Not to be cited withaIt pgrmi ASicn of the autbors1

canadian Atlantic Fisheries scientific AdvisOJy oanittee

CAFSAC Researcb Dcoment 9015

He pas citer sans autorlsatiat des auteurs1

Cklrdte scientifique CXIlSUltatif des pidles canadiennes dans lAtlantique

cscR Ikament de redlerche 9015

by

HMC DJplis ani CJ MacDcu1all

SCierat Branch GUlf Region Deparbnent of Fisheries amp oceans

Po Ebx 5030 Mcntal New Brunswick

E1C 956

~ series dooDDPnts the scientific basis for fisheries management advice in Atlantic canada As such it addresses the issJeS of the day in the time frames required ani the Researcb J))ounents it contains are not intended as definitive statements on the subjects addressed but rather as progress repgtrts on arJOirg investigations

l cette serie doomente les bases scientif1ques des calSeils de qestion des pamphes sur la c6te atlantique du Canada Qume telle elle cawre les prcigtlemes actuels seloo les edleaxiers voulus et les Dcxunents de red1erche quelle CXll1tient ne doivent pas etre consideres cxmne des ~ finals surles sujets traites mais plutOt came des rAIPJrts detape sur les etudes en cxm-s

Research Dcxunents are prodllCed in Lea IDcuments de red1erche sont the official lanJ1la98 in Which they plblies dans la laDJUe officielle are pxovided to the Secretariat by utUisee par les auteurs dans Ie the author manuscrit erwoye au secretariat

Abstract

The catch-at-age matrices for spring- and fall-spawningherring in NAFO division 4T were revised for the years 1974 to1988 and the procedures used are documented here Commercialsamples were handled by three laboratories at various timesduring this period resulting in inconsistent age or spawning-group assignment between years and laboratories Errors in datafiles were corrected and landings were updated before proceedingwith the catch-at-age calculations Fish sampled in 1985-1988were reassigned to a spawning group by gonadal maturity stage orby visual inspection of otolith characteristics to be consistentwith prior methodology

The new catch-at-age differs slightly from previousmatrices the more notable differences being found at extremeages Despite the spawning-group reassignment applied to someyears the numbers-at-age for those years were not altered to anygreater extent than those of other years

Resume

Les matrices de capture a lage pour les geniteurs printempset automne du hareng de la division 4T de 1OPANO ont etereconstruites pour les annees 1974-1988 et les methodesemployees pour ce faire sont presentees Les echantillons tiresde la peche commerciale ont ete analyses par trois laboratoiresdifferents au cours de cette periode ce qui a cree desdifferences dans la classification du type geniteur et de lageentre les laboratoires et au cours des annees Les erreursdecouvertes dans les fichiers ont ete corrigees et les prises ontete mises a jour avant de proceder a la revision des matricesLe type geniteur des poissons echatillonnes entre 1985 et 1988 aete revise en se servant soit du stade de maturite des gonades oudes caracteres de lotolithe pour mieux se conformer auxmethodes employees par le passe

Les nouvelles matrices different quelque peu desantecedentes notamment pour les ages extremes Les changementsapportes a la methode de classification du type geniteur pour lesannees 1985-1988 ne semblent guere avoir augmente les differencesentre les nouveaux et anciens chiffres de capture a lage

2

Introduction

The CAFSAC Advisory Document 8619 (Anon 1986) recommendedthat the catch- and weight-at-age matrices for 4T herring bereconstructed Several factors were suspected of havingintroduced inconsistencies in the matrices The problems stemmedin part from successive transfers of responsibilities andinformation lack of documentation about laboratory proceduresand differences in year-class and spawning-group assignmentbetween laboratories and years The 1974-1980 commercial samplesfor 4T were processed in St Andrews and aged in St Johns The1981 samples were processed in St Andrews and aged in both StAndrews and St Johns (the final ages on file were from StAndrews) The 1982 samples were processed in St Andrews up toSeptember and from then on in Quebec The St Andrews sampleswere aged by the Gulf Regions ager (transferred from StAndrews) Starting in 1983 4T samples were processed and agedin Moncton and the data files were stored on site Starting in1984 sampling for detailed information switched from random tostratified subsampling (Ahrens 1985b)

Many changes in methodology for calculating weights-at-agetook place between 1971 and 1985 (Ahrens 1985a) From 1971 to1973 weights for 4T herring were derived from samples drawn fromthe winter purse-seine fishery in Subdivision 3Pn From 1974 to1980 weights-at-age were estimated from the spring fishery alongthe Edge of the Laurentian Channel When this fishery endedweights were obtained from spring gillnet samples (1981 and1982) For the 1983 and 1984 fisheries weights were derivedfrom the weighted averages of fish weights in all 4T fisheriesSuch inconsistencies yielded a weight-at-age matrix that did notmatch reported landings when multiplied by the catch-at-agematrix In 1985 the weight matrix was revised by applying theobserved average weights for each year between 1971 and 1976 andby averaging the 1981-to-1984 weights for the period 1977 to1984 In subsequent assessments no mention was made of themethodology used to derive weights-at-age and previous symptomsin the matrices persisted

Furthermore changes in methodology used over the years todetermine the spawning group of sampled individuals added to themounting evidence against the reliability of the matricesStarting with 1974 spawning group was assigned mainly by gonadmaturity using otolith type when maturity stage was ambiguous(Cleary et al 1982) When the two methods gave differentanswers the one that seemed most definite took precedence (thegonad-maturity schedule adopted by CAFSAC was followed but theoverwintering stages were considered less certain and were

3

therefore likely to be discarded in favour of a clear-cutotolith type In 1985 the Gulf Region started usingDiscriminant Analysis of otolith morphometrics which was meantto replace the agers evaluation of otolith type when maturitystage was ambiguous It was felt that using a Discriminant scorewould be an objective way of evaluating otolith morphology notsubject to within- and among- ager variability The region wasasked to document and justify this method Messieh et al (1989)have done so but provide little information on the selection ofthe classification data sets which we know was inconsistent fromyear to year and often included the agers opinion of otolithtype (thus generating tautological classifications) The methodwas to be re-examined and documented here as well but this couldnot be done in the time available and had to be postponedInstead spawning group was reassigned for samples collectedbetween 1985 and 1988 by a method more consistent with previousmethodology namely gonad ripeness in season or alternatelyotolith morphology as evaluated by the ager (see Methods below)

This document outlines the revisions to the previous (ineffect up to 1989) catch- and weight-at-age matrices Unexpectederrors in the data files and in the landings were discovered andwere edited before reconstructing the matrices The effect ofeach revision step on the matrices is being documented with therevised 1989 assessment (Claytor and Dupuis 1990)

Methods

a) Catch data

The landings for particular gear types fishing seasons andunit areas (or cells) that had been used to generate the catch-at-age for the 4T herring assessments up to 1989 rarely added upto the total landings for that year The compilation method wasnot documented in the assessment documents Landing statisticswere therefore recompiled Updated landing statistics for 4Tfrom 1974 to 1982 were drawn from McMillan et al (1984)Quebecs 1982 statistics were not included in this report andwere therefore obtained from Table 5 in the NAFO StatisticalBulletin (NAFO 1984) Starting with 1983 Maritime landingswere compiled from the Gulf Regions purchase slip files towhich Quebec totals were added The latter were obtained eitherfrom Statistical Bullletins (1983-1985) NAFO data tape (1986) orZIFF file (1987) Landings for 1987 and 1988 are preliminary

Landings (round weight in kg) for each gear month and unitarea were pooled into cells defined by gear type (fixed and

4

mobile) fishing season (spring and fall) and aggregatedstatistical unit areas (431-435 436 437-439) for each yearThey were later rounded off to the nearest metric tonne (MT)When the statistical unit area of a landing was unknown (4Tu)as sometimes occurred in the Maritimes or Quebec Regionsreports it was reallocated proportionally to specific 4T pooledareas fished with the same gear and in the same season In 1975and 1977 there were no landings reported for fixed gear duringany month in any specific area but such landings listed under4Tu were considerable They were reallocated to specificpooled areas by calculating for 1974 1976 and 1978 the averagerelative catches in the three areas and applying theseproportions to the catch recorded for 4Tu The averages werebased on only those years because the relative importance of thetwo fishing seasons and gear types shifted in subsequent years

b) Commercial sampling

Data files consisting of random length-frequency samplesfrom which random detailed samples were drawn were copied fromthe CYBER at BSO for 1974 to 1982 except for the years 1978-1981 which were obtained from the St Andrews magnetic tapecollection Data files generated before 1974 cannot be usedbecause the errors missing data and unreliable spawning-groupand age assignments cannot be corrected

