annotated checklist of vascular...

National Park ServiceU.S. Department of the Interior

Natural Resource Program Center

Annotated Checklist of Vascular FloraPipe Spring National Monument

Natural Resource Technical Report NPS/NCPN/NRTR–2008/131

ON THE COVER

Wetland below West Cabin, Pipe Spring National Monument, Arizona. Photo by Walter Fertig.

Annotated Checklist of Vascular FloraPipe Spring National Monument

Natural Resource Technical Report NPS/NCPN/NRTR–2008/131

AuthorsWalter Fertig Moenave Botanical Consulting1117 W. Grand Canyon Dr.Kanab, UT 84741

Jason AlexanderCurator, UVU HerbariumUtah Valley University800 West University ParkwayOrem, UT 84058

Editing and DesignAlice Wondrak BielNorthern Colorado Plateau NetworkP.O. Box 848Moab, UT 84532

October 2008

U.S. Department of the InteriorNational Park ServiceNatural Resource Program CenterFort Collins, Colorado

ii Annotated Checklist of Vascular Flora: Pipe Spring National Monument

The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientifi c community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer-reviewed to ensure that the information is scientifi cally credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner.

The Natural Resource Technical Report series is used to disseminate the peer-reviewed results of scientifi c studies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service’s mission. The reports provide contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. Current examples of such reports include the results of research that addresses natural resource management issues; natural resource inventory and monitoring activities; resource assessment reports; scientifi c literature reviews; and peer-reviewed proceedings of technical workshops, conferences, or symposia.

Views, statements, fi ndings, conclusions, recommendations, and data in this report are solely those of the author(s) and do not necessarily refl ect views and policies of the U.S. Department of the Interior, National Park Service. Mention of trade names or commercial products does not constitute endorsement or recommendation for use by the National Park Service.

Printed copies of reports in this series may be produced in limited quantity, and are only available as long as the supply lasts. You may send a request to:

Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

This report is also available electronically from the Northern Colorado Plateau Network website, http://science.nature.nps.gov/im/units/NCPN/, or athttp://www.nature.nps.gov/publications/NRPM.

When original printed copies are exhausted, copies can be obtained from: Technical Information Center (TIC) Denver Service Center National Park Service P.O. Box 25287 Denver, CO 80225-0287

Please cite this publication as:Fertig, W., and J. Alexander. 2008. Annotated checklist of vascular fl ora: Pipe Spring National Monument. Natural Resource Technical Report NPS/NCPN/NRTR–2008/131. National Park Service, Fort Collins, Colorado.

NPS D-222, October 2008

Contents iii

Contents

Figures ...........................................................................................................................................vTables ...........................................................................................................................................viiAbstract ........................................................................................................................................ ixAcknowledgements .....................................................................................................................xi

1 Introduction .............................................................................................................................11.1 Background and justifi cation .......................................................................................................11.2 Objectives ......................................................................................................................................11.3 Study area .....................................................................................................................................1

1.3.1 History and setting ........................................................................................................................ 11.3.2 Geology ......................................................................................................................................... 21.3.3 Vegetation ..................................................................................................................................... 21.3.4 Previous botanical studies ............................................................................................................ 4

2 Methods .....................................................................................................................................7

3 Results ........................................................................................................................................93.1 Summary of the fl ora of Pipe Spring National Monument .......................................................93.2 Life form and biogeographic diversity ......................................................................................103.3 Plants of special concern ............................................................................................................103.4 Noxious weeds ............................................................................................................................11

4 Discussion ................................................................................................................................13

5 Literature Cited .......................................................................................................................15

6 Appendices ..............................................................................................................................17Appendix A. Confi rmed, historical, and reported vascular plant taxa of Pipe Spring National Monument, organized by family and scientifi c name with taxonomic and biogeographic notes. .20Appendix B. Potential (unconfi rmed) taxa of Pipe Spring National Monument. .........................44Appendix C. Rejected (falsely reported or questionable) taxa of Pipe Spring National Monument. ........................................................................................................................................47Appendix D. Confi rmed, historical, and reported taxa of Pipe Spring National Monument organized by life form and common name with ecological notes. ...............................................48

iv Annotated Checklist of Vascular Flora: Pipe Spring National Monument

Contents v

FiguresFigure 1.3.1. Location of Pipe Spring National Monument, Mohave County, Arizona. Map created by

Aneth Wight of NCPN (derived from Evenden et al. 2002). ............................................................... 3

Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Pipe Spring National Monument by decade, 1930–2008. ...................................................................................................... 6

vi Annotated Checklist of Vascular Flora: Pipe Spring National Monument

Contents vii

TablesTable 1.3.4. Chronology of plant-collecting efforts in Pipe Spring National Monument. ............................... 5

Table 3.1. Statistical summary of the fl ora of Pipe Spring National Monument. ............................................ 9

