaTmercial Fisheries Branch of the Ministry of Agriculture and
Fisheries. The Ministry provided field sampling data and aerial
photography under contract to IEC CollaOOrative Marine Research and
Developnent Ltd .for CCITIpletion of the inventory .
Accurate and CCITIprehensi ve data on the standing crop of kelp in
British Columbia provide a basis for allocating these resources through
licensing and for establishing area specific harvest quotas. Because
kelp beds are ~rtant to other marine species, kelp inventory charts
and data will be of value to those preparing envirornnental impact
statanents, or conducting surveys of herring spawn , abalone and sea
urchins (Coon, 1977).
-
3 -
METHODS
The standing crop bia1lasS of Nereo~stis and kelp bed areas of
both Nereo~stis and Macro~stis were esti1Tlated by a m:>dification of the
kelp inventory method (Krn-1) developed by Fore:nan ( 1975) .Modifications
of this method as stated by Coon et al ( 1977) and Field et al ( 1977 )
were used in this study .A further change was introduced in this
inventory by estimating the total standing crop bia1lasS of Nereo~stis
rather than the harvestable bia1lasS at various cutting levels. This
method was first used in the Porcher Island area by Coon ( 1981) .
Briefly the Krn-1 technique involves obtaining 24 an. format
black and white infrared (IR) aerial photography of the kelp bed and
shoreline in the desired region. The black and white IR negatives are
used to prepare charts of the coastline and offshore kelp beds .On
these charts the survey area is di vided to 1 kilaneter wide statistical
blocks .Bed areas for each of the six bed types described below are
dete.rmined for each block. The density of kelp is deteImined directly
fraT! the photographs with the aid of a microscope. Field crellTS obtain
samples of kelp fraT! the area for mean weight per plant ( Nereo~stis ) or
frond (Macro~stis) deteImination, near the time that the beds are
photographed. The kelp bia11asS per block is dete.rmined by multiplying
the mean weight per plant or frond values by the observed plant or frond
densities and multiplying this product by the observed bed areas .
The Krn-1 technique identifies six bed types on the basis of :
a) species -Macro~stis or Nereo~stis or mixed (42%
Nereo~stis and 58% Macro~stis; Fore:nan,
1975); and
b) plant or frond density -la..l density (less than 10 plants
or fronds per 10 square meters) or high
( greater than 10 plants or fronds per 10 square
meters).The kelp bed area and density esti1Tlates given in this reI;XJrt are
derived fran August 27, 1988 photography. IDcal tidal characteristics
canbined with the large distances covered by this inventory resulted in
photography of the more easter 1 y area at 1 meter la..ler than the opti1Tlal
-
5
RESULTS
Charts A through F (Appendix) illustrate the distX)sition of kelp
bed resources by bed type along the Canadian shore of Juan de Fuca
Strait. Coverage extends fran Race Passage to Clo-oose. It will be
noted fran the charts and Figure I that tX)rtions of the a)astlines of
Becher Bay and Port San Juan are not represented .This is due to
ina:lnplete photographic ooverage. Sufficient space and block numbers
have been reserved for these areas should the need arise for their
inclusion in a later inventory .
Table I presents the field-determined mean bianass per plant (wet
weight) estimate obtained fran means of eight sampling stations in the
inventory area September 12, 13 and 14, 1988. A total of 110 plants
were sampled .
A sUIm1ary of Ctnnulati ve numbers of Nereo~stis plants oollected
in the above sampling is expressed as percent of sample totals in Table
2. This table shC1ofls the vertical distribution of kelp numbers in I
meter increnents above and below mean water level (MWL ) , and was used
to calculate the density oorrection factors .
A sUIm1ary of the tide level at the time of photography and the
oonversion factor used to change the observed density to total (or
bot tan) density for each block is presented in Table 3.
Tables 4 to 9 present estimates of kelp bed areas, density and
total Nereo~stis bianass for each block in Charts A through F (Figure
1, and Appendix) .Macro~stis density measurenents and bianass samples
were not taken for this area due to the small quantity present, but bed
area has been retX)rted .Macro~stis acoounted for only 9 .5 hectares, or
slightly less than 2%, of the total kelp bed area of 510.6 hectares.
Nereo~stis mean bianass per plant estimate fran Septanber12, 13 and 14, 1988 field samples -used to calculate bianassestimates for the 1988 Juan de Fuca Strait inventory area.
1:
4.9 kg. SD = 1.8Nereo~stis mean biaTlass/plant = 8 stations
-
6
Table 2: Cumulative percent of plants present at one meterincrements relative to mean water (MWL) for samplesof Nereocystis from the inventory area September 12,13 and 14, 1988.
Mean PercentPlants inincrement1 2Station:
Meter+6+5+4+3+2+1MWL-1-2-3-4-5-6-7-8-9-10
3 4 5 6 7 8
4789999
1010101010101010
11268999999
101010
73693
100100100100100100100100100100
1
1
1
1
1
1
1
1
35888999
101010
479
101010101010101010
3889
1010101010101010
54100100100100100100100100100100
5689
100100100100100100100100100
Table 3: The tidal level at the time of photography and theNereo~stis conversion factors used to convert densityvalues observed on the photography to total density as usedin bianass calculations .
Block Nos. Tidal I.evel(m) Conversion Factor
1-7,9-20 -1 1.16
21-65 MWL 1.49
66-75,89-118 +1 3.70
Mix 00 beds were not found in this region. Tables 10, 11 and 12 slUm\arize
the J:::6j area and bianass estimates by bOO type, by percent Nereo~stis
bianass and by percent CCI11position of bOO area for each bed bed type in
each Chart area, respectively.