Attempts to match length-frequency with detailed biologicaldata revealed pervasive errors and missing information foridentifying samples in these data files Length-frequency andbiological samples were therefore renumbered and obviouskeypunching and coding errors were corrected by checkingphotocopies of data sheets filled out by samplers and laboratorytechnicians For 1980 and 1981 several data sheets had not beenkeypunched or had been deleted and therefore were restoredOnce data files had been reorganized length-weight regressionswere executed to flag outliers Such data points were examinedin concert with gonad maturity and weight and were eitherretained (if the length-weight relationship seemed plausible)edited (if a keypunching error was found) or deleted (if theerror was untraceable)

There have been several changes in the procedures used todetermine age and spawning-group affinity over the years whichreduce consistency across years Thus the reconstruction of thecatch-at-age matrices was based on the method most consistentlyapplicable to all years namely maturity stage if unambiguous(stages 56 and 7 with July 1 delineating the spring and fallspawning seasons) or alternately otolith type as evaluated bythe ager (Cleary et al 1982) This technique was applied to

9

the 1985-1988 samples to regenerate age and spawning-group dataIn 1985 however maturity stages were determined in part byunsupervised temporary helpers with the result that maturitystages seemed unreliable and were deleted from the current filealtogether For 1985 therefore all fish were assigned to aspawning group by the agers evaluation of otolith type whereasnormally 50-80 the fish sampled in a given year get classifiedaccording to maturity stage (weighted average for the 1980sexcluding 1985 is 63)

After editing the data files the length-frequency sampleswere matched to their corresponding biological data to create amaster data file for each spawning group of numbers aged atlength in each sample Samples were then pooled by area fishingseason and gear type (or cells) The corrections outlinedabove usually increased the effective sample sizes butunfortunately did not rectify the sampling deficiency for certaincells (Table 1)

The detailed samples were used to construct age-lengthrelationships specific to each cell (some of the fish that wereused for regressing weight on length could not be aged and weretherefore subsequently excluded from the age-length key) Whencell samples were missing or too small to be reliable (because ofa low r for the length-weight regression or very few lengthgroups represented in the sample) other samples had to besubstituted to construct the age-length key The substitutionsto be applied were determined by the considerations listed below

1 - The unknown spawning-group composition (percentage bynumber) of the landings from one cell was estimated by taking theaverage (weighted by sample size) of the percentages in theequivalent cell for the two most recent and the two subsequentyears available (Table 1) As an example the South SpringMobile catch in 1976 would be estimated by averaging thepercentages observed in the equivalent cell in 1974 1975 1977and 1978 Table 1 indicates however that there was no samplingin 1974 or 1977 limiting the number of most recent cells toone (1975) and shifting the two subsequent available years to1978 and 1979 The percentage estimate is required to adjust thenumbers-at-age calculated with the substitute sample as thelatter is rarely composed of the same proportions of spring andfall spawners as those estimated for the missing sample Weassume that population composition for a cell is similar acrossyears This seems more likely than relying on a population froma different area season or gear For unit area 436 and unitareas 437-439 however the Spring Mobile-gear fishery was neversampled preventing any estimation of spawning-group compositionbased on adjacent years (Table 1) In this instance thepercentage of spring spawners in the catch had to be estimated

C

from the Southern samples collected sporadically over the years

2 - The age-length key for an empty cell is preferablyderived from the samples from another cell in that year ratherthan from the equivalent cell in another year to avoid thepossibility of interdependent age or size shifts between yearsThe possible choices in selecting a substitute cell were

a) the same season and gear in a different areab) the same season and area but different gearc) the same season but a different area and geard) for mobile gear only the same area and gear but a

different season ore) the same season gear and area in a different year

Because fixed and mobile gears usually targetted fish ofdifferent maturity stages within the same area the order ofpreference for these choices was usually as listed above but wasdetermined in each case by comparing the size distributions andthe regression coefficients of length-weight relationships in therelevant cells from better-sampled years The final choice of asubstitute sample was therefore particular to that case and wasaffected by the similarities noted in other years

3 - Partial substitutions were also required when a samplewas available (and could therefore reveal spawning-groupcomposition) but consisted almost exclusively of one group Insuch cases the sample size of the rare spawning-group was ofteninadequate for its age-length determination The decision stepsoutlined in 2) above were followed and another subsample wassubstituted

c) Catch-at-age matrices

Length-weight regressions were computed with the SAS packagefor both spring and fall spawners in each cell The percentagesof spring and fall spawners at each length were also calculatedThe options defining a particular cell (area fishing season andgear type) selected the appropriate samples in the master filewhich along with the corresponding regression parameters totallandings and percentages-at-length were input into the FORTRANprogram AGELEN (Wright 1990) This set of programs calculatesthe catch-at-age from equations found in Gavaris and Gavaris(1983) It generated an age-length key for each spawning groupand projected it onto the landings for that cell Thenumbers-at-age thus obtained were then summed to yieldcorresponding annual totals for fixed mobile and all gears

Average weight-at-age was simultaneously computed by AGELEN

7

for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Introduction

The CAFSAC Advisory Document 8619 (Anon 1986) recommendedthat the catch- and weight-at-age matrices for 4T herring bereconstructed Several factors were suspected of havingintroduced inconsistencies in the matrices The problems stemmedin part from successive transfers of responsibilities andinformation lack of documentation about laboratory proceduresand differences in year-class and spawning-group assignmentbetween laboratories and years The 1974-1980 commercial samplesfor 4T were processed in St Andrews and aged in St Johns The1981 samples were processed in St Andrews and aged in both StAndrews and St Johns (the final ages on file were from StAndrews) The 1982 samples were processed in St Andrews up toSeptember and from then on in Quebec The St Andrews sampleswere aged by the Gulf Regions ager (transferred from StAndrews) Starting in 1983 4T samples were processed and agedin Moncton and the data files were stored on site Starting in1984 sampling for detailed information switched from random tostratified subsampling (Ahrens 1985b)

Many changes in methodology for calculating weights-at-agetook place between 1971 and 1985 (Ahrens 1985a) From 1971 to1973 weights for 4T herring were derived from samples drawn fromthe winter purse-seine fishery in Subdivision 3Pn From 1974 to1980 weights-at-age were estimated from the spring fishery alongthe Edge of the Laurentian Channel When this fishery endedweights were obtained from spring gillnet samples (1981 and1982) For the 1983 and 1984 fisheries weights were derivedfrom the weighted averages of fish weights in all 4T fisheriesSuch inconsistencies yielded a weight-at-age matrix that did notmatch reported landings when multiplied by the catch-at-agematrix In 1985 the weight matrix was revised by applying theobserved average weights for each year between 1971 and 1976 andby averaging the 1981-to-1984 weights for the period 1977 to1984 In subsequent assessments no mention was made of themethodology used to derive weights-at-age and previous symptomsin the matrices persisted

Furthermore changes in methodology used over the years todetermine the spawning group of sampled individuals added to themounting evidence against the reliability of the matricesStarting with 1974 spawning group was assigned mainly by gonadmaturity using otolith type when maturity stage was ambiguous(Cleary et al 1982) When the two methods gave differentanswers the one that seemed most definite took precedence (thegonad-maturity schedule adopted by CAFSAC was followed but theoverwintering stages were considered less certain and were

3

therefore likely to be discarded in favour of a clear-cutotolith type In 1985 the Gulf Region started usingDiscriminant Analysis of otolith morphometrics which was meantto replace the agers evaluation of otolith type when maturitystage was ambiguous It was felt that using a Discriminant scorewould be an objective way of evaluating otolith morphology notsubject to within- and among- ager variability The region wasasked to document and justify this method Messieh et al (1989)have done so but provide little information on the selection ofthe classification data sets which we know was inconsistent fromyear to year and often included the agers opinion of otolithtype (thus generating tautological classifications) The methodwas to be re-examined and documented here as well but this couldnot be done in the time available and had to be postponedInstead spawning group was reassigned for samples collectedbetween 1985 and 1988 by a method more consistent with previousmethodology namely gonad ripeness in season or alternatelyotolith morphology as evaluated by the ager (see Methods below)

This document outlines the revisions to the previous (ineffect up to 1989) catch- and weight-at-age matrices Unexpectederrors in the data files and in the landings were discovered andwere edited before reconstructing the matrices The effect ofeach revision step on the matrices is being documented with therevised 1989 assessment (Claytor and Dupuis 1990)

Methods

a) Catch data

The landings for particular gear types fishing seasons andunit areas (or cells) that had been used to generate the catch-at-age for the 4T herring assessments up to 1989 rarely added upto the total landings for that year The compilation method wasnot documented in the assessment documents Landing statisticswere therefore recompiled Updated landing statistics for 4Tfrom 1974 to 1982 were drawn from McMillan et al (1984)Quebecs 1982 statistics were not included in this report andwere therefore obtained from Table 5 in the NAFO StatisticalBulletin (NAFO 1984) Starting with 1983 Maritime landingswere compiled from the Gulf Regions purchase slip files towhich Quebec totals were added The latter were obtained eitherfrom Statistical Bullletins (1983-1985) NAFO data tape (1986) orZIFF file (1987) Landings for 1987 and 1988 are preliminary

