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15.1 Darwin’s Theory of Natural Selection
Darwin on the HMS Beagle
Evolution
Darwin’s role on the ship was as naturalist and companion to the captain.
His job was to collect biological and geological specimens during the ship’s travel.
Chapter 15
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The Galápagos Islands
Evolution
Darwin began to collect mockingbirds, finches, and other animals on the four islands.
He noticed that the different islands seemed to have their own, slightly different varieties of animals.
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Chapter 15
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Almost every specimen that Darwin had collected on the islands was new to European scientists.
Evolution
Populations from the mainland changed after reaching the Galápagos.
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Chapter 15
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Darwin Continued His Studies
Evolution
Darwin hypothesized that new species could appear gradually through small changes in ancestral species.
Darwin inferred that if humans could change species by artificial selection, then perhaps the same process could work in nature.
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Chapter 15
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Natural Selection
Evolution
Individuals in a population show variations.
Variations can be inherited.
Organisms have more offspring than can survive on available resources.
Variations that increase reproductive success will have a greater chance of being passed on.
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Chapter 15
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Darwin’s theory of natural selection is not synonymous with evolution.
The Origin of Species
Evolution
Darwin published On the Origin of Species by Means of Natural Selection in 1859.
It is a means of explaining how evolution works.
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Chapter 15
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15.2 Evidence of Evolution
Support for Evolution
Evolution
The fossil record
Glyptodont
Fossils provide a record of species that lived long ago.
Fossils show that ancient species share similarities with species that now live on Earth.
Armadillo
Chapter 15
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Evolution
Derived traits are newly evolved features, such as feathers, that do not appear in the fossils of common ancestors.
Ancestral traits are more primitive features, such as teeth and tails, that do appear in ancestral forms.
Anatomically similar structures inherited from a common ancestor are called homologous structures.
15.2 Evidence of Evolution
Chapter 15
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Evolution
Vestigial Structures
Structures that are the reduced forms of functional structures in other organisms.
15.2 Evidence of Evolution
Chapter 15
Evolutionary theorypredicts that features of ancestors that no longer have a function for that species will become smaller over time until they are lost.
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Evolution
Show that functionally similar features can evolve independently in similar environments
Analogous structures can be used for the same purpose and can be superficially similar in construction, but are notinherited from a common ancestor.
15.2 Evidence of Evolution
Chapter 15
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Evolution
Vertebrate embryos exhibit homologous structures during certain phases of development but become totally different structures in the adult forms.
Comparative Embryology
15.2 Evidence of Evolution
Chapter 15
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Evolution
Common ancestry can be seen in the complex metabolic molecules that many different organisms share.
Comparative Biochemistry
15.2 Evidence of Evolution
Chapter 15
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Evolution
Comparisons of the similarities in these molecules across species reflect evolutionary patterns seen in comparative anatomy and in the fossil record.
Organisms with closely related morphological features have more closely related molecular features.
15.2 Evidence of Evolution
Chapter 15
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Review
• Explain how mimicry and camouflage help species survive.
• How do homologous structures provide evidence for evolution?
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Geographic Distribution
Evolution
The distribution of plants and animals that Darwin saw first suggested evolution to Darwin.
Rabbit Mara
15.2 Mechanisms of Evolution
Chapter 15
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Evolution
Patterns of migration were critical to Darwin when he was developing his theory.
Evolution is intimately linked with climate and geological forces.
15.2 Evidence of Evolution
Chapter 15
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Evolution
Types of Adaptation
An adaptation is a trait shaped by natural selection that increases an organism’s reproductive success.
Fitness is a measure of the relative contribution an individual trait makes to the next generation.
15.2 Evidence of Evolution
Chapter 15
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Evolution
Camouflage
Allows organisms to become almost invisible to predators
Leafy sea dragon
15.2 Evidence of Evolution
Chapter 15
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Evolution
Mimicry
One species evolves to resemble another species.
Western coral snake California kingsnake
15.2 Evidence of Evolution
Chapter 15
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Evolution
Consequences of Adaptations
Some features of an organism might be consequences of other evolved characteristics.
They do not increase reproductive success.
Features likely arose as an unavoidable consequence of prior evolutionary change.
15.2 Evidence of Evolution
Chapter 15
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Populations, not individuals, evolve
• Gene pool – all of the alleles in a population’s genes
• Allelic Frequency – the percentage of any specific allele in the gene pool
• Genetic Equilibrium – a population in which the frequency of alleles remains the same over generations
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Evolution
Genetic Drift
A change in the allelic frequencies in a population that is due to chance
In smaller populations, the effects of genetic drift become more pronounced, and the chance of losing an allele becomes greater.
15.3 Shaping Evolutionary Theory
Chapter 15
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Evolution
Gene Flow
Increases genetic variation within a population and reduces differences between populations
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Chapter 15
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Evolution
Natural Selection
Acts to select the individuals that are best adapted for survival and reproduction
15.3 Shaping Evolutionary Theory
Chapter 15
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Evolution
Stabilizing selection operates to eliminate extreme expressions of a trait when the average expression leads to higher fitness.
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Chapter 15
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Evolution
Directional selection makes an organism more fit.
15.3 Shaping Evolutionary Theory
Chapter 15
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Evolution
Disruptive selection is a process that splits a population into two groups. It tends to eliminate the intermediate phenotypes.
15.3 Shaping Evolutionary Theory
Chapter 15
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Evolution
Sexual selection operates in populations where males and females differ significantly in appearance.
Qualities of sexual attractiveness appear to be the opposite of qualities that might enhance survival.
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Chapter 15
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Evolution
Prezygotic isolation prevents reproduction by making fertilization unlikely.
Prevents genotypes from entering a population’s gene pool through geographic, ecological, behavioral, or other differences
Eastern meadowlark and Western meadowlark
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Chapter 15
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Postzygotic isolation occurs when fertilization has occurred but a hybrid offspring cannot develop or reproduce.
Evolution
Prevents offspring survival or reproduction
Liger
15.3 Shaping Evolutionary Theory
Chapter 15
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Physiological Adaptations
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Evolution
Geographic Isolation
A physical barrier divides one population into two or more populations.
Abert squirrel Kaibab squirrel
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Chapter 15
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Evolution
Adaptive Radiation
Follows large-scale extinction events
Can occur in a relatively short time when one speciesgives rise to many different species in response to the creation of new habitat
Chapter 15
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Evolution
Unrelated species evolve similar traits even though they live in different parts of the world.
Convergent Evolution
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Chapter 15
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Divergent Evolution
• A type of adaptive radiation
• Species that once were similar to an ancestral (older) species diverge
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Evolution
Rate of Speciation
Evolution proceeds in small, gradual steps according to a theory called gradualism.
Punctuated equilibrium explains rapid spurts of genetic change causing species to diverge quickly.
15.3 Shaping Evolutionary Theory
Chapter 15
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Evolution
15.3 Shaping Evolutionary Theory
Chapter 15
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Review
• Why is rapid evolutionary change more likely to occur in small populations?
• Explain why the evolution of resistance to antibiotics in bacteria is an example of directional natural selection.