8
8
6
7
5
22
2
2
0
0
0
0
0
0
0
O
1
1
6
O
27
7
6
5
8
8
9
9
9
0
0
0
5790000000O
7
7O
1
3
0
0
0
0
0
0
0
0
58
7
1
6
6
0
0
0
0
0
O
531OOOOOOOOO
3O7
3
0
0
0
0
0
0
0
0
-
-7 -
A total of 50,148 tonnes of Nereo~stis were estimated to lie
along the 104 kilaneters of shoreline inventoried. The major
roncentrations were found to be at Muir Point (Blocks 17-19; 6,262
tonnes) , Orveas Bay (Blocks 28,29 ; 2,707 tonnes) , immediately west of
Sambrio Point (Blocks 62-64; 6,827 tonnes) and near Bonilla Point
(Blocks 102-106; 11,807 tonnes).It should be noted that large bianass figures reported for
Blocks 66 to 75 and 89 to 118 are largely the result of the application
of the ronversion factor fran Table 3. This is evident in the density
increase between the "Photo XD" and the "Bot tan XD" figures reported in
Tables 8 and 9 .A high proportion of Nereo~stis plants in the bianass
samples were found to extend only to MWL or lC1tller ( see Table 2 ) , a
situation Irore typical of early to mid SUImler. Only 27% of the plants
sampled extended to the MWL+ 1 tide level. This has resulted in the
large ronversion factor of 3.70 for the +1 level (Table 3) .Acrording
to this method, Charts E and F rontain 43% of the biomass but only 30%
of the area for Nereo~stis in this inventory .
-
V1
~ -0 V1
"'C))
C))
0- 0
~
"N ~ 1.11 mN , ~ -- .0 ~ .0 O '.I$ VI -..1n ~ 01 , ... ~ --1 O ... 01 .- ."
NN
~l:-
l:-N
l:-
o-O
Nl:-
Oo-
(X)N
--.j-
oN...
l: (X
)~
ol:-
V10
--.j-
-.jN
~~
~~
~~
IV-O
o-~
-0""
'~IV
--IV
IV
IV
O
'"'
~
V1
~~
~~
~~
~~
,-.j
,-.j
,-.j
,-.j
,-.j
,-.j
,-.j
,-.j
l-"l-"
l-"l-"
l-"l-"
l-"l-"
,N
,N
,N
,N
,N
,N
,N
,N
~
N~
~~
~
~~
~<
»V1~
~~
O-~
~~
~~
<»~
o-~
~~
V1-
oN<
»
-0-0
-0
N
\-,\-
,C»I
.nI.n
-O--
,jC»
.o~
:0,..
"!",
..,,.0
~I.n
NN
N~
IXII.
n~
,0-
,0-
,0-
,0-
,0-
,0-
,0-
,0-
C»
C»
.0
.0
.0
.0
.0
.0C
»C»V
IVIV
IVIV
IVI
aaaa
nnnn
VI
VI
VI
VI
VI
VI
VI
VI
"""" -0
-0
-0
-0
-0
-0
-0
-0
~~
-0-0
-0-0
-0-o
0000
0000
--N
--~
~N
---0
~--
~V
1--C
»--~
-oO
~
""""
""""
""""
"' "'
l:-
l:-
l:-
l:-
l:-
l:-
,
\H~
I\J
\HI\J
1»
I»O
o-o-
V1~
O~
-,j--
I\JC
»~0
"""
~~
~~
~~
~o-
V1~
\HN
~C
).O
8
--~
~~
,J
CX
I~
VI-
-~--
Vl~
aI:O
~
~!"
CX
I-.,J
-.,J
o-I\J
VI"
,
NC
XIl:
-V1-
-V1V
1O
NC
XI-
-V1-
-CX
I"'
N
N
m
-n
"' ~
NN
NN
NN
N~
~~
NN
NN
"""' -o-o
-o§
ooo
0000
00
000
000
nnn
-0-
--,j
--,j
--,j
--,j
--,j
--,j
.O0-
0-0-
0-C
»V1
V1.
0.0.
0.O
lHrv
0000
000
--0-
lHlH
lH
rvrv
-,
j ~
~
.0
--
C»
.0
--rv
-,
j C
» rv
0-
,V1
,cr-
,c
r-
,cr-
,c
r-
,cr-
,c
r-"'
cr-
cr-
cr-
cr-
.."
~"'
""
""
""
""
V1
(»O
oooo
oa
-..N
-..N
N
",8
°'0-
..~V
lO-
VlO
"'o-O
())
'0
I» I
» "'
"' -.
."'
-0-0
-0
NN
.."V
I~.."
o-O
-o~
--0
OV
I
NN
NN
NN
N"'-
I ."
'-I
"'-I
VI
VI
VI
VI
N
VI
~~
VlN
~~
0 ~
N(X
)Oo-
~~
.0
(X)
.0
(X)
.0
~
-0
\H
\H
\H
\H
\H
\H
\H~
(X
lo-
""
""
""
"'"
"'
-0
"' "'
"..
...",
--
'
ID~ >
-:1"
..,
01
It~
"'
~
>~
,
QI
~~
Q
I
~
>:T
..,
01
IU~
01
IJI
~
>:3
" ,
Dl
ID'-'
D
l
XI
~ , :3"
Dl
~ OX
I...
0 ... 0 ~~ :3
"X
I O
o ... "
ID~ >
:J"
,01
In
~
01
XI
III :J"
01
g'X
I -+
0 -+ 0 ~"0 J
x,
Oo
-+ ()
m O ~
n z~
~ ~ -'
~ z \C :T 0 IV ~ "' ~.
.-+
'
K"c
",..
tC
~
, C
'0
~.
"'n
,..
C"
, "
,..
Q.
O
:J
-0
Q/
:J
CX
) ,
" C
X)
""'
Q/
VJ
Q/
" :J
" Q
.