Landings (round weight in kg) for each gear month and unitarea were pooled into cells defined by gear type (fixed and

4

mobile) fishing season (spring and fall) and aggregatedstatistical unit areas (431-435 436 437-439) for each yearThey were later rounded off to the nearest metric tonne (MT)When the statistical unit area of a landing was unknown (4Tu)as sometimes occurred in the Maritimes or Quebec Regionsreports it was reallocated proportionally to specific 4T pooledareas fished with the same gear and in the same season In 1975and 1977 there were no landings reported for fixed gear duringany month in any specific area but such landings listed under4Tu were considerable They were reallocated to specificpooled areas by calculating for 1974 1976 and 1978 the averagerelative catches in the three areas and applying theseproportions to the catch recorded for 4Tu The averages werebased on only those years because the relative importance of thetwo fishing seasons and gear types shifted in subsequent years

b) Commercial sampling

Data files consisting of random length-frequency samplesfrom which random detailed samples were drawn were copied fromthe CYBER at BSO for 1974 to 1982 except for the years 1978-1981 which were obtained from the St Andrews magnetic tapecollection Data files generated before 1974 cannot be usedbecause the errors missing data and unreliable spawning-groupand age assignments cannot be corrected

Attempts to match length-frequency with detailed biologicaldata revealed pervasive errors and missing information foridentifying samples in these data files Length-frequency andbiological samples were therefore renumbered and obviouskeypunching and coding errors were corrected by checkingphotocopies of data sheets filled out by samplers and laboratorytechnicians For 1980 and 1981 several data sheets had not beenkeypunched or had been deleted and therefore were restoredOnce data files had been reorganized length-weight regressionswere executed to flag outliers Such data points were examinedin concert with gonad maturity and weight and were eitherretained (if the length-weight relationship seemed plausible)edited (if a keypunching error was found) or deleted (if theerror was untraceable)

There have been several changes in the procedures used todetermine age and spawning-group affinity over the years whichreduce consistency across years Thus the reconstruction of thecatch-at-age matrices was based on the method most consistentlyapplicable to all years namely maturity stage if unambiguous(stages 56 and 7 with July 1 delineating the spring and fallspawning seasons) or alternately otolith type as evaluated bythe ager (Cleary et al 1982) This technique was applied to

9

the 1985-1988 samples to regenerate age and spawning-group dataIn 1985 however maturity stages were determined in part byunsupervised temporary helpers with the result that maturitystages seemed unreliable and were deleted from the current filealtogether For 1985 therefore all fish were assigned to aspawning group by the agers evaluation of otolith type whereasnormally 50-80 the fish sampled in a given year get classifiedaccording to maturity stage (weighted average for the 1980sexcluding 1985 is 63)

After editing the data files the length-frequency sampleswere matched to their corresponding biological data to create amaster data file for each spawning group of numbers aged atlength in each sample Samples were then pooled by area fishingseason and gear type (or cells) The corrections outlinedabove usually increased the effective sample sizes butunfortunately did not rectify the sampling deficiency for certaincells (Table 1)

The detailed samples were used to construct age-lengthrelationships specific to each cell (some of the fish that wereused for regressing weight on length could not be aged and weretherefore subsequently excluded from the age-length key) Whencell samples were missing or too small to be reliable (because ofa low r for the length-weight regression or very few lengthgroups represented in the sample) other samples had to besubstituted to construct the age-length key The substitutionsto be applied were determined by the considerations listed below

1 - The unknown spawning-group composition (percentage bynumber) of the landings from one cell was estimated by taking theaverage (weighted by sample size) of the percentages in theequivalent cell for the two most recent and the two subsequentyears available (Table 1) As an example the South SpringMobile catch in 1976 would be estimated by averaging thepercentages observed in the equivalent cell in 1974 1975 1977and 1978 Table 1 indicates however that there was no samplingin 1974 or 1977 limiting the number of most recent cells toone (1975) and shifting the two subsequent available years to1978 and 1979 The percentage estimate is required to adjust thenumbers-at-age calculated with the substitute sample as thelatter is rarely composed of the same proportions of spring andfall spawners as those estimated for the missing sample Weassume that population composition for a cell is similar acrossyears This seems more likely than relying on a population froma different area season or gear For unit area 436 and unitareas 437-439 however the Spring Mobile-gear fishery was neversampled preventing any estimation of spawning-group compositionbased on adjacent years (Table 1) In this instance thepercentage of spring spawners in the catch had to be estimated

C

from the Southern samples collected sporadically over the years

2 - The age-length key for an empty cell is preferablyderived from the samples from another cell in that year ratherthan from the equivalent cell in another year to avoid thepossibility of interdependent age or size shifts between yearsThe possible choices in selecting a substitute cell were

a) the same season and gear in a different areab) the same season and area but different gearc) the same season but a different area and geard) for mobile gear only the same area and gear but a

different season ore) the same season gear and area in a different year

Because fixed and mobile gears usually targetted fish ofdifferent maturity stages within the same area the order ofpreference for these choices was usually as listed above but wasdetermined in each case by comparing the size distributions andthe regression coefficients of length-weight relationships in therelevant cells from better-sampled years The final choice of asubstitute sample was therefore particular to that case and wasaffected by the similarities noted in other years

3 - Partial substitutions were also required when a samplewas available (and could therefore reveal spawning-groupcomposition) but consisted almost exclusively of one group Insuch cases the sample size of the rare spawning-group was ofteninadequate for its age-length determination The decision stepsoutlined in 2) above were followed and another subsample wassubstituted

c) Catch-at-age matrices

Length-weight regressions were computed with the SAS packagefor both spring and fall spawners in each cell The percentagesof spring and fall spawners at each length were also calculatedThe options defining a particular cell (area fishing season andgear type) selected the appropriate samples in the master filewhich along with the corresponding regression parameters totallandings and percentages-at-length were input into the FORTRANprogram AGELEN (Wright 1990) This set of programs calculatesthe catch-at-age from equations found in Gavaris and Gavaris(1983) It generated an age-length key for each spawning groupand projected it onto the landings for that cell Thenumbers-at-age thus obtained were then summed to yieldcorresponding annual totals for fixed mobile and all gears

Average weight-at-age was simultaneously computed by AGELEN

7

for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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therefore likely to be discarded in favour of a clear-cutotolith type In 1985 the Gulf Region started usingDiscriminant Analysis of otolith morphometrics which was meantto replace the agers evaluation of otolith type when maturitystage was ambiguous It was felt that using a Discriminant scorewould be an objective way of evaluating otolith morphology notsubject to within- and among- ager variability The region wasasked to document and justify this method Messieh et al (1989)have done so but provide little information on the selection ofthe classification data sets which we know was inconsistent fromyear to year and often included the agers opinion of otolithtype (thus generating tautological classifications) The methodwas to be re-examined and documented here as well but this couldnot be done in the time available and had to be postponedInstead spawning group was reassigned for samples collectedbetween 1985 and 1988 by a method more consistent with previousmethodology namely gonad ripeness in season or alternatelyotolith morphology as evaluated by the ager (see Methods below)

This document outlines the revisions to the previous (ineffect up to 1989) catch- and weight-at-age matrices Unexpectederrors in the data files and in the landings were discovered andwere edited before reconstructing the matrices The effect ofeach revision step on the matrices is being documented with therevised 1989 assessment (Claytor and Dupuis 1990)

Methods

a) Catch data

The landings for particular gear types fishing seasons andunit areas (or cells) that had been used to generate the catch-at-age for the 4T herring assessments up to 1989 rarely added upto the total landings for that year The compilation method wasnot documented in the assessment documents Landing statisticswere therefore recompiled Updated landing statistics for 4Tfrom 1974 to 1982 were drawn from McMillan et al (1984)Quebecs 1982 statistics were not included in this report andwere therefore obtained from Table 5 in the NAFO StatisticalBulletin (NAFO 1984) Starting with 1983 Maritime landingswere compiled from the Gulf Regions purchase slip files towhich Quebec totals were added The latter were obtained eitherfrom Statistical Bullletins (1983-1985) NAFO data tape (1986) orZIFF file (1987) Landings for 1987 and 1988 are preliminary

Landings (round weight in kg) for each gear month and unitarea were pooled into cells defined by gear type (fixed and