Table 3.3. Plant species of conservation concern from Pipe Spring National Monument. ........................... 11

viii Annotated Checklist of Vascular Flora: Pipe Spring National Monument

Contents ix

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Pipe Spring National Monument. This entailed a thorough review of existing literature, re-examination of specimens from the Pipe Spring National Monument herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports or locate new species. Based on this study, 277 vascu-lar plant taxa have been confi rmed or reliably reported as occuring in the monument (215 are known from validated herbarium specimens and 62 from unvouchered literature reports). Another 15 species formerly considered part of the monument fl ora have now been shown to be falsely reported. The current fl ora of Pipe Spring National Monument represents 6.5% of the known fl ora of Arizona and 11% of the fl ora of northern Arizona and the Arizona Strip. Although no federally listed threatened or endangered species are known from the monument, two occur in the vicinity. Three plant species from Pipe Spring National Monument are currently tracked as species of concern by the State of Arizona’s Natural Heritage Program. Introduced plants make up nearly 21% of the monument fl ora (58 species), of which only 4 species are considered noxious weeds by the State of Arizona. Further surveys in Pipe Spring National Monument are likely to fi nd additional new species (46 are known from similar habi-tats outside the monument boundary), especially among late-season fl owering, weedy, or wetland taxa.

x Annotated Checklist of Vascular Flora: Pipe Spring National Monument

Contents xi

Acknowledgements

We would like to thank the following individuals for their assistance with this project. Botanist Dr. Angela Evenden, former leader of the Northern Colorado Plateau Network in Moab, Utah, was an active proponent of this project and helped secure funding for fi eld work and herbarium studies in 2006. John Hiscock and Andrea Bornemeier provided data from monument archives and assisted with research permits. Cheryl Decker and Denise Louie, of Zion National Park, assisted with fi eld surveys of the monument in 2001. Heather Karnes and An Glorieux helped with fi eld work and data entry in 2004 and 2005, respectively. Libby Nance, database specialist with the NCPN, assisted with development and quality control of the monument’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the monument. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hartman and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Thanks to the technical reviewers (Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others whom we may have inadvertently forgotten, please accept our apologies (and thanks).

Research was conducted under permit # PISP-2006-SCI-0002 for study number PISP-00005. Funding was provided by the National Park Service through an agreement with the Rocky Mountains Coopera-tive Ecosystem Studies Unit and the University of Wyoming.

xii Annotated Checklist of Vascular Flora: Pipe Spring National Monument

Chapter 1: Introduction 1

1 Introduction In 2006, the National Park Service (NPS) estab-lished a cooperative agreement with the Univer-sity of Wyoming, through the Rocky Mountains Cooperative Ecosystem Studies Unit, to produce a series of technical reports summarizing the vas-cular plant fl ora of each of the 16 national park units of the Northern Colorado Plateau Network (NCPN). These reports are based on existing in-formation in the park service’s NPSpecies data-base system and augmented by further data min-ing, reviews of state and regional herbaria, and new fi eld surveys to resolve the status of many reported and unconfi rmed species. The objec-tive of these reports is to summarize the current state of knowledge of the fl ora of each park for the benefi t of NPS biologists and interpretive staff , outside researchers, and the general public. The following report addresses the vascular fl ora of Pipe Spring National Monument, in northern Arizona. Other NCPN park units are addressed in separate reports.

1.1 Background and justifi cationOne of the primary missions of NPS is “to con-serve unimpaired the natural and cultural resourc-es and values of the national park system for the enjoyment of this and future generations” (NPS 2000). Due to their high degree of protection and distribution across a wide variety of habitat types, the nation’s national park units make a signifi -cant contribution to the protection of biological diversity (Noss and Cooperrider 1994). Unfortu-nately, the full contribution of the NPS system is not known, as many park units have traditionally lacked comprehensive biological inventories or monitoring programs to identify the composition and condition of their biota. Knowing the status of park fl ora and fauna is critical for eff ective park management and for systematically identifying and fi lling gaps in the nation’s protected area net-work (Margules and Pressey 2000).

To rectify this shortcoming, Congress passed the National Parks Omnibus Management Act in 1998 to provide guidance and funding for an inventory and monitoring program across the entire Na-tional Park System. Since 2000, the NCPN Inven-tory and Monitoring Program has been develop-ing and verifying species lists of vertebrates and vascular plants for each park unit in the network

(Evenden et al. 2002). This eff ort has entailed re-viewing existing species checklists, corroborating the identifi cation of specimens in park museum and herbarium collections, locating additional specimen records from other museums, mining relevant literature and datasets, and conducting fi eld surveys to locate new taxa1 or confi rm un-substantiated reports. All of these data have been synthesized in NPSpecies, the park service’s stan-dardized database for documenting park status (present, probably present, unconfi rmed, falsely reported), abundance, residency, nativity, and rarity. With the production of a technical series of reports on the fl ora of each park, the NPSpecies data will now be available to a broader audience.