'"'
C"
:J"
~.
QI
O:J
" ,
3Q
I ,..
Q
I"'
>
"'
:J"
~
" >
O
n '0
,
,..
'0Q
I "
,..,
:J
:J"
" Q
. "
X
'"'~
Q
I:J Q
IQ
.
QI
:J
o "' :J
"O ,
0 "
" :J
O"' ,..
'-
'<
C QI
~
:J:J O
Q
.." 0
" C n'0
Q
IQ
;"
VJ
:J
,..,..
,
"' Q
I~
.:J
" ,..
" n ,.. :0
QI
QI
, n
" "
~"U Q
I"' "' Q
ItC " ,
.. 0-. Qj
t:r ~ ~
--"-
-"--
"-"-
0..,
...,..
.,...,
...,..
.,...,
...,.
oooo
o""o
n
0000
0000
~~
~~
~~
~~
0000
0000
1»1»
1»1»
1»1»
1»1»
NN
NN
NN
NN
-' -'
-' -'
-' -'
-'\.I
I\.11
\.11N
NN
N0-
0-
0-
V
I V
I V
I V
I
\.11\
.11\
.110
000
00
00
00
VI
VI
VI
VI
1\)1
\)1\
)1\)
1\)1
\)1\
)oo
oaaa
a
-
0......~c...~
4141
L~
10
O
...O
U
"' 41.r:
...C/)
.~
...10
CL
10'
"' c.
10 -.-
U
O;]
O41
C"U
~
C
>-
10 ...
;] .~
-, C
/)C
-.-41O
Q
41LO Q
.r:"'C10 ~X
"U
.~10
"UC
C
10 41
U
c.c.41
<.r:
~...41
aJ L
L 10
0 -.-L
U10
41C
/) .r:
.r:C
/) U
10E 41
04110.~
"'
.r:.D"U
~
C
~10
0-.-~10
C/)
41 ,
41L
...C10
C/)
C;]
0"U
C)...
41 ;]
.D
<
Uc.
, L
-' 41C41
~
.~
E0
~-.-a.0
C/)
E
C/)
C/)
10 10
41 .r:
E...C
) 0
10 C
.~
E
.~
aJL
...41C
/) .r:
W
"' aJ
U"\
41-'.0101-
.-'o
lIO
G
I ...
Lo O
GI
~Z
QU
)~
.c
i 01
U);
.~
~X
'-U Q
~I
~O-J -'10...O1->-+
"'"'c.,Q3O-'I.~I
~~I
>-
...0)c4)Q.c01~0)'.~
I
!Z1-~< aI
~'--I
10 ""'
4) 10
'- .r.
<
'""
"' ~
v "'
'- .c
<
~
m
'"'Q
J m
'- .c
d ~
0...o0
Ix.c" EO+-'
Q+
-' IX
OID
ID.c,III
Ix
ID
"v
ID
;;: 5
O...Q0
IX.cno EO...Q...IXOIII
m~a)IX
m
~Q
J m
'- ~
4 ~ mm~uO~ m
o-"'0--oo~~
OO
OC
x)Cx)"'-o
.-°"'N"'.-N
".. C
X)
".. ,
.-N.-
-0 .-IX
)
-.1" IX)
-.1"0.--00'
I/\ "'"
'.-
.-0' -.1"
cxil/\cxiN
-00-0
0 1"
","'..'-0.-
0001/\1/\-0.--0.-
-0' ,.::;
-.1"'"1"""1
0- "'
InN
.0
O
"1 "'
"'°.-"'.-~~
NN
~~
N.-"'
"' .". .--0
0' "'
.--0
I\J -0
00 "'
"' -0
00000I\J0'",.-00
I\J 0'
001n O
'OO
"'O'O
OO
OO
OI\JI\J
"""""".".Inln
000~
0-
"""""'C
X>
N'
0' 0'
NN
N
1'-- ~
0-
O
...N
1'1 ~
"'
-0 1'--
~
0- O
...N
NN
NN
NN
NN
NN
I'1I'1I'1
008
°000
..;t1J"\ 001'11'1
0-0-,...,..., IJ"\'
.0' .0'
.0' .0'
,...' ,...'
1O1'-0tt\N
~.-
1O~
0-I'-0-.01"1
1"1 N
tt\
N
1"1 N
00001"1000-.0
1'-1'-10 N
N10
0 1'-
1'- 0-
.0 .0
.0' 10' 1'-' 1'-' 0-'
0' O
9
,.,~
N
O
~1t1.-
000"'
"' -0
CX
) CX
) 0
0' 0'
0'N
NN
o o
oooj
ooj 0-
OO
W
.-' .-'
0-'""
"" N
«I &
I\ ~
0- «I
"' 0-
"' &
I\ O
«I -0
.-"' «I
.-~~
"',
N
~Ln'...I'")-o
~I'")LnC
X!.-
-oCX
!.-OI'")
.-Ln.-~
.-
LnLnLn~-o
-o-o-oCX
!o-~
~~
1'")~
~'...o-oC
X!
~Ln'...o-C
X!
0.-
00000o-o-0-~
1'")C
X)C
X)C
X)N
Ln
o-~
o-~
o-~
CX
)~
o~N
N
N
N
"'
00000000
"" 0
CX
I C
XI
CX
I ~
C
XI
,,--' ,,--'
,,--' ,,--'
,,--'
.." I--N
P'1",N
CO
-0
.-"'-ON
.-
0008
°N
NN
N
-0-0-0.--0""'
o-o-0-~o-
-o-o-o~-o
", "'
"' ",
"'
..;rO.O
.ON
0- .0
.0 1'-
0-.-«J
0- ;r
N",.-
00000-0
-0 -0
OJ\
0-0000-..;t
;3
~ 0-~"'~U'\"'0--0"' ~..0H" ~~,On.J 0I')""',0"'
-00-~.-r-~900,"-
ON.0,...