4

mobile) fishing season (spring and fall) and aggregatedstatistical unit areas (431-435 436 437-439) for each yearThey were later rounded off to the nearest metric tonne (MT)When the statistical unit area of a landing was unknown (4Tu)as sometimes occurred in the Maritimes or Quebec Regionsreports it was reallocated proportionally to specific 4T pooledareas fished with the same gear and in the same season In 1975and 1977 there were no landings reported for fixed gear duringany month in any specific area but such landings listed under4Tu were considerable They were reallocated to specificpooled areas by calculating for 1974 1976 and 1978 the averagerelative catches in the three areas and applying theseproportions to the catch recorded for 4Tu The averages werebased on only those years because the relative importance of thetwo fishing seasons and gear types shifted in subsequent years

b) Commercial sampling

Data files consisting of random length-frequency samplesfrom which random detailed samples were drawn were copied fromthe CYBER at BSO for 1974 to 1982 except for the years 1978-1981 which were obtained from the St Andrews magnetic tapecollection Data files generated before 1974 cannot be usedbecause the errors missing data and unreliable spawning-groupand age assignments cannot be corrected

Attempts to match length-frequency with detailed biologicaldata revealed pervasive errors and missing information foridentifying samples in these data files Length-frequency andbiological samples were therefore renumbered and obviouskeypunching and coding errors were corrected by checkingphotocopies of data sheets filled out by samplers and laboratorytechnicians For 1980 and 1981 several data sheets had not beenkeypunched or had been deleted and therefore were restoredOnce data files had been reorganized length-weight regressionswere executed to flag outliers Such data points were examinedin concert with gonad maturity and weight and were eitherretained (if the length-weight relationship seemed plausible)edited (if a keypunching error was found) or deleted (if theerror was untraceable)

There have been several changes in the procedures used todetermine age and spawning-group affinity over the years whichreduce consistency across years Thus the reconstruction of thecatch-at-age matrices was based on the method most consistentlyapplicable to all years namely maturity stage if unambiguous(stages 56 and 7 with July 1 delineating the spring and fallspawning seasons) or alternately otolith type as evaluated bythe ager (Cleary et al 1982) This technique was applied to

9

the 1985-1988 samples to regenerate age and spawning-group dataIn 1985 however maturity stages were determined in part byunsupervised temporary helpers with the result that maturitystages seemed unreliable and were deleted from the current filealtogether For 1985 therefore all fish were assigned to aspawning group by the agers evaluation of otolith type whereasnormally 50-80 the fish sampled in a given year get classifiedaccording to maturity stage (weighted average for the 1980sexcluding 1985 is 63)

After editing the data files the length-frequency sampleswere matched to their corresponding biological data to create amaster data file for each spawning group of numbers aged atlength in each sample Samples were then pooled by area fishingseason and gear type (or cells) The corrections outlinedabove usually increased the effective sample sizes butunfortunately did not rectify the sampling deficiency for certaincells (Table 1)

The detailed samples were used to construct age-lengthrelationships specific to each cell (some of the fish that wereused for regressing weight on length could not be aged and weretherefore subsequently excluded from the age-length key) Whencell samples were missing or too small to be reliable (because ofa low r for the length-weight regression or very few lengthgroups represented in the sample) other samples had to besubstituted to construct the age-length key The substitutionsto be applied were determined by the considerations listed below

1 - The unknown spawning-group composition (percentage bynumber) of the landings from one cell was estimated by taking theaverage (weighted by sample size) of the percentages in theequivalent cell for the two most recent and the two subsequentyears available (Table 1) As an example the South SpringMobile catch in 1976 would be estimated by averaging thepercentages observed in the equivalent cell in 1974 1975 1977and 1978 Table 1 indicates however that there was no samplingin 1974 or 1977 limiting the number of most recent cells toone (1975) and shifting the two subsequent available years to1978 and 1979 The percentage estimate is required to adjust thenumbers-at-age calculated with the substitute sample as thelatter is rarely composed of the same proportions of spring andfall spawners as those estimated for the missing sample Weassume that population composition for a cell is similar acrossyears This seems more likely than relying on a population froma different area season or gear For unit area 436 and unitareas 437-439 however the Spring Mobile-gear fishery was neversampled preventing any estimation of spawning-group compositionbased on adjacent years (Table 1) In this instance thepercentage of spring spawners in the catch had to be estimated

C

from the Southern samples collected sporadically over the years

2 - The age-length key for an empty cell is preferablyderived from the samples from another cell in that year ratherthan from the equivalent cell in another year to avoid thepossibility of interdependent age or size shifts between yearsThe possible choices in selecting a substitute cell were

a) the same season and gear in a different areab) the same season and area but different gearc) the same season but a different area and geard) for mobile gear only the same area and gear but a

different season ore) the same season gear and area in a different year

Because fixed and mobile gears usually targetted fish ofdifferent maturity stages within the same area the order ofpreference for these choices was usually as listed above but wasdetermined in each case by comparing the size distributions andthe regression coefficients of length-weight relationships in therelevant cells from better-sampled years The final choice of asubstitute sample was therefore particular to that case and wasaffected by the similarities noted in other years

3 - Partial substitutions were also required when a samplewas available (and could therefore reveal spawning-groupcomposition) but consisted almost exclusively of one group Insuch cases the sample size of the rare spawning-group was ofteninadequate for its age-length determination The decision stepsoutlined in 2) above were followed and another subsample wassubstituted

c) Catch-at-age matrices

Length-weight regressions were computed with the SAS packagefor both spring and fall spawners in each cell The percentagesof spring and fall spawners at each length were also calculatedThe options defining a particular cell (area fishing season andgear type) selected the appropriate samples in the master filewhich along with the corresponding regression parameters totallandings and percentages-at-length were input into the FORTRANprogram AGELEN (Wright 1990) This set of programs calculatesthe catch-at-age from equations found in Gavaris and Gavaris(1983) It generated an age-length key for each spawning groupand projected it onto the landings for that cell Thenumbers-at-age thus obtained were then summed to yieldcorresponding annual totals for fixed mobile and all gears

Average weight-at-age was simultaneously computed by AGELEN

7

for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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mobile) fishing season (spring and fall) and aggregatedstatistical unit areas (431-435 436 437-439) for each yearThey were later rounded off to the nearest metric tonne (MT)When the statistical unit area of a landing was unknown (4Tu)as sometimes occurred in the Maritimes or Quebec Regionsreports it was reallocated proportionally to specific 4T pooledareas fished with the same gear and in the same season In 1975and 1977 there were no landings reported for fixed gear duringany month in any specific area but such landings listed under4Tu were considerable They were reallocated to specificpooled areas by calculating for 1974 1976 and 1978 the averagerelative catches in the three areas and applying theseproportions to the catch recorded for 4Tu The averages werebased on only those years because the relative importance of thetwo fishing seasons and gear types shifted in subsequent years

b) Commercial sampling

Data files consisting of random length-frequency samplesfrom which random detailed samples were drawn were copied fromthe CYBER at BSO for 1974 to 1982 except for the years 1978-1981 which were obtained from the St Andrews magnetic tapecollection Data files generated before 1974 cannot be usedbecause the errors missing data and unreliable spawning-groupand age assignments cannot be corrected

Attempts to match length-frequency with detailed biologicaldata revealed pervasive errors and missing information foridentifying samples in these data files Length-frequency andbiological samples were therefore renumbered and obviouskeypunching and coding errors were corrected by checkingphotocopies of data sheets filled out by samplers and laboratorytechnicians For 1980 and 1981 several data sheets had not beenkeypunched or had been deleted and therefore were restoredOnce data files had been reorganized length-weight regressionswere executed to flag outliers Such data points were examinedin concert with gonad maturity and weight and were eitherretained (if the length-weight relationship seemed plausible)edited (if a keypunching error was found) or deleted (if theerror was untraceable)

There have been several changes in the procedures used todetermine age and spawning-group affinity over the years whichreduce consistency across years Thus the reconstruction of thecatch-at-age matrices was based on the method most consistentlyapplicable to all years namely maturity stage if unambiguous(stages 56 and 7 with July 1 delineating the spring and fallspawning seasons) or alternately otolith type as evaluated bythe ager (Cleary et al 1982) This technique was applied to

9

the 1985-1988 samples to regenerate age and spawning-group dataIn 1985 however maturity stages were determined in part byunsupervised temporary helpers with the result that maturitystages seemed unreliable and were deleted from the current filealtogether For 1985 therefore all fish were assigned to aspawning group by the agers evaluation of otolith type whereasnormally 50-80 the fish sampled in a given year get classifiedaccording to maturity stage (weighted average for the 1980sexcluding 1985 is 63)

After editing the data files the length-frequency sampleswere matched to their corresponding biological data to create amaster data file for each spawning group of numbers aged atlength in each sample Samples were then pooled by area fishingseason and gear type (or cells) The corrections outlinedabove usually increased the effective sample sizes butunfortunately did not rectify the sampling deficiency for certaincells (Table 1)