1.2 ObjectivesThe objectives of this project are to:

1. Document the current status (present, reported, potential, or falsely reported) of all vascular plant species that have been cited as occurring in Pipe Spring National Monument based on a review of herbarium collections and past literature; and

2. Develop an annotated checklist of the fl ora of Pipe Spring National Monument with supplemental information on taxonomic synonyms, distribution, within-park abun-dance, fl owering period, growth form, nativ-ity, habitat, and sources for each record.

1.3 Study area

1.3.1 History and setting

As one of the few reliable water sources in a large desert area, Pipe Spring has been a focal area of human activity for many centuries. Prior to about 300 B.C., the Arizona Strip was inhabited by semi-nomadic hunter-gatherers who left little physical evidence behind. Over the next thousand years, the region was inhabited by Ancestral Puebloan peoples who engaged in agriculture and hunting and produced abundant artifacts, including pit-house sites, woven baskets, pottery sherds, and arrowheads. Prolonged drought was one likely factor that brought an end to the Puebloan cul-ture in the local area by 1250 A.D. Subsequent inhabitants of the Arizona Strip were semi-no-madic Southern Paiute bands; the Kaibab Paiute occupied the area at the time of contact with

1Taxa refer to all named species, subspecies, and varieties.

2 Annotated Checklist of Vascular Flora: Pipe Spring National Monument

European cultures (McKoy 2000, Leach 1999).

Mormon missionary Jacob Hamblin passed through Pipe Spring in 1858; fi ve years later, St. George, Utah, resident James Whitmore estab-lished a cattle ranch there. In January 1866, Whit-more was murdered during a trip to retrieve live-stock driven off by an Indian party (Leach 1999, McKoy 2000). The Church of Jesus Christ of Latter-Day Saints acquired the property in 1870, constructed a fort (Winsor Castle) that controlled the outlet of Pipe Spring, and managed the site as a tithing ranch. During the 1870s, the native desert grasslands were subjected to grazing by more than 2,500 head of cattle and 11,000 sheep (Alexander 1998a). The ranch passed into private ownership in 1895, and remained as such with the establish-ment of the Kaibab Paiute Indian Reservation in 1909 (McKoy 2000).

While traveling to Zion National Park in 1920, National Park Service Director Stephen Mather hatched the idea of preserving the site as a nation-al monument. With the support of local boosters anticipating improved road access and tourism income, President Warren Harding established Pipe Spring National Monument in May 1923, to “serve as a memorial of western pioneer life.” (McKoy 2000). The monument’s mission has since been expanded to also convey American In-dian culture and history and to protect the spring and associated natural environment (Evenden et al. 2002).

Pipe Spring National Monument is located about 15 miles west of Fredonia in eastern Mohave County, Arizona (Figure 1.3.1). The monument is situated on the northern end of the Uinkaret Plateau, below the Vermilion Cliff s within the Ari-zona Strip (the northern portion of the state cut off by the Grand Canyon). The entire monument covers 40 acres (16 ha) and is embedded within the Kaibab Paiute Indian Reservation. Elevation within Pipe Spring National Monument ranges from 4,923 to 5,100 feet (1,500–1,555 m). Winter temperatures vary from 0 to 40°F, while summer temperatures reach highs of 90–115°F (Evenden et al. 2002).

1.3.2 Geology

The sandstone ridge to the northwest of Win-sor Castle is comprised of Jurassic-age Navajo Sandstone deposited as part of a Sahara-like dune system that extended from Nevada to Wyoming approximately 180 million years ago. The Navajo

formation overlies Kayenta strata derived from early Jurassic river sediments. Navajo Sandstone is more permeable than the Kayenta, and the con-tact between the two formations is notable as a source of springs throughout the Colorado Pla-teau. The juxtaposition of this contact zone with the Sevier Fault (separating the mesa from the adjacent Moenkopi and Chinle beds to the east) accounts for the size and fl ow of the three springs in the monument (Pipe Spring, itself, and the smaller Tunnel and Cabin springs). Holocene-age alluvium covers the eastern and southern ends of the monument, burying the Triassic Chinle and Moenkopi formations (Billingsley 2004).

1.3.3 Vegetation

Unfortunately, the pre-settlement vegetation of Pipe Spring and the Arizona Strip was never stud-ied in detail using modern scientifi c methods. Anecdotal reports by local ranchers describe the Strip as a “sea of grass” in the late 1860s (Alex-ander 1998a). In 1882, geologist Charles Dut-ton wrote that the Pipe Spring area had changed from being “covered with abundant grasses, af-fording rich pasturage to horses and cattle” to a place where “hardly a blade of grass is to be found within ten miles of the spring, unless upon the crags and mesas of the Vermilion Cliff s behind it” in just 10 years (Dutton 1882, cited in Alexander 1998a). Based on historic diaries, archives, and photographs, Alexander (1998a) reported that the mesa northwest of Pipe Spring was devoid of trees by the time NPS took over management in 1923, and the fl ats surrounding the fort were mostly bare ground.