;g
CX
J
-.1
8o.--.1,..,0-"'1'-N.-~+ 0~"'~", "1 0"'-0O~.- «>.-"'1'-'~(1!-'cu...O1-aI...tocu~"
I'\~N
N
.-"'~
"'
I/\
N
N
-01/\1/\...'.
0'...' """0
CX
)CX
)Or'-.~
r'-.OO
r'-.1L\
N-O
r'-."'"'
0' 0'
0' «)'
0'N
NN
.-N
-
0......cO n.
Q)
'-e
1010
....c
U01
Q)
C
.c.~
'- cn
Q)
....c
Ccn
~Q.
....~
't-
10 O
'-cn OC
10 ~
U" >
-
Q)
cn~
CQ
)C
O
10" II
-,O
't-oQ) '-
X
O
.~.c
~cn
CQ)
C
Q.
10 Q
..~
C
~
~10C
U10U
...'-
Q)
10.c
.c...u
'-Q)Q
)O
Q
) '-
't- cn
10...
cn .U
cn C
X)
Q)
10 C
X)
.ce
0-0
.-
.D
, 10
c~
cn
C
"10
01" 10
C
cn
Q)
Q)
'- ,
C10
...CC
0
~
.~
...Q
) 0
.D
n. U.~
Q.
10 '-
-' C
...
Q)Q
)Q)
~
-' e
10 ~
't- ~
0 01
cn10
cncn
E:
10Q
) e
...010
X
.~e
0 ID
.~
'-...Q
. II
cn Q
.W
10
ID
.0.,~.0tU~
.-'0
110
QI
+-'
...°~
1-Q
I/)~, .c
...' C
I
~ ~:f
U
Q
10E
:z0.
~IU...O1-1-~< ~ >
-...C/)c4!
c::0O-J.~t;1>-1
~I
~ >,
....,cGJO~C
I.~:r.".~
I
El~IGJ
~j
2:0
QO
Ix
.1=:
n EO...o...IXO"1 10
'"'"
10"-
~cc
~
III ~
QJ
III"-
0!:...~
10 ~
GJ
10...r;
~
~
0...oolx
*' EO+-' Q
+-'IX
0M
.0.c,ID
IX
.0 ~
., .0
Lo .c
4 ~ II!.c[C
IX
10 ~
GJ
10'-
.c6
~ m~ "'oWu2In
-0-0
0-0-~
~
-01nO
~
N
0""
0::8"'
,.,0-N
U
'\
.-"'
.-0-.-0-oU
'\"'
." .".
&r\
-0 r--
«) 0-
O
.-N
." .".
&r\
-0 r--
«) 0-
O
.-N"'
"' "'
"' "'
"' "'
-.1" 1"
~
-.1" ~
&
r\ &
r\ &
r\
88",' ",'
00gg",' ",'
, ..;r
"""
~
~...,
.",.
~
..".
N
..".U
'\«>
000- 0-
~
~
0' 0'
NN
00""""~
~
0' 0'
"' "'
«1",' 0-«1«1-0«1
.-N~
N'...'..."'.-0-~
....-~
N~
"'«)N
O
rt1 0-
0- C
X) 0
~
.-"' 1--
«i N
ci
~
0: 0:
.-0: .--0
.-N
"' 0
rt1 0-
CX
) "'
0-"'
~
O~
0"0"CX
)00"-0 0"1'-
CX
) 0"
O
""' 0"
CX
) .-C
X)
~
-0N
"'
O
.-1'- 1'-
""' 0"
N
,,",
88888888~~
CX
) "'
-n' ",'
-n' ",'
-n' -n'
-n' -n'
",' ",'
00000000000-
0- 0-
0- 0-
0- 0-
0- II\
.0000000001'-N
",' n-'
",' ",'
n-' ",'
n-' n-'
"1' ro'
CX
I ...1'-
"' -0
0-...N
~N
I'--oCX
IOC
XI"''.N
"'..'.I'-N
"""""""""""""""
-.1" -.1"
-.1" -.1"
-.1" -.1"
-.1" -.1"
N
O~
-':~-':~
-':~~
-':~
.00-~~NO O"'~~,ON O0'\I/\,0"'
-10
O-~
"'~00-0-0N
.-
00 00
0-0-0-0-~
~
~
~
0' 0'
0' 0'
NN
NN
..;tt'l1'-CO
01'-
0000
~!"2~
!"2
-0-0-0-0
0-0-0-0--4"
-4" -4"
-4"
10 N
N
",""n
.-~~
o-.-",""'0
, ,
, ,
0~"'O...,
-0"'~.-1'-- ~"'t\.I
1'1 ,...
,... -4"
(X)
.-1'1
.0 1'1
.0 .0
O
I'1N
..;r~I'-O
l'-o-
N
1'-1'-~1'-1'-
I'-N
NO
11'.-
000000..q
..q ..q
..q ..q
..qU
'\U'\U
'\U'\U
'\U'\
"""""""""""""""
000000-0-0-0-0-0-0N
NN
NN
N
I')' I')'
«I' I')'
I')' «I'
-0 r--
0- r--
.-«>.--0.-""".-
.-N.-
11\11\11\11\11\11\
000000~
~~
~~
~
O-.-I'-0~
«>-o1'-1'-~
"'1'-"'
N
~
"' "'
~...'.,
O".
CX
)pt1",pt1.-N
"' .
o o
0C
--o-o-4"
/'- /'-
0001"1
-C
-C"'1"11"1
0' .-'
.-'1"1
...r ...r
-C1'--1'--
o-NN
~
O
O.-N
N
~
1'-- "'
aoo
O
\J .-
-o~o
0-0-
",,«I1'
'.