The detailed samples were used to construct age-lengthrelationships specific to each cell (some of the fish that wereused for regressing weight on length could not be aged and weretherefore subsequently excluded from the age-length key) Whencell samples were missing or too small to be reliable (because ofa low r for the length-weight regression or very few lengthgroups represented in the sample) other samples had to besubstituted to construct the age-length key The substitutionsto be applied were determined by the considerations listed below

1 - The unknown spawning-group composition (percentage bynumber) of the landings from one cell was estimated by taking theaverage (weighted by sample size) of the percentages in theequivalent cell for the two most recent and the two subsequentyears available (Table 1) As an example the South SpringMobile catch in 1976 would be estimated by averaging thepercentages observed in the equivalent cell in 1974 1975 1977and 1978 Table 1 indicates however that there was no samplingin 1974 or 1977 limiting the number of most recent cells toone (1975) and shifting the two subsequent available years to1978 and 1979 The percentage estimate is required to adjust thenumbers-at-age calculated with the substitute sample as thelatter is rarely composed of the same proportions of spring andfall spawners as those estimated for the missing sample Weassume that population composition for a cell is similar acrossyears This seems more likely than relying on a population froma different area season or gear For unit area 436 and unitareas 437-439 however the Spring Mobile-gear fishery was neversampled preventing any estimation of spawning-group compositionbased on adjacent years (Table 1) In this instance thepercentage of spring spawners in the catch had to be estimated

C

from the Southern samples collected sporadically over the years

2 - The age-length key for an empty cell is preferablyderived from the samples from another cell in that year ratherthan from the equivalent cell in another year to avoid thepossibility of interdependent age or size shifts between yearsThe possible choices in selecting a substitute cell were

a) the same season and gear in a different areab) the same season and area but different gearc) the same season but a different area and geard) for mobile gear only the same area and gear but a

different season ore) the same season gear and area in a different year

Because fixed and mobile gears usually targetted fish ofdifferent maturity stages within the same area the order ofpreference for these choices was usually as listed above but wasdetermined in each case by comparing the size distributions andthe regression coefficients of length-weight relationships in therelevant cells from better-sampled years The final choice of asubstitute sample was therefore particular to that case and wasaffected by the similarities noted in other years

3 - Partial substitutions were also required when a samplewas available (and could therefore reveal spawning-groupcomposition) but consisted almost exclusively of one group Insuch cases the sample size of the rare spawning-group was ofteninadequate for its age-length determination The decision stepsoutlined in 2) above were followed and another subsample wassubstituted

c) Catch-at-age matrices

Length-weight regressions were computed with the SAS packagefor both spring and fall spawners in each cell The percentagesof spring and fall spawners at each length were also calculatedThe options defining a particular cell (area fishing season andgear type) selected the appropriate samples in the master filewhich along with the corresponding regression parameters totallandings and percentages-at-length were input into the FORTRANprogram AGELEN (Wright 1990) This set of programs calculatesthe catch-at-age from equations found in Gavaris and Gavaris(1983) It generated an age-length key for each spawning groupand projected it onto the landings for that cell Thenumbers-at-age thus obtained were then summed to yieldcorresponding annual totals for fixed mobile and all gears

Average weight-at-age was simultaneously computed by AGELEN

7

for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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the 1985-1988 samples to regenerate age and spawning-group dataIn 1985 however maturity stages were determined in part byunsupervised temporary helpers with the result that maturitystages seemed unreliable and were deleted from the current filealtogether For 1985 therefore all fish were assigned to aspawning group by the agers evaluation of otolith type whereasnormally 50-80 the fish sampled in a given year get classifiedaccording to maturity stage (weighted average for the 1980sexcluding 1985 is 63)

After editing the data files the length-frequency sampleswere matched to their corresponding biological data to create amaster data file for each spawning group of numbers aged atlength in each sample Samples were then pooled by area fishingseason and gear type (or cells) The corrections outlinedabove usually increased the effective sample sizes butunfortunately did not rectify the sampling deficiency for certaincells (Table 1)

The detailed samples were used to construct age-lengthrelationships specific to each cell (some of the fish that wereused for regressing weight on length could not be aged and weretherefore subsequently excluded from the age-length key) Whencell samples were missing or too small to be reliable (because ofa low r for the length-weight regression or very few lengthgroups represented in the sample) other samples had to besubstituted to construct the age-length key The substitutionsto be applied were determined by the considerations listed below

1 - The unknown spawning-group composition (percentage bynumber) of the landings from one cell was estimated by taking theaverage (weighted by sample size) of the percentages in theequivalent cell for the two most recent and the two subsequentyears available (Table 1) As an example the South SpringMobile catch in 1976 would be estimated by averaging thepercentages observed in the equivalent cell in 1974 1975 1977and 1978 Table 1 indicates however that there was no samplingin 1974 or 1977 limiting the number of most recent cells toone (1975) and shifting the two subsequent available years to1978 and 1979 The percentage estimate is required to adjust thenumbers-at-age calculated with the substitute sample as thelatter is rarely composed of the same proportions of spring andfall spawners as those estimated for the missing sample Weassume that population composition for a cell is similar acrossyears This seems more likely than relying on a population froma different area season or gear For unit area 436 and unitareas 437-439 however the Spring Mobile-gear fishery was neversampled preventing any estimation of spawning-group compositionbased on adjacent years (Table 1) In this instance thepercentage of spring spawners in the catch had to be estimated

C

from the Southern samples collected sporadically over the years

2 - The age-length key for an empty cell is preferablyderived from the samples from another cell in that year ratherthan from the equivalent cell in another year to avoid thepossibility of interdependent age or size shifts between yearsThe possible choices in selecting a substitute cell were

a) the same season and gear in a different areab) the same season and area but different gearc) the same season but a different area and geard) for mobile gear only the same area and gear but a

different season ore) the same season gear and area in a different year

Because fixed and mobile gears usually targetted fish ofdifferent maturity stages within the same area the order ofpreference for these choices was usually as listed above but wasdetermined in each case by comparing the size distributions andthe regression coefficients of length-weight relationships in therelevant cells from better-sampled years The final choice of asubstitute sample was therefore particular to that case and wasaffected by the similarities noted in other years

3 - Partial substitutions were also required when a samplewas available (and could therefore reveal spawning-groupcomposition) but consisted almost exclusively of one group Insuch cases the sample size of the rare spawning-group was ofteninadequate for its age-length determination The decision stepsoutlined in 2) above were followed and another subsample wassubstituted

c) Catch-at-age matrices

Length-weight regressions were computed with the SAS packagefor both spring and fall spawners in each cell The percentagesof spring and fall spawners at each length were also calculatedThe options defining a particular cell (area fishing season andgear type) selected the appropriate samples in the master filewhich along with the corresponding regression parameters totallandings and percentages-at-length were input into the FORTRANprogram AGELEN (Wright 1990) This set of programs calculatesthe catch-at-age from equations found in Gavaris and Gavaris(1983) It generated an age-length key for each spawning groupand projected it onto the landings for that cell Thenumbers-at-age thus obtained were then summed to yieldcorresponding annual totals for fixed mobile and all gears

Average weight-at-age was simultaneously computed by AGELEN

7

for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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from the Southern samples collected sporadically over the years

2 - The age-length key for an empty cell is preferablyderived from the samples from another cell in that year ratherthan from the equivalent cell in another year to avoid thepossibility of interdependent age or size shifts between yearsThe possible choices in selecting a substitute cell were

a) the same season and gear in a different areab) the same season and area but different gearc) the same season but a different area and geard) for mobile gear only the same area and gear but a

different season ore) the same season gear and area in a different year

Because fixed and mobile gears usually targetted fish ofdifferent maturity stages within the same area the order ofpreference for these choices was usually as listed above but wasdetermined in each case by comparing the size distributions andthe regression coefficients of length-weight relationships in therelevant cells from better-sampled years The final choice of asubstitute sample was therefore particular to that case and wasaffected by the similarities noted in other years

3 - Partial substitutions were also required when a samplewas available (and could therefore reveal spawning-groupcomposition) but consisted almost exclusively of one group Insuch cases the sample size of the rare spawning-group was ofteninadequate for its age-length determination The decision stepsoutlined in 2) above were followed and another subsample wassubstituted

c) Catch-at-age matrices

Length-weight regressions were computed with the SAS packagefor both spring and fall spawners in each cell The percentagesof spring and fall spawners at each length were also calculatedThe options defining a particular cell (area fishing season andgear type) selected the appropriate samples in the master filewhich along with the corresponding regression parameters totallandings and percentages-at-length were input into the FORTRANprogram AGELEN (Wright 1990) This set of programs calculatesthe catch-at-age from equations found in Gavaris and Gavaris(1983) It generated an age-length key for each spawning groupand projected it onto the landings for that cell Thenumbers-at-age thus obtained were then summed to yieldcorresponding annual totals for fixed mobile and all gears