Vegetation has slowly recovered in the monu-ment. Today, three main plant communities can be recognized. Sandy fl ats and south-facing mesa slopes are inhabited by desert shrub vegetation, dominated by greenish rabbitbrush (Chrysotham-nus nauseosus), fourwing saltbush (Atriplex ca-nescens), and sand sagebrush (Artemisia fi lifolia) (Alexander 1998b). Steeper Navajo Sandstone slopes are dominated by open Utah juniper (Juni-perus osteosperma) and two-needle pinyon (Pinus edulis) woodlands with an understory of turbi-nella live oak (Quercus turbinella) and Utah ser-viceberry (Amelanchier utahensis). Wet meadow vegetation consisting of baltic rush (Juncus arcti-cus), rabbitbrush, and scratchgrass (Muhlenbergia asperifolia) occurs below Cabin Spring (by West Cabin). A human-made pond, located just out-side Winsor Castle, is dominated by planted white poplar (Populus alba), Fremont cottonwood (P.


Figure 1.3.1. Location of Pipe Spring National Monument, Mohave County, Arizona. Map created by Aneth Wight of NCPN (derived from Evenden et al. 2002).

To St. George, UT

Visitor Center

To Fredonia, AZ389

Established May 31, 19231,500−1,555 meters elevation(4,923−5,100 feet elevation)

16 hectares (40 acres)0 0.1 0.2 0.3

Kilometers

0 0.1 0.2 0.3Miles

Legend:

Intermittent Streams

Paved Road

Unpaved Road

Unimproved/4WD Road

Park Boundary (NPS)

Kaibab Reservation (IR)

Pipe Spring National MonumentUtah


fremontii), Carolina poplar (P. x canadensis), Si-berian elm (Ulmus pumila), and tree-of-heaven (Ailanthus altissima). Much of the remainder of the grounds is dedicated to cultivated orchards and a replica pioneer garden.

1.3.4 Previous botanical studies

The fi rst botanist to visit Pipe Spring was proba-bly Ellen Powell Thompson, sister of John Wesley Powell and resident of Kanab, Utah, in the early 1870s (Alexander 2004). It is not known wheth-er Thompson collected specimens at the future monument. Marcus E. Jones investigated the fl ora of Kanab and the Arizona Strip in the 1880s, but it is also unknown whether he specifi cally collect-ed at Pipe Spring. Albert Siler, a pioneer rancher and naturalist in southern Utah, collected the type specimen of Utahia sileri (later renamed Pediocactus sileri), Siler’s pincushion cactus, from “Cottonwood Springs and Pipe Springs, southern Utah” (actually Arizona) in 1883 (Benson 1982). This collection is now believed to have come from just outside the monument on the Kaibab Paiute Indian Reservation, as the exacting habitat condi-tions required by this rare cactus are not present within monument boundaries.

The holotype of Pipe Spring pricklypear (Opun-tia erinacea var. aurea, O. basilaris var. aurea, or O. aurea, depending on one’s taxonomic pref-erence) was collected “1/2 mile north of Pipe Spring” by McCabe and McCabe in 1930 (Ben-son 1982). This locality is either within or barely outside the monument’s boundary, but the spe-cies is well documented elsewhere in Pipe Spring National Monument. If accepted as being col-lected in the monument, the McCabe collection would stand as the earliest known specimen from the area (Table 1.3.4, Figure 1.3.4).

Leonard Heaton, custodian of Pipe Spring Na-tional Monument (PISP), began collecting plants for the PISP herbarium in 1934. Over the next seven years, Heaton and J. Whitehead collected 112 specimens for the herbarium, representing 98 species. Heaton (1935) compiled the fi rst check-list of monument fl ora in 1935, citing 42 species and estimating that another 100 remained to be

discovered (in hindsight, a reasonable prediction). Robert Peebles (co-author of Arizona Flora) and A. R. Purchase added six new species in 1936, and later found two more in a 1940 visit—including the fi rst report of Opuntia pinkavae (Pinkava’s pricklypear), not recognized as a new species to science until 1997. Utah botanist Walter Cottam and Kathy Parker also collected a few new taxa in the 1940s.

Over the next quarter century, relatively little bo-tanical work was done in the monument, except for some sporadic study in the early to mid-1960s. Ruth Nelson visited Pipe Spring twice starting in 1971, and added fi ve new species to the fl ora. Richard King, a range conservationist with the Fredonia offi ce of the Soil Conservation Service, compiled a species list in 1977 that included 82 plant species (with an emphasis on shrubs and grass species) (King 1977). King’s list contained 39 new species for the monument, of which all but 18 have since been confi rmed with vouchers.

The most intensive botanical surveys of Pipe Spring National Monument since the mid-1930s have taken place in the last decade. Zion National Park biologists Karen Mason and Bill LaBarre, who conducted a noxious-weed inventory in 1997, reported 13 new species for the monument (Kurth 1998). In 1998, Jason Alexander was hired by the NPS to research vegetation change within the monument and compile a revised species list. Alexander (1998b) documented 166 taxa for Pipe Spring National Monument, of which 63 repre-sented new species. In 2001, Alexander, Denise Louie of Zion National Park, and Angie Even-den of the NCPN staff in Moab, Utah, collected 116 vouchers for the PISP herbarium (more than doubling it in size) and recorded 22 new species for the fl ora. This team also verifi ed 60 previous literature reports with specimen vouchers (Table 1.3.4). More recently, we, along with Janet Coles of the NCPN vegetation staff , found 10 new spe-cies for the monument in fall 2007, and relocated 18 more species known only from historical or unvouchered reports.