CX
)
~O.n ~.,-
.0U'\
N .-.-"""~ ~ro"..N.""'0--D00"'
~-0-0,
.- ro0-"""'-'10+-'O1-U+-'..10.&
:u
0' 0'
0' 0'
"""""""'
O
r--' r--'
""' "'
""'
-
0......c.~
~
O
4J11.
L.1010
...C
U
4J 4J
-' .c
10 "0
'"
C1
...10
C~
~Q
.....~
'+
-10
OL.
"' OC
10 '""'
u:I >
,"-
....~
4J '"
"0 C4J
C
Q10:I-,
Q'+
-04JL.0.c'"C10.~
~
"0 X
10 .~
C
"010
CU
4JQ.
4J Q
..c
<...'""'
L. Q
4J
0 L.
'+-
...10L.
...,"IO
U'"
.c 4J
IOU
.Ce0
4J.~
4J
.Q
"' 10
.c"0
.C
IX
)10
IX)
0- ~
10 .-'"
4J 4J
L. ,
C10
...C'"
0"0
:I ...
4J C
1.Q
:lu<
.~
Q.
L.-'
, ...
4J4J4J~
>
e
o '""'
'+-
U0
'"4J
'","ulO4J
C
e...4J
010
"0 .~
e .~
ID
>...0'"
L.
f'-GI
~.010
01
"' G
1 ...
'- °
G1
1-Z
c~
I~
...01
cn .~
>,X
UO...U C
"'~
30;
-'10...0'->
-....~(/)c:0>
o~O-'~~I
~I
>..
<I)
c41o.c01.~%<
I),
.~~I
1-~< Q
~'-,
EO+' Q
+'
IXO~
10.cC
DIx
10 ~
GI
10Lo
.cd
~
10 ~
QJ
10...L:cr
~
"' ~
«I "'
'- .c
<
~
10 ~
aJ 10
'- .c
<
~
O...QO
IX
.I::n EO..,
C..,
IX0".,
10.caJ
IX
10 ~
QJ
10L
.c<
~
0c
O
IX.c,.,
[[1/11
rn~..,O-'..,
No-r ..
-C"'N
..'-.,-
Ir1 .,-
O
N-o
..;r..;r
~
~
.-
000 0
-oI\JI\JI\JI\J C
X)'
0' 0'
0'
---
0"1-00..;r
1"1 "I
.-
~-0-0-o
00-0-0-~
~
No-",O
.0 "'
N
"'.-~
-n
~
..t U
'\ .0
"'" 10
0- O
...N
~
..t
U'\
.0 "'"
10 0-
OU
'\U'\U
'\U'\U
'\U'\U
'\~~
~~
~~
~~
~~
"'"
~
~
~
~""
1'- 1'-
I'-",'
",' ",'
",'
00-0-0!'-!'-
!'-' !'-'
!', "I
o o
-0 -0
I') I')
, ,
~
~
1'-- 1'--I\J
I\J0
OI\J
I\J
O
-.t~
0
I')~0«1N
11 -
-0 cx>
I"
-0 1'--
0- ..;t
-0 ...0-
..;t 1'--
CX
>1"C
X>
0- .1"..;tN
I"I'--0.....cx>
...
N
I'--N
NI"N
N
"' -0
...o o
«>
"' r-.
...0- "'
1'"1
~...oN
:
:N-o"".;oo
"' r-.
0- 1'"1 ...r-.
~
N
1'"1 r-. ~
...
...0-, -0, N
. N
N
1'"1 N, 1'"1, ~
NN
...
000000000000-0
-0 -0
-0 -0
0- 0-
0- 0-
0- 0-
0-1'-
1'- 1'-
1'- N
""
0- 0-
0- 0-
0- 0-
1'-' 1'-'
1'-' 1'-'
0-' 0'
..;t' ..;t'
..;t' ..;t'
..;t' ..;t'
NN
NN
N""""""""""""""
1'1 0- 0-
.-"' r-
r- «)
r- "'
0- -0
N
O
..i O
8...;
O
riI "'
..i riI
,...:(\J
r- -0
~
r- "'
CX
I .-"'
0- r-
~
(\J (\J
"' "
(\J(\J.- r-
r- r-
r- -0
0- "'
~
~
~
~
~
N
(\J N
N
..;
...: "'
..i ..i
..i ..i
..i0000.-(\J"'1'11'11'11'11'1(\J(\J(\J(\J(\J(\J(\J-o-o-o-o-o
.-"' (\J
~
0 0-
0- ~
~
(\J
0- C
XI
.-1'1 1'1
0 1'1
(\J "'
.-1'1 C
XI
CX
I (\J
(\J 0
"'
000000000000-0
-0 -0
-0 0
CX
I ~
-0
-0 -0
-0 -0
1'1 1'1
1'1 1'1
-0 (\J
.-~
~
~
~
~'
' 1'1' ",'
(\J' 0-' 0-'
0-' 0-'
0-'~
~
~
~
~
~
"'
(\J (\J
(\J (\J
(\J.,-
.,- .,-
.,- .,-
0000000000000-
0- 0-
0- 0-
0- 0-
0- 0-
0- 0-
0-r--
r-- r--
r-- r--
r-- r--
r-- r--
r-- r--
r--
-n' -n'
-n' -n'
-n' -n'
-n' -n'
",' -n'
.a' .a'
000000000000N
NN
NN
NN
NN
NN
N
-0 -0
-0 -0
-0 -0
-0 o-o-o-o-o-o-o-lt'Ilt'Ilt'Ilt'Ilt'I
-.t-.t-.t-.t-.t-.t-.tNN
NN
N
"' ,... ""
0. "'
.--,... 0.
~
~
-0 -0
,... 8"'
.--0 ~
.-,...