Average weight-at-age was simultaneously computed by AGELEN

7

for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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for each spawning group in each cell Annual weight estimatesfor fixed mobile and all gears combined were obtained byaveraging the weights-at-age (weighted by their correspondingcatch) from all appropriate cells

Results and Discussion

Updated landings for NAFO Division 4T are grouped by areafishing season and gear type in Table 1 along with the revisedspawning-group composition and size of the biological samplesavailable for each cell In most cases the landings haveincreased from those previously used but some values most oftenin the north show a decline The most probable explanation forsuch reductions is the 4Tu landings which may have beenallocated to specific areas differently in the past (theprocedures were not documented)

In general the corrections made to the data files havealtered sample sizes and spawning-group composition by only asmall percentage when compared with Table 2 in CAFSAC Res Doc8963 (Chadwick et al 1989) Sample sizes increasedsubstantially in 1980 and 1981 due to newly keypunched samplesOther notable changes were found in 1985 and 1986 a few samplesoriginally allocated to 4T1 or to 4Tq in fact belonged to 4Tj or4Tg respectively thereby increasing the souths sample size by200 The percentage of spring spawners in this area alsochanged from 100 to 80 This drop and the 10-15 differencesobserved in most cells of 1986 are the result of the spawning-group reassignment It was discovered while editing thebiological files that the 1987 and 1988 results of theDiscriminant Analyses were inadvertently omitted from theassignment process such that the reassignment incorporated intothe revised matrices affected the spawning-group composition of1985 and 1986 samples only

The catch-at-age matrices (Tables 2-4) begin with 1974although it is apparent from Table 1 that many cells were notsampled that year and therefore several substitutions for age-length keys were required Mean weights-at-age are found inTable 5-7 The unusual changes in weight observed as one followssome cohorts across the years are attributable either to a lowsample size for that cohort in a given year or to the cohortspoor fit with the length-weight relationships observed in a givenyear The new catch-at-age for spring and fall-spawning herringdiffers somewhat from that found in previous 4T herringassessments (Table 8) and the largest deviations tend to occurat extreme ages and represent minor catches Despite the

8

reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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reassignment of spawning group using a more traditional methodinstead of Discriminant Analysis on otolith morphometrics thenumbers-at-age for 1985 and 1986 have not been altered to anygreater extent than those of other years

The impact of edited data files updated landings andspawning-group reassignment upon the revised catch-at-agematrices are being documented individually in conjunction withthe revised 1989 assessment (Claytor and Dupuis 1990) Inaddition otolith characteristics the application ofDiscriminant Analysis to otolith morphometrics and the GSI model(McQuinn 1989) as methods of assigning spawning group are beingcompared by the Gulf Region The method deemed most useful willbe subject to CAFSAC approval before being applied retroactivelyto historical data sets

Acknowledgements

We thank Gloria Nielsen for compiling and updating landingsfrom the Gulf Regions ZIFF and Purchase-slip data files andJohn Wright for repeatedly editing the AGELEN program to suit ourspecific needs Ross Claytor and David Cairns made valuablecomments on an earlier draught

Literature Cited

Ahrens M 1985a Annual assessment of herring in NAFO Division4T CAFSAC Res Doc 8545

Ahrens M 1985b Two phase sampling of the herring fishery inNAFO division 4T CAFSAC Res Doc 8546

Anonymous 1986 Advice on the management of some herring stocksin the Gulf of St Lawrence CAFSAC Adv Doc 8619

Chadwick EMP DK Cairns and HMC Dupuis 1989Assessment of Atlantic herring in NAFO division 4T 1988CAFSAC Res Doc 8963

Claytor RR and HMC Dupuis 1990 Effects of revising 4Therring catch-at-age matrices CAFSAC Res Doc (inprep)

9

Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Cleary L JJ Hunt J Moores and D Tremblay 1982 Herringaging workshop St Johns Newfoundland March - 1982CAFSAC Res Doc 8241

Gavaris S and CA Gavaris 1983 Estimation of catch at ageand its variance for groundfish stocks in the Newfoundlandregion Can Spec Publ Fish Aquat Sci 66 178-182

McMillan JI RN OBoyle L Cleary and SN Messieh 1984Compilation of landing statistics of the southern Gulf ofSt Lawrence herring ( Clupea harengus harengus ) for 1967-1982 Can Data Rep Fish Aquat Sci 468 231 pages

McQuinn IH 1989 Identification of spring- and autumn-spawning herring ( Clupea harengus harengus ) using maturitystages assigned from a gonadosomatic index model Can JFish Aquat Sci 46 969-980

Messieh SN C MacDougall and R Claytor 1989 Separation ofAtlantic herring ( Clupea harengus harengus ) stocks in thesouthern Gulf of St Lawrence using digitized otolithmorphometrics and discriminant function analysis Can TechRep Fish Aquat Sci 1647 22 pages

NAFO 1984 (-1987) Fishery statistics for 1982 (-1985) StatBull 32 (-35)

Wright J 1990 AGELEN-- A system of programs for computingestimates of age and length distributions in fishpopulations Can Tech Rep Fish Aquat Sci (in prep)

10

Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 1 Landings in metric tonnes (MT) for NAFO Division 4T by area fishing season and gear type 1987 -

1988 landings are provisional Spring fishing January-June Fall fishing July-December Most fixed gears

are gillnets mobile gears are mainly purse seines XP Percentage (by numbers) of spring spawners (P) in

biological samples N sample size NS no sample available () or inadequate for one of the spawning group

(A or P)

SOUTH (4Tf-4Tk) MIDDLE (4Tl) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

1974 MT 3572 8762 497 17 2767 5 792 0 1954 31 2914 13974

P 96 12 11 39 99 12 0 -- 100 12 0 39

N 1096 658 234 -- 344 439 1994

NS A

1975 MT 1939 18565 573 0 3539 0 619 0 1539 0 2605 14149

P 100 16 0 -- 98 -- 3 -- 100 -- 0 83

N 1592 660 100 -- 2691 -- -- -- 808 780

NS

1976 MT 1766 16871 527 173 2940 0 604 0 1875 345 2286 12034

P 100 20 22 39 87 -- 3 -- 98 20 0 56

N 2276 98 717 -- -- 156 598 1018

NS A

1977 MT 1361 19887 492 0 2484 0 531 106 1080 0 2237 16645

P 100 20 2 -- 100 -- 3 48 99 -- 6 70

N 1189 -- 671 -- 239 -- 298 2040

NS P

1978 MT 1114 4648 933 4885 6261 0 630 3491 1172 3399 3578 23379

P 97 9 0 39 81 -- 3 31 100 20 2 52

N 4275 1090 121 516 489 -- 292 89 945 1487

NS

1979 MT 1241 13901 2305 78 5246 0 1069 5738 1137 0 2701 14803

P 90 31 1 39 85 -- 5 18 88 -- 12 66

N 6082 1246 1311 499 -- 192 345 -- 287 3520

NS P

1980 MT 1994 13897 2786 320 3604 20 1826 793 1674 0 1933 13699

P 99 24 1 39 100 22 5 19 98 -- 7 66

N 4780 1488 1100 186 97 300 -- 297 2346

NS P A

11

62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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62 6636 2569

27 2 12

1867 798

0 7091 148

-- 1 49

-- 1151 1200

P

0 6697 2164

- 6 63

925 992

Table 1 (contd)

SOUTH (4Tf-4Tk) MIDDLE (4TL) NORTH (4Tm-4To)

SPRING FALL SPRING FALL SPRING FALL

FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE FIXED MOBILE

0 5572 93

-- 3 16

-- 2291 759

1981 MT 2386

P 97

N 3157

NS

1982 MT 2015

YP 98

N 4069

NS

1983 MT 1911

P 97

N

NS

1984 MT 663

P 76

N 157

NS

1985 MT 2352

80

N 240

NS

1986 MT 3336

P 93

N 192

NS

1987 MT 3600

P 94

N 258

NS

1988 MT 2050

P 93

N 572

NS

21 3272 3081 4028

27 1 39 84

400 494 p

0 5241 0 2836

-- 0 -- 100

-- 298 -- 397

0 5177 85 5097

-- 0 22 93

-- 743

683

3 7939 0 4192

27 0 -- 96

446 - -

0 9362 0 3902

-- 8 -- 99

-- 449 -- 70

A

0 12265 44 3389

-- 0 22 100

-- 663 36 204

0 18198 78 3738

-- 0 22 100

-- 1692 232

0 15034 226 3967

-- 1 22 94

-- 685 425

P

0 2381 14 1087

- 1 18 83

- 694 1053

P

0 1105 9 1072

- 0 18 99

- 75 772 A

19 1572 3256 1515

27 5 18 96

175 114 p A

1 1338 114 427

27 1 18 85

406 143

P

0 1413 0 963

-- 2 -- 65

77

0 1570 0 1825

-- 2 -- 85

-- 281

0 1970 13 4386

-- 0 18 93

-- 32 325

1 3785 1 6616

27 0 18 70

95 527

0 12932 7022

-- 29 43

-- 336 429

0 26056 10871

-- 13 62

-- 578 693

0 31862 13613

-- 6 49

-- 1309 445

0 22266 17423

-- 14 48

-- 763 530

12

Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 2 Annual catch-at-age (1000s) of spring and fall spawners in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 107 0 35 1131 1434 6787 701 85 7 0 0 190 47 0 2467