Additional details on plant-collecting eff orts at PISP can be found in Table 1.3.4.


Table 1.3.4. Chronology of plant-collecting efforts in Pipe Spring National Monument.

Year Collector(s)# of new

taxa Comments

1883 A. L. Siler 0 Albert L. Siler collected the holotype of Pediocactus sileri from “Cottonwood Springs and Pipe Springs, southern Utah” [Arizona] in 1883. Based on the specifi c habitat requirements of this species, it is now believed that the collection came from the adjacent Kaibab Paiute Indian Reservation rather than Pipe Spring National Monument, itself.

1930 McCabe and McCabe

1 McCabe and McCabe collected the holotype of Opuntia erinacea var. aurea “1/2 mile north of Pipe Spring” within or very near Pipe Spring National Monument.

1934–1939 L. Heaton, J. Whitehead, A. R. Purchase, and R. H. Peebles

98 + 7 reported

Leonard Heaton, custodian of Pipe Spring National Monument, compiled a vascular plant species list with 42 species in 1935 and noted “there are about 100 more plants that have not been [n]amed or classifi ed...” (Heaton 1935). All but 7 of these species have been subsequently vouchered. From 1934–1939, Heaton and J. Whitehead collected 104 voucher specimens for the PISP herbarium and documented 92 plant taxa. In 1936, Robert H. Peebles discovered 4 new species for the monument and A.R. Purchase added 2 new grass species.

1940–1946 R. H. Peebles, W. Cottam, L. Heaton, and K. F. Parker

9 R. H. Peebles discovered 2 new species for PISP in 1940, including the fi rst record of Opuntia pinkavae (later described as a new taxon by Bruce Parfi tt in 1997). Walter Cottam and Leonard Heaton found 6 new taxa in 1941, including the earliest report of Lepidium montanum var. stellae (Stella’s pepperwort; later described as a new taxon by Stan Welsh in 1978). Kathy Parker discovered Convolvulus arvensis (fi eld bindweed) in the monument in 1946.

1961 F. Fanning 1 F. Fanning collected Amsonia jonesii (Jones’ amsonia) in the monument (specimen at the Desert Botanical Garden herbarium).

1965–1967 E. Lehto and M. Ganz

5 Ganz and Lehto made at least 16 collections at Pipe Spring (deposited at Arizona State University, University of Arizona, and Northern Arizona University).

1971–1973 R. A. Nelson and M. G. MacLeod

5 Ruth Nelson and M. G. MacLeod made collections at PISP during visits in 1971 and 1973 (specimens are deposited at Arizona State University and the University of Wyoming).

1977 R. J. King 21 + 18 reported

Richard King, range conservationist with the Soil Conservation Service in Fredonia, Arizona, compiled an annotated checklist of 82 species for the monument (King 1977), including 39 that had not previously been reported. In subsequent years, 21 of King’s new taxa were documented with vouchers.

1995 A. M. Phillips III 1 Arthur Phillips made 9 collections in the monument, including the fi rst of Rhus aromatica var. simplicifolia (squawbush).

1997 K. Mason and B. LaBarre

8 + 5 reported

Zion NP biological science technicians Karen Mason and Bill LaBarre conducted a weed survey of PISP, reporting 13 new species for the area (Kurth 1998), of which 8 have been subsequently vouchered.

1998 J. Alexander 34 + 29 reported

Alexander (1998b) examined historic records and conducted a ground survey of the monument to compile a revised checklist of 166 vascular plant taxa for PISP. This list included 63 new species, of which 33 were later vouchered (29 are still known only from literature reports).

2001 J. Alexander, D. Louie, and A. Evenden

22 Jason Alexander, Denise Louie, and Angie Evenden collected 116 voucher specimens (doubling the size of the PISP herbarium) and documented 22 new species as well as making vouchers of over 60 previously reported taxa.

2007 W. Fertig and J. Coles

10 Walter Fertig and Janet Coles conducted surveys and vegetation sampling and added 33 specimens for the PISP herbarium, including 10 new to the monument and 18 that were previously considered historical or had been reported without a voucher.


Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Pipe Spring National Monument by decade, 1930–2008.