0. 0.
.-"'""
"'0 oo.""
~""
-3 -3
"'
0 0 C()
~O.-,
.-.-II\,.,r--"'~,...
"'"' «I1'-.-.-~~
~
....0"'...,
or--C
X)
0.,C
X)
In~10+
"'O1-Q+"'t-10
~'-'
0' 0'
0' 0'
O
O
0' .;;
",' ",'
",' ",'
NN
NN
N
ll\1"1.-NO
OI1\N
I'--I'--I1\0-1'--
O
II\ 0-
..;r .-«I
.-0- 0-
.-«I
N
o-«1N
-
0...GI>0
,...u
GI
L.G
I m
(.) ...
c (.)
GI
GI
"t) .r;
.~ >
.,
0 ...
L. C
~
~c.....~
-.-
m
0L.
(/) oC
m
~(.)~
>
-
GI
.:;;"t)
CGI
C
Qm~
II
-,Q
-.-0GIL.0.r;.,Cm
"t)m ,...
C
Xm
.~
U
"t)CG
I G
I.r;
a....a.
<L.
~
GI
0 L.
-.-UJ
m....,
...(.).'L.G
Im
m
.c
e .r;
0 u
.DG
ImII
.r;"t)
(/)Cm
.
CX
) ,...
m
CX
) .,
II 0-
IIL.
.-Cm
C
, 0
"t) II
.,.D
~
(.)
C)
.~c.
~
L.-'
<
..,G
I G
I~
,
e~
~
-.-GI
011.'L.
.,.,
u m
II e
...C
0m
m
.~
e L.
m.~
.D'
., m
UJ
3 m
(X)
~.c101-
.-'0
110
L °
~II-
-'10...O1->-
"'cIIQ
.cCI
:1:
.~~
IIL~I>..+
'.~I/)c"Q::tO--1I/).~
,
+,1
1/)1
~I
21..I"I~
!
1-~< w
~,"'
10 "
41 10
'- .r.
...~O...Q0
ix.&
:"- eo
O,XO""
10.£:0,ID
ix
10 ~
GI
10Lo
.£:0...~
ItI ~
Q)
ItI'-
.1::<
~
10 ~
GJ
10'-
.c<
~
EO...Q...IX0a10..,
oO
IX
.$:;""
..,In1\1.caJIX
1\1 ~
QJ
1\1...c
d ~ "'..-u2..,
~0-"'...o
"""0-~~
"' ."
"' -~
~N
~o-o-
~"'.-N
.-"'°o-O
~'...~
'D~
N,
,
000001'--1'--1'--1'--1'--0-0-0-0-0-
""'"""""""'00000
0-0-~0'...
~N
~O
.-C
X)
0 0
,... 0-
~~
.-N~
~
~~
~~
~
0-0-0-0-0-00000~
~~
~~
~~
,...""CX
).-0-.-~
.-
.-NN
000000-0-0-0-0-1'--
1'-- 1'--
1'-- 1'--
.0' .0'
.0' .0'
.0'
00000N
NN
NN
""'IJ'\IJ'\IJ'\IJ'\IJ'\N
NN
NN
tI'ItI'ItI'ItI'ItI'I1'11'11'11'11'1
o-"'o-o-t\i
CX
)8
"'t\lO~
C
X)
"' "'
.,- ...,
", -.,-
00008
0000
.-N'.'~
"'
CX
) -.2"
-.2" U
'\ -0
P')
-.2" C
X)
0-U
'\U'\C
X)P
')-oNN
ON
T
"'.0"'N.-O
.O~
.O-ci0.:N
N.';0.:0.:.-cxi
"' .-O
"'
.0 .0
.-"' N
NN
NN
""""'
o o
o 0
'...0000000-00
00 00
, ,
, ,
"" o
o o
0000000
""""""0--0-0-000-0-0-""""""
11' .0
1'-- 11'
.-~1'--~
~..;rN
I1'
000000000N
N
1'1
1'1 1'1
1'11'11'11'1,
.0 .0
.0 .0
.0 .0
.0"""C
()C()C
()C()C
()C()C
()N
No-o-o-o-o-o-o-
CO
O-oI'-.-0--O
O-O
~-0~
~"'-oo-coco
1'-I'-I'-0--0-0.-CO
N""
"' '..'
000 0
0-0-0-0.-C
X)
CX
) C
X)
CX
)' .-'
.-' .-'
"' "'
"' ~
,
O
.-N0-
0 .-N
P
'1 ~
""'
-0 ,...
IX>
0-
0 0
0IX
>
0-0-0-0-0-0-0-0-0-0
000000000o-0-~
~~
~~
~~
!'-!'-~~
~~
~~
~""""'
-o-o"""""""""U"\
12
00000,OIr'
"'-00-"'.-", .-~N~CX)
.-"-1
I"\
-00-1"\
-0 ~""~O-0
-
0..."«J «J
«J '-
'- IU
U
U
C
«JIU
.c
'- ,
.tJ c/I
-' IU
C
3 ~a.
.~
...
IU
O'-
"' OC
IU
~
U~
>,
«J c/I
"C
C«JC
O
IU~ II
-,
0...O«J'-0.cc/I
CIU
"CIUCIU
U«J "
.c X~"C
'- C
«J
0 «J
'-
...a. IU
a. c/I
<
U
c/I ~
«J
IU
.ce
...
0.~ .tJ
'- IU
IU
.c
"C.cC
U
IU
«J ~
IU
«J c/I
«J "'
«J
'- C
IU
.C
CX
) 0
"C
CX
) «J
0-
.tJ .-U
a. ,
'--' «J
c/I «J
...~
e
O)~
...~0 <
c/IU
Ic/I
, m
«J «J
e
c/I 0
IU
0 .~
e 0
aJ
I
0
c/I -'
W
U
aJ
0-41~.0C
U1-
"0..