2 2616 1058 16335 3541 16542 15870 12366 1674 772 305 237 1587 1639 306 5697

3 4932 20105 6679 58151 13469 14435 24371 12588 24424 16343 5366 8782 5379 2360 5797

4 I 4081 10670 36486 8676 52354 10846 11445 8131 4339 26786 16610 19159 24684 9357 108835 I 1978 2053 5117 8359 3736 37186 6957 2044 1063 2017 9884 16581 16783 29599 111326 22693 11944 1528 888 5814 5795 19340 1921 465 492 702 5804 17456 16648 19027

7 I 7646 21774 204 2569 1401 4393 9782 3258 350 64 86 1693 3561 14614 124588 4209 3082 7439 315 1123 1817 4482 1160 348 23 50 779 818 5572 7878

9 4968 3314 860 3584 181 772 2649 699 126 5 17 260 245 750 2544

10 J 605 4802 480 3873 4125 772 1185 512 91 0 5 4 131 509 19611+1 5363 5837 3736 3459 2070 3125 1142 763 587 0 0 155 197 202 1878

ALL GEARS - FALL SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 101 0 0 0 0 1115 148 0 0 0 0 0 0 0 66

2 2421 23 71 195 1512 2747 2364 358 203 210 36 264 557 62 3984

3 4478 4149 164 6995 23044 7010 39878 10107 7606 6850 1127 4353 2510 9090 3714

4 26000 5110 1976 10091 29774 38935 18275 30067 18424 26606 26157 16904 35492 39403 23612

5 I 9632 30464 5872 19178 15160 26002 23878 7476 21118 11157 14436 30191 20940 33195 435386 J 5032 8896 17483 8318 4672 12342 7705 3255 5488 14225 8892 13562 40368 25761 248467 I 18672 4122 4491 14338 5452 3392 4648 1585 2703 2152 6526 8188 21094 41792 229508 974 7228 3022 6408 19244 3374 1482 882 1118 1753 1588 3594 10967 18992 20477

9 4621 3049 5327 2484 1847 6294 1726 236 492 548 541 823 4185 10713 11800

10 I 3082 4490 1181 4884 2377 1282 1202 241 225 149 211 894 562 4526 433811 +1 8962 19984 33949 19963 16893 9695 358 309 181 194 75 295 870 1778 3054

13

Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 3 Annual catch-at-age (1000s) for spring and fall spawners in fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

ii 1 0 0 17 0 105 0 85 7 0 0 0 0 0 57

2 107 51 0 40 4 17 205 947 52 73 11 360 50 150 487

3 3454 14335 1410 18158 5544 5974 11194 11285 23679 14226 3549 6085 3026 1325 3972

4 I3160 3189 16902 2901 25449 2292 7078 7363 4246 25857 14001 16610 17926 7697 89425 1352 926 2610 4121 1142 17595 2693 2041 1057 1892 8501 15127 11485 22395 9895

6 15869 1737 528 489 1631 552 9204 1385 461 474 462 5145 11994 10559 13598

7 2674 6581 114 90 227 1214 1869 2929 331 54 44 1418 1675 9722 7842

8 2952 1360 3737 117 257 180 892 480 331 5 31 643 383 3504 5807

9 2951 1043 505 1082 88 131 764 459 125 1 2 75 24 681 1385

10 332 1497 300 22 1379 303 456 187 91 0 5 4 117 441 196

11 +1 541 1211 2632 803 764 1266 683 761 583 0 0 9 137 169 598

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977

1 0 0 0 0

2J 0 0 0 0

31 157 0 8 434

4 5577 1437 344 3164

5 1764 6601 1845 480

6 534 968 5052 384

7 2125 634 625 4013

8 145 605 221 222

9 I 1635 435 573 92101 383 475 72 664

11 +1 2582 1671 2500 1000

1978 1979

0 903

80 6

3520 532

5608 10754

3593 5789

904 2672

958 907

5103 1068

117 958

521 265

2589 710

1980 1981 1982 1983 1984 1985 1986 1987 1988

0 0 0 0 0 0 0 0 0

91 281 0 0 0 31 403 15 0

8149 5406 2492 993 920 1966 1606 8166 1401

5460 25698 15876 22006 25228 14054 33820 38450 21118

6285 7187 18618 10225 13303 25547 17678 30658 40919

1023 3142 5152 12739 8235 10342 35098 20148 21742

1244 1457 2586 1927 6162 6622 18454 36671 14437

269 863 1048 1690 1529 2829 9483 14711 14841

293 152 389 524 517 541 3402 9517 7868

302 240 114 108 211 481 362 4340 2832

52 309 58 162 74 27 745 1764 1475

14

0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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0

77

4702

4369

289

112

129

19

84

1

0

0

203

5114

2548

2500

336

116

70

103

111

123

0

233

2387

2850

4644

3220

1566

765

282

413

268

0

154

903

1672

3262

5271

2640

1485

783

200

125

Table 4 Annual catch-at-age (1000s) for spring and fall spawners in mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGEI

1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

106 0 35 1114 1434 6681 701 0

2508 1007 16335 3501 16538 15853 12161 727

1478 5771 5269 39993 7925 8461 13177 1303

921 7481 19584 5775 26905 8555 4366 768627 1127 2507 4238 2594 19591 4265 3

6824 10207 1000 400 4184 5243 10136 536

4971 15193 90 2479 1174 3179 7913 329

1257 1722 3702 199 866 1637 3590 679

2017 2271 355 2501 93 641 1885 239

273 3305 179 3851 2746 469 729 326

4822 4625 1104 2656 1306 1859 459 2

0 0 0 190 47 0 2410

720 232 226 1227 1589 155 5210

745 2117 1818 2697 2353 1035 1825

93 928 2609 2549 6758 1659 1941

6 125 1383 1454 5298 7204 1237

3 18 240 659 5463 6089 5429

19 10 42 275 1886 4892 4616

17 17 19 136 435 2068 2070

1 4 14 185 220 69 1159

1 0 0 0 14 68 0

5 0 0 146 59 32 1280

1i21

3

41

5

61

71

8

91

10

11 +1

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 101 0 0 0 0 212 148

21

2421 23 71 195 1432 2741 2273

31

4320 4149 155 6561 19524 6478 31729

41 20423 3673 1632 6927 24166 28182 12815

51

7868 23863 4027 18698 11567 20212 17593

61

4498 7928 12431 7934 3767 9670 6682

71 16547 3488 3866 10325 4494 2485 3404

81

829 6623 2801 6185 14142 2306 1213

91

2985 2614 4754 2392 1730 5336 1433

10

2699 4015 1110 4220 1856 1017 900

11 +1

6380 18314 31449 18963 14305 8986 305

0 0

210 36

5857 207

4600 929

933 1133

1486 657

225 364

62 60

23 25

42 0

32 1

0 66

47 3984

924 2313

954 2494

2537 2619

5613 3104

5121 8514

4281 5635

1197 3932

186 1506

13 1580

15

Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 5 Mean weight-at-age (kg) of spring and fall spawners caught in 4T all gears combined

ALL GEARS - SPRING SPAWNERS

AGEJ 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0064 0101 00779 00953 01361 01128 00316 0 0 00872 0089 0 00834

2 0125 0086 01405 01401 01307 01654 01629 01573 0142 01376 01481 01702 01404 01408 0116

3 I 01606 01526 01626 01832 01704 01483 01658 01834 01747 01556 01809 01886 01796 01982 01716

4 02084 01742 01888 01965 02268 01665 01852 02338 02077 02099 01991 02231 02137 01990 02168

5 02299 02091 02108 02347 0245 0243 02342 02636 02624 02386 02179 02492 02498 02348 02401

6 02537 02026 02301 02638 02789 02349 02662 03288 03175 02738 03011 02803 02771 02748 02837

7 02818 02547 02674 02847 03103 02859 02555 0326 03816 03326 0373 03379 03156 02928 03058

8 02864 02937 02963 02786 03439 03018 02728 03016 03801 02528 04025 03767 03466 03052 03209

9 02963 03212 03169 0311 03311 03231 03095 03411 04028 0426 03401 04793 03152 03158 03609

10 I 03077 03176 03185 03238 03563 0328 03546 03279 04058 0 03818 0386 03143 03287 0395911+ 03429 03623 03189 03468 03876 03793 0394 03973 04445 0 0 03262 04046 04163 03922