Nu

mb

er o

f va

scu

lar

pla

nt

taxa

250

200

150

100

50

01930 1940 1950 1960 1970 1980 1990 2000 2008

Year

Chapter 2: Methods 7

2 MethodsUnder contract with the National Park Service, Jason Alexander reviewed all mounted speci-mens in the PISP herbarium in 2004 to determine whether each was correctly identifi ed to species, subspecies, or variety following the nomenclature of Welsh et al. (2003). As necessary, he made cor-rections and annotations to each specimen and to the voucher records in PISP’s Automated Nation-al Cataloging System (ANCS+) and NPSpecies database. Based on Alexander’s herbarium work, we developed a revised checklist of species that were present in the monument (confi rmed by an authenticated voucher) and those that were re-ported in the literature but not corroborated with a voucher (equivalent to “probably present” in NPSpecies terminology). Species that were now known to be misidentifi ed or highly questionable were moved to a third list of rejected taxa. Finally, we developed a list of potential species (“uncon-fi rmed” in NPspecies) known from the vicinity of Pipe Spring National Monument using an unpub-lished list generated by Dr. N. Duane Atwood of Brigham Young University.

Each list was annotated with supplemental infor-mation on life form, geographic range (within the context of Arizona), population size in the monu-ment (derived from Jason Alexander’s NPSpecies dataset for PISP and fi eld observations), fl ower-ing period, distribution across general vegetation types, nativity, data source (collector and collec-tion number from a sample specimen or literature citation), year of earliest collection (from herbar-

ium records), and relevant synonymy and taxo-nomic problems (based on review of Barkworth et al. 2003, 2007, Barneby 1989, Cronquist 1994, Cronquist et al. 1972, 1977, 1984, 1997, Dorn 2001, Flora of North America Editorial Commit-tee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c, Holmgren et al. 2005, Mc-Dougall 1973, and Weber and Wittmann 2001). The annotations to these lists and label data from the PISP herbarium review were then incorpo-rated into the NPSpecies database for the monu-ment.

Work in 2007 focused on confi rmation of report-ed and potential species for the monument fl ora through fi eld surveys and review of collections from state and regional herbaria. We made one collecting trip to PISP, in October 2007, to target missing species, emphasizing under-sampled wet-land, weedy, and late-fl owering taxa. Herbarium studies were undertaken at Brigham Young Uni-versity, Utah State University, and the University of Wyoming in March, June, and October 2006, and February and March 2007. We also docu-mented monument specimens from the digital herbarium database of Arizona State University (http://seinet.asu.edu/seinet/collections/selec-tion.jsp?cat=plantae) and its links to comparable databases for the Desert Botanical Garden, Uni-versity of Arizona, and Northern Arizona Univer-sity. Finally, we conducted an internet search of the virtual herbarium database of the New York Botanical Garden (http://sciweb.nybg.org/sci-ence2/hcol/intf/index.asp) and examined plot data from the NCPN vegetation mapping crew (NCPN, in ed.).

Chapter 3: Results 9

3 Results

3.1 Summary of the fl ora of Pipe Spring National Monument

In 2004, Jason Alexander completed a thorough review of the 227 vascular plant vouchers in the PISP herbarium (only 112 of which were col-lected prior to 2001). He found that 8 specimens were misidentifi ed (3.5%), 3 needed to be revised to a more current name (1.3%), 27 needed vari-ety or subspecies names added (11.9%), and 189 were correctly determined (83.3%). Based solely on these collections, the confi rmed fl ora of Pipe Spring National Monument stood at 162 taxa. From 2005 to 2007, we located an additional 27 taxa from collections at the University of Wyo-ming’s Rocky Mountain Herbarium and speci-

mens found through a database search of the Arizona State University digital database. Field surveys in 2007 netted another 26 new species (including 16 that were previously reported with-out a voucher). Overall, the documented fl ora of Pipe Spring National Monument has increased by 53 species (24.7%), bringing the total number of vouchered taxa to 215 (Table 3.1, Appendices A, D).

Currently, 62 additional taxa have been reported for PISP but have yet to be corroborated with a voucher specimen (Table 3.1, Appendix A). Of these, 7 were reported by Heaton (1935), 18 by King (1977), 5 by Kurth (1998), and 32 by Alexan-der (1998b). Taken together, the number of con-fi rmed and reported vascular plant taxa in Pipe Spring National Monument stands at 277 taxa (Table 3.1, Appendix A).

Table 3.1. Statistical summary of the fl ora of Pipe Spring National Monument.

CategoryNumber

confi rmed in park

Number additionally

reported for parkTotal

Taxonomic diversity

Total taxa (including varieties and subspecies)

215 62 277

Full species (excluding varieties and subspecies)

203 59 262

Families 54 2 56

Life form diversity

Tree taxa 8 2 10

Shrub taxa 34 6 40

Perennial forb taxa 72 26 98

Annual forb taxa 72 9 81

Perennial graminoid taxa 21 15 36

Annual graminoid taxa 8 4 12

Fern taxa 0 0 0

Biogeographic diversity

Introduced taxa 35 23 58

Native taxa

Locally endemic taxa 4 0 4

Regionally endemic taxa 14 7 21

Disjunct taxa 0 0 0

Peripheral taxa 4 1 5

Sparse taxa 1 0 1

Widespread taxa 157 31 188

Total native taxa 180 39 219The number of taxa and families is based on taxonomic concepts of Welsh et al. (2003). See Appendices, “Codes and Defi nitions”, for an explanation of the various categories.


Entering the 2007 fi eld season, 41 taxa were rec-ognized as potentially occurring in PISP based on the proximity of the monument to known populations (Alexander NPSpecies database). Including fi ve additional species identifi ed by Duane Atwood, the current number of potential species for Pipe Spring National Monument is 46 (Appendix B).

Based on his evaluation of the PISP herbarium, Jason Alexander found 8 taxa that were falsely re-ported due to misidentifi cation. We consider an additional seven species cited (but not vouchered) by Heaton (1935) and King (1977) to be question-able based on the absence of suitable habitat or the distance of Pipe Spring National Monument to the known or expected range of each species (Appendix C).

The confi rmed and reported fl ora of Pipe Spring National Monument represents 6.5% of the 4,241 native and naturalized taxa documented for the state of Arizona by Kartesz (2003) and 34.8% of the state’s 161 plant families (Table 3.1). Within Mohave County, the monument fl ora captures 17.4% of the 1,588 taxa reported by Kartesz (2003). At the ecoregional scale, Pipe Spring Na-tional Monument contains 11.0% of the 2,517 species of seed plants and pteridophytes reported for northern Arizona by McDougall (1973) and the Flora of North America Editorial Committee (1993).

3.2 Life form and biogeographic diversity

Although ecologically dominant, tree and shrub species make up only 18% of the confi rmed and reported plant taxa of Pipe Spring National Mon-ument (Table 3.1). Perennial forbs are the most species-rich life form in the monument’s fl ora, with 98 known and reported taxa (35.4% of the total fl ora). Annual forbs and grass-like plants are unusually species-rich in the monument (relative to other fl oras in the Colorado Plateau), compris-ing 33.5% of all taxa. Like trees and shrubs, pe-rennial graminoids are ecologically important but relatively species-poor, representing just 13% of the total fl ora. No ferns or fern-allies have been found in PISP (Table 3.1).

Of the monument’s 219 native taxa, 188 (85.8%) are widespread across Arizona and western North America. Only four species (1.8% of the monument’s native fl ora) have extremely limited global distributions (local endemics found within

an area of less than 1° of latitude × 2° of longi-tude; see Appendix A for complete defi nitions of biogeographic categories). Regional endemics (largely restricted to the Colorado Plateau) com-prise an additional 9.6% of the monument fl ora (Table 3.1). Species at the edge of their range (pe-ripheral) or found sparsely across the state make up just 2.8% of the total fl ora.

3.3 Plants of special concernPipe Spring National Monument has no plants listed as threatened or endangered under the U.S. Endangered Species Act (ESA). Siler’s pincush-ion cactus (Pediocactus sileri), listed as threatened under the ESA, is known from adjacent areas of the Kaibab Pauite Indian Reservation (Siler’s type specimen comes from “Pipe Spring”), but is not thought to occur within the monument itself due to a lack of suitable gypsiferous Moenkopi soils (ARPC, no date). The federally threatened Jones’ cycladenia (Cycladenia humilis var. jonesii) is also known from Chinle soils west of Pipe Spring, but is unlikely to be found in the monument.

The Arizona Game and Fish Department (AGF)’s Heritage Data Management System (2008) tracks three cactus species from PISP as species of con-cern (Table 3.3). All are listed as Salvage Restrict-ed (collection prohibited except with a permit) under the 1993 Arizona Native Plant Law (ARPC, no date). Atwood’s wild-buckwheat (Eriogonum thompsoniae var. atwoodii) and Morton wild-buckwheat (E. mortonianum) are also tracked as species of concern by AGF and known from the Kaibab Paiute Indian Reservation, but have not been documented in Pipe Spring National Monu-ment.

Two recently described species with narrow geo-graphic ranges currently lack any protective status but might warrant management attention. Stella’s pepperwort (Lepidium montanum var. stellae) is restricted to southern Utah and northern Arizona (Welsh et al. 2003), where it typically occurs on gypsum-rich, Moenkopi-derived soils. Leon-ard Heaton collected this species at or near Pipe Spring in 1941 (suitable habitat is more likely just outside the monument), but it remained uniden-tifi ed until annotated by Jason Alexander in 2004. Pinkava’s pricklypear (Opuntia pinkavae) is also restricted to the Utah–Arizona border and was fi rst recorded at Pipe Spring in 1940 by Robert Peebles. The holotype of O. pinkavae comes from Bulrush Canyon, south of Pipe Spring (Parfi tt 1997).

Chapter 3: Results 11

3.4 Noxious weedsIntroduced plant species (not historically native to Arizona or North America) make up 20.9% of the confi rmed and reported fl ora of Pipe Spring National Monument (Table 3.1). This fi gure is substantially higher than the statewide average of 12.7% non-native species (Kartesz 2003). Of the 58 introduced plant species confi rmed or re-ported for PISP (Table 3.1), only four are listed as noxious by the State of Arizona (ADAPSD 2005). Scotch thistle (Onopordum acanthium) is on the state’s prohibited and restricted noxious weed lists, while fi eld bindweed (Convolvulus arven-sis), common purslane (Portulaca oleracea), and puncturevine (Tribulus terrestris) are listed as prohibited and regulated noxious weeds.

Kurth (1998) cited 34 non-native species of man-agement concern, based on a 1997 survey of the monument by Karen Mason and Bill LaBarre of Zion National Park. These consisted primar-ily of introduced ornamental trees that have be-come weedy or exotic forbs and grasses that are common agricultural or rangeland pests. Using a matrix of signifi cance of impact and feasibility of control, Kurth classifi ed these species into four groups: low threat/easy to control, low threat/hard to control, serious threat/easy to control, and serious threat/hard to control. Four species were recommended for immediate control: tree-of-heaven (Ailanthus altissima), white poplar (Populus alba), tamarisk (Tamarix chinensis), and Siberian elm (Ulmus pumila).

Table 3.3. Plant species of conservation concern from Pipe Spring National Monument.

Family Species Common nameTNC global rank1 Federal legal status AZ status2

Cactaceae Opuntia erinacea var. aurea (O. basilaris var. aurea)

Pipe Spring pricklypear

G3 None Salvage Restricted, S3

Cactaceae Opuntia whipplei var. whipplei Whipple’s cholla G4?T4? None Salvage Restricted, S1

Cactaceae Sclerocactus whipplei var. roseus (S. parvifl orus ssp. intermedius)

small-fl ower fi shhook cactus

G4T3? None Salvage Restricted, S2

This list is derived from Arizona Game and Fish Department Heritage Data Management System (2008) and Arizona Rare Plant Committee (no date).1TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank. A comparable scoring system is used to measure state abundance (S) in Arizona under the column “AZ status”.2AZ status includes protection afforded under the 1993 Arizona Native Plant Law.

Chapter 4: Discussion 13

4 DiscussionBeginning with Leonard Heaton’s fi rst species list in 1935, the known fl ora of Pipe Spring National Monument has grown from 42 species to 277 taxa (Tables 1.3.4 and 3.1, Figure 1.3.4, Appendix A). Based on available habitats and proximity, at least another 46 plant taxa may occur within the monument (Appendix B). Should all of these po-tential species be found, the documented fl ora of PISP would increase by nearly 14%. Many of the “missing” potential or reported taxa consist of cryptic species that fl ower late in the growing sea-son, or occur in undersampled wetland or weedy habitats. Confi rmation of reported species should remain a high priority, as some of these taxa may ultimately prove to be erroneous. Pipe Spring also has a high percentage of plants known only from

historic records (pre-dating 1970) that should be a priority for re-location to confi rm that they are still extant. Further examination of monu-ment collections from other herbaria in Arizona or major national collections (such as Harvard University’s Gray Herbarium, the New York Bo-tanical Garden, and the Smithsonian Institution’s U.S. National Herbarium) may result in the confi -mation of additional records of historical species for the monument. Additional fi eld studies would also be useful to better refi ne the abundance and distribution of species of high management interest (especially rare plants and invasive non-natives). Surveys following wet winters would likely lead to the discovery of new annual native and exotic species. Lastly, better documentation of plant species previously or currently grown in the monument’s orchard and pioneer garden is needed.

Chapter 5: Literature Cited 15

5 Literature CitedAlexander, J. 1998a. Pipe Spring National Monu-

ment presettlement vegetation literature survey. Pipe Spring National Monument, Az. 43 pp.

—— . 1998b. Pipe Spring National Monument master vegetation species list. Report pre-pared for the National Park Service, Pipe Spring National Monument.

—— . 2004. A summary of fl oristics at Pipe Spring National Monument. Pipe Spring National Monument, Az. 3 pp.

Arizona Department of Agriculture, Plant Ser-vices Division (ADAPSD). 2005. Prohibited, regulated and restricted noxious weeds.

Arizona Game and Fish Department Heritage Data Management System. 2008. Special status species by taxon, scientifi c name. http://www.azgfd.gov/w_c/edits/hdms _spe-cies_lists.shtml

Arizona Rare Plant Committee (ARPC). No date. Arizona rare plant fi eld guide: A collabora-tion of agencies and organizations.

Barkworth, M. E., K. M. Capels, S. Long, and M. B. Piep, eds. 2003. Magnoliophyta: Com-melinidae (in part): Poaceae, part 2. Flora of North America north of Mexico, volume 25. New York: Oxford University Press. 783 pp.

Barkworth, M. E., K. M. Capels, S. Long, L. K. Anderton, and M. B. Piep, eds. 2007. Mag-noliophyta: Commelinidae (in part): Poa-ceae, part 1. Flora of North America north of Mexico, volume 24. New York: Oxford University Press. 911 pp.

Barneby, R. C. 1989. Volume 3, Part B. Fabales. In Cronquist, A., A. H. Holmgren, N. H. Holmgren, J. L