.~I
.I:'"'
01..
1.~
>-.1:
~I"O
E
33
.-"'01
1\10>
1 Lo
00>
1-
Z,
>-
+'
.~.,c41"0.c0),~:t:I'�1~ ~10O1-...~<
...:1:u >
-....~"'c4!"t)3O.J"'.~
I
~I~I
,.,
10 "
QJ
10...0!:
<0:
'-'
10 ~
II 10
..0.:<
'-'
0+' Q
OIX
.r:"'- EO+-' C
+-'
I X
Orn
cu.caJ
I )(
CU
~
41 C
U'-
.c<
~
0Q
OIX
.c0..
EO+" o
+"1
XOm
m.caIIx
m
~«I
m'-
.c<
..."
ID
~~
ID
'- .c
d ~ C
Q1!1
m~u.?rn
P'1P
'1P'1P
'1-0-0-0-00-0-0-0-P
'1 P
'1 P
'1 P
'1
"'0-01'-
CX
IOO
~.--001'--0
CX
I C
XI
"'
C(I
C(I
-0 1'1
0-
r--C(lO
-oO-oI'1N
C(lr--
~r--",O
, ,
, ,
.-1'11'1 N
"' ~
"'
.0 r--
00 0-
0 ...N
"'
~
"'0000000
00000.o.oI"1N
N.0
1'-- .0
"' "'
",' ",'
.0' .0'
.0'
-000.--00-
0-000.-0
-.;tr--N.-r--
0000r--1n,
, ,
NN
.-
-0-.;tNN
N
N-.;tO
0000O
ON
00r--,..,0-0
NIn-or---0
oor--NN
N
N
.-
00000-.;t
.-,.., N
N
0-'..'1n 0'
.-' -.;t'
-.;t' -.;t'
"I ~
I "'
~.
,,!
o o
o 0
-D-D
-D-D
CX
JCX
JCX
JCX
J,
, ,
,
"' ".
~, ".
9 1r\10!'-I')
-D
I')
~
I')
~P
'1-O
N-O
N.-LnP
'1p'1.-
-13
o o
o..,.
..,. ..,.
P"\P
"\P"\
, ,
,~
~
~
IXIIX
IIXI
""""'1'-.1'-.1'-.
' ,
,0-0-0----
.0 C
X)
r-ooN
.0
,.,
..., "'
...
(X)
(X)
(X)
-4:1 -4:1
-4:1""
"" ""
~"'-o
N-O
N"'
"'-.3: ~
...
"' .0
0",
N.-.n
.-"- N
""".-N
"--0
"' -",
o o
01'-
1'- N
-0-01'-
it\' it\'
-.1"'
000roro-D0-
0- ..;r
0' 0'
I'-'N
N
...
CX
J CX
J .0
NN
"'O
OC
XJ
~
~
.-I--N
1'"\ 1'"\
.-N
I---D
N.-"'
1'- ~
~ «1 "'-0"'"'
"'N
'-o,...«)~
~
...
OI\J1'-
-3 ~~,1'-.- ,...-~ CX
I
.0"'"'-0"'m
0(X)
(X)
OIr1
~-00-~~... I/\I/\"'~00.-"1
~"' 0-0000-"' ~ 0 ~-0"'~,"'~0N"'0.,-'mO1-Lom~ 1'-1'-"'"' N,..0-2"~0...
...,.0-"'.- N~0'-<
0,~N ,...0- 1'-"' 0CX
)..,..-0"'..,..00.-",...I<
~0-"'
I...~...10
"'.c
4!U
-".0
"' 10
-" 1-
(0 ,
...~ -0.-1'-1(1"' CX
I"'"'~"'N
.0 ,...
N
""""
~
0 ~
~
N
"' .,-
~~
"'~o
IXI'
D
N-D
o-0-~~
~N
.-
000 0
0000000000000000
0' 0'
0' O
m
m
m
m
,... 0-
-0.--0
"'
""' ..."'
-
14 -
Table 10: Summary of biomass and kelp bed area estimates bychart area (see Figure 1) and bed type for theCanadian shore of Juan de Fuca Strait.
Biomass(metrictonnes)
Area(hectares)Chart Geographic area
611
3339
ABCDEF
Low Density NereocystisRace Passage to Sooke In.Sooke In. to Sheringham Pt.Sheringham Pt. to Magdalena Pt.Magdalena Pt. to Providence c.Providence C. to Walbran Cr.Walbran Cr. to Clo-oose
High Density NereocystisRace Passage to Sooke In.Sooke In. to Sheringham Pt.Sheringham Pt. to Magdalena Pt.Magdalena Pt. to Providence C.Providence C. to Walbran Cr.Walbran Cr. to Clo-oose
19.041.3
4.033.6
8.77.3
ABCDEF
.05.04.4
.0
.0
.1
ABCDEF
Low and High Density MacrocystisRace Passage to Sooke In. N/ASooke In. to Sheringham Pt. N/ASheringham Pt. to Magdalena Pt. N/AMagdalena Pt. to Providence C. N/AProvidence C. to Walbran Cr. N/AWalbran Cr. to Clo-oose N/A
Totals by speciesNereocystis 50,148Macrocystis N/A
501.19.5
2,5124,7521,6762,1183,948
10,247
9.63.18.73.76.35.8
2,8845,253
6028,9874,2602,909
-
UJ
.rl.j.JUJ.j.JQ)
~.c=
UQ
).j.JO
U
Q)
~
~~
Q
) O
Q)
~4-1
ZU
J U
J:>
IQ)U
J.j.J:>
IU.rl
.rl E
UJtJ10
~
.rlQ
)~.O
'OE
~
UJ
.c=
.-t .rl
.
tJ10.j.JIU.rl
U
UJ
Q)
.c=
~.j.JU
IU-aU
JO~
1UQ
):>1
1U.-t~
~Q
) O
~Q
)Z.j.J
O.C
=
~.-tE
-t.-tQ)IU
:>
4-1..j.J~O
.j.J O
.rl
~.j.J
~IU
.j.J
O
.c=
Q) .rl
.rl U
.c=
IU
.j.J .j.J~
.rl .c=
.j.J
UJU
4-1CI)
O
IU
O~
Q)
IUE
~
U
O
~
O
~U
.rl
.rl ~
.j.J.j.J
UJ .rl
Q)
~U
JUJ'O
Q)IU
OuE
~~
~O
EIU
Q) .rl
O
~~
.OU
I-)
Q)
.-t.0IUF-t
G>
'O.r::~
~E
-tO~
4-1
t/)+.J~
..-I ~
G
>+
.J~~
t/).r::~uu
'0o.r::G
>G
>U
.o~
~
G>
G>
.-1z
~~
+.J
:>t..-1
O+
.J +
.J..-I
~t/)G
>4-1
~
~
O
.G
> ~
~
'0 ~
G>
G>
O
~.r::
U
..-I ~
O'~
+.J
..-I 4-1 ..-I
:>t
.r::~t/)~~
O
O
'Ot/)~
+.J
~
E
~~
4-10G>
O
u >
~
~O
t/)
+.J ..-I
.-IG>
~+.JG>
+.J
4-1 ~
u
..-IO
E
~
~
..-I G
> ~
~
+.J
~+
.Jot/)
C/)
..-I G
> t/)
+.J
G>
~..-I
'0 >
U
t/) G
> ..-I
~0.0
O'~
~Et/)~
G>
O ..-I
E
'0U
+.J~t/).-1~
+.J
O~
~
U
U
~G
>O
/"j
U~
+.J
~
U
t/) G
>G
>~
~.r::
P.~
.-1+.J
N.-IG>.-I.0~E-t
15
-
-16 -
DISCUSSION
The largest kelp beds were located at Muir Point, Orveas Bay,
Sa11brio Point and Bonilla Point. Within the inventory area, 55% of the
kelp bed area and 55% of the estimated Nereo~stis bianass lie in these
beds .The primary canopy forming kelp was Nereo~stis, but small areas
of Macro~stis were located at Muir Point and in the vicinity of San
Si1ron Point .
NarrON fringe beds follON the majority of the shoreline fran Race
Passage to Sooke Inlet. With the exception of the large Muir Point and
Orveas Bay beds , only occasional small beds are found between SCX)ke
Inlet and Sa11brio Point. Westward fran the large Sanbrio Point bed, a
narrON fringe again follONs the shore as far as CUllite Creek. Fran
CUllite Creek to the end of the inventory area near Clo-oose, occasional
beds, of which the Bonilla Point bed is the largest, extend out fran
shore.
-
17
REFERENCES
Coon, L.M. 1977. Marine Plant Manage:nent program in BritishColumbia. A paper presented at the IXth International Seaweed~sium at Santa Barbara, California, U.S.A. in August, 1977(Mimeo) .17 pp.
Coon, L.M.data.
1981. Porcher Island. UnpublishedKelp Inventory, 1981.
L.M. , E.J. Field and Canadian Benthic Ltd. 1977. Nootka SoundKelp Inventory , 1975. British Columbia Marine Resources Branch,Fish. Managanent Rep. No.2 (2nd Edition) .27 pp.
L.M., W.G. Roland, E.J. Field, and W.E.L. Clayton. 1979. KelpInventory , 1976, Part 3. North and West Coasts Graham Island(Q.C.I. ) .British Oolumbia Marine Resources Branch, Fish.Managenent Rep. No.13. 26 pp.
Ooon, L.M., W.G. Roland, E.J. Field, W.E.L. Clayton, v. Jenson. 1980Kelp Inventory , 1976, Part 4. Goschen Island to the Tree KnobGroup. British Columbia Marine Resources Branch, Fish.Managanent Rep. No.19 .18 pp.
Goon, L.M., W.G. Roland, E.J. Field, W.E.L. Clayton, V. Jenson. 1981.Kelp Inventory , 1976, Part 5. North Vanoouver Island, Hope ,Nigei and Balaklava Islands .British Columbia Marine ResourcesBranch, Fish. Management Rep. No.20. 20 pp.
Goon, L.M., W.G. Roland, I.R. Sutherland, R.A. Hall. 1982. KelpInventory , 1978, Northwest Ooast of Vancouver Island. BritishColumbia Marine Resources Branch, Fisheries DeveloJ;:lnent ReportNo.28. 16pp.
Field, E.J. and E.A.C. Clark. 1978. Kelp Inventory, 1976, Part 2.The Dundas Group. British Columbia Marine Resources Branch,Fish Management Rep. No.11. 21 pp.
Field, E.J., L.M. Coon, W.E.L. Clayton and E.A.C. Clark. 1977. KelpInventory , 1976, Part 1. The Estevan Group and Campania Island.British Oolumbia Marine Resources Branch, Fish.Managenent Rep. No.9. 19 pp.
Foreman, R.E. , 1975. KIM-1. A method for inventory of floating kelpsand its application to selected areas of Kelp Licence Area 12.
Benthic ECXJlogical Research Program RefXJrt 75-1. RefXJrt toFederal Fisheries and Marine Service and Provincial MarineResources Branch. 81 pp .
Foretlan, R.E., 1979.Nereo~stis.16 pp.
Supplanentary Evaluation of the Kll1-1 Method forBenthic Eoological Research program Ref:X:)rt 79-2.
-
18
APPENDIX
Charts A through F are enclosed in the following envelope .