ALL GEARS - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00498 0 0 0 0 00311 00318 0 0 0 0 0 0 0 00726

2 00665 00566 00887 01178 00972 01128 01023 01284 00949 01073 01047 01176 01558 01408 00934

3 I 01352 01103 01802 0138 01458 01218 01596 01931 01819 01742 02208 02242 01845 02313 01981

4 01968 02 01866 0214 02208 01984 02036 02425 02486 02439 02455 02647 02476 02469 02608

5 02333 02388 02219 02249 02634 02368 02676 03065 02729 02816 02824 02928 02867 02810 02909

6 02637 02662 02722 02583 02793 02697 02921 03601 03175 03126 03182 03224 03206 03145 03244

7 I 02629 02994 02875 02974 02949 03095 0309 03873 03659 03453 03443 03548 03588 03414 03404

8 0316 03028 03192 02995 03351 03426 03281 04054 03677 03636 03924 037370375903663 03676

9 I 0314 03269 0328 03203 03173 0355 03455 03809 03594 03701 04079 03972 03892 03793 03925

10 03071 03337 03551 0328 03194 03673 03836 0472 03386 03929 03918 03938 04027 03889 04008

11+1 0352 03653 03812 03545 0386 03824 04155 04246 04236 04311 04869 04597 04138 04305 043

16

Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 6 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with fixed gear

FIXED GEAR - SPRING SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 00537 0 0 0101 0 00194 0 01128 00316 0 0 0 0 0 00386

2 01129 01146 0 016 00798 01171 02265 01511 01762 01376 01123 02193 01811 00906 00818

3 01573 01536 01594 01562 0149 01673 01573 01812 01735 0151 01684 0184 01575 01752 01663

4 I 02057 01941 01854 01882 01852 01997 01719 0231 02072 02093 01939 02209 02007 01883 02073

5 02254 02292 02177 02217 02038 02319 02182 02636 02623 02367 02128 02472 02389 02159 0236

6 02507 02493 02488 02611 02449 0235 02664 03106 03175 02716 03032 02774 02645 02463 02622

7 I 02735 02773 02587 02629 02912 03047 02933 03304 03768 03358 03937 03384 03001 02703 02849

8 02849 02893 02868 0274 0299 03404 03268 03412 03833 02528 04444 03647 03448 02744 0299

9 I 02937 03022 03087 02969 03271 0345 03653 03725 04027 0426 03903 04283 03285 03028 03347

10 I 03043 03115 02965 02998 03199 0318 03723 03946 04057 0 03819 0386 02981 03060 03959

11+1 03281 03383 03115 03239 03614 03507 03845 03973 04445 0 0 03414 04169 03991 03177

FIXED GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1 1 0 0 0 0 0 00231 0 0 0 0 0 0 0 0 0

2 0 0 0 0 00808 01003 01922 01325 0 0 0 02332 01745 01405 0

3 01669 0 02134 01705 01378 02027 02072 02087 02094 01863 0234 02629 02007 02345 02454

4 I 02227 02383 02609 02375 02373 02547 02565 0246 0255 02492 02466 02733 02488 02469 026475 I 02696 02772 02898 02791 02825 02945 03113 03087 02748 02844 02852 02988 0292 02805 02921

6 02972 03155 03142 02873 0306 03239 03597 03627 03201 03156 03217 03322 03263 03192 03247

7 I 03089 03341 03429 03222 03153 03583 03575 03906 03658 03488 03466 0364 03665 03439 03512

8 03363 03436 03489 03375 03642 03992 03857 0409 03694 03637 03953 03824 03849 03729 03774

9 I 03501 03732 03775 03598 03734 04143 04082 04482 0374 03683 04104 04005 04001 03814 04013

10 03381 03783 03604 03699 03382 04288 04474 04712 03495 04588 03918 0383 04488 03878 03973

11+1 03753 04007 04074 03927 04195 04201 04677 04246 0388 0427 04884 04741 04157 04306 04504

17

Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 7 Mean weight-at-age (kg) of spring and fall spawners caught in 4T with mobile gear

MOBILE GEAR - SPRING SPAWNERS

AGES 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

1i 00537 0 0064 0101 00779 00965 01361 00281 00316 0 0 00872 0089 0 00845

2J 01255 00846 01405 01399 01307 01654 01618 01652 01395 01376 01499 01558 01391 01893 01192

3j 01683 01499 01634 01955 01853 01348 0173 02031 02135 01862 02053 0199 02081 02276 01831

41 02178 01657 01917 02006 02661 01576 02066 02602 02298 02245 02269 02373 02484 02486 02606

5 02396 01927 02036 02474 02632 0253 02443 02604 0274 02669 02497 027 02735 02934 02732

61 02605 01947 02203 02672 02921 02349 0266 03759 03208 03322 0297 03033 03049 03242 03376

02862 02449 02784 02854 0314 02787 02466 02863 0465 03147 03509 03354 03293 03376 03412

81 02901 02972 03059 02814 03572 02975 02594 02736 03198 02528 03364 04335 03482 03574 03823

91 03 03298 03285 03171 03349 03187 02868 02807 04132 0426 03324 04999 03137 04454 03923

10

03119 03203 03553 0324 03745 03344 03435 02897 04322 0 03515 0 04544 04769 03394

11+1 03445 03686 03365 03537 04029 03987 04081 03966 04456 0 0 03253 0376 05062 0427

MOBILE GEAR - FALL SPAWNERS

AGEI 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

11 00498 0 0 0 0 00652 00318 0 0 0 0 0 0 0 0072621 00665 00566 00887 01178 00981 01129 00987 01133 00949 01073 01047 01024 0107 01409 00934

31 0134 01103 01784 01358 01473 01152 01473 01751 01685 01722 01617 01923 01558 02032 01694

41 01897 0185 01709 02033 0217 0177 0181 02218 02082 02186 02168 0222 02234 02493 02278

5 02252 02281 01908 02235 02575 02202 0252 02516 02584 02507 02484 02598 02575 02869 02725

61 02597 02602 02551 02569 0273 02547 02818 02872 02775 02875 02736 02909 02833 02974 03223

71 0257 02931 02786 02878 02905 02918 02913 03497 03684 03161 03053 03157 03053 03238 03219

81 03125 0299 03169 02981 03246 03163 03153 02383 03407 03618 03163 03417 03188 03436 03418

91 02942 03192 0322 03188 03135 03444 03326 02583 03043 04101 03558 0391 03417 03623 03748

10 03027 03284 03548 03214 03141 03513 03622 06919 03275 02231 0 04064 03192 04143 04072

11+1 03426 0362 03791 03525 03799 03794 04065 0 04404 04518 04143 04583 0403 0417 04109

18

Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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Table 8 Ratio of new annual catch-at-age numbers to old catch-at-age (see CAFSAC Res Doc 8963)

SPRING SPAWNERS - ALL GEARS

AGEI 1978 1979 1980

2 11460 7300 5783

3 9538 10545 5724

4 I 8017 18521 201185 7962 10952 22471

6 8358 27207 12265

7 I 10971 14300 299248 9501 25700 22046

9 9476 38030 35797

10 I 11509 10752 3703111 +1 10392 8959 3924

1981

2726

7082

9856

15675

22131

7331

15364

9246

47407

6369

1982

8355

7316

6997

7202

13798

16129

10265

11053

454998

53364

1983 1984 1985

7193 11449 126960

15103 15437 10365

8584 15055 16910

5127 7143 14163

4457 4652 10812

9143 7414 8315

4600 45455 21401

2941 15455 10442

0000 2273 40000

0000 0000 1549985

1986

52032

10713

14297

8127

10877

6143

4907

4554

11197

4273

1987 1988

11250 130068

10328 12199

11118 7379

10171 7898

9523 9267

9736 11663

9556 9692

10260 12272

11114 4516

10306 11804

FALL SPAWNERS - ALL GEARS

AGEI 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988

2 I 9987 9453 17268 32844 11033 60000 40000 88000 16828 10877 12072723 I 11910 11276 12297 10032 8202 14325 9930 11651 5741 9709 238694 10875 11114 18284 9055 8559 11142 9505 9554 9879 9942 8696

5 I 10758 9411 10258 12521 8077 10170 8885 12542 7973 10004 82906 11759 11110 9235 12490 9691 10427 6855 10742 11523 9975 9522

7 15734 14602 11254 16207 11532 8933 9714 14122 10506 10076 11922

8 13892 10786 23265 9028 11135 9389 11457 16763 10812 09971 11158

9 I 11501 12007 20354 10926 7676 8796 11271 19095 12651 10059 1158510 26708 18262 37562 22315 17045 13070 13701 44039 10505 09725 14407

11+ 10390 9335 1207 3544 11173 6278 4310 2949971 13043 10131 13794

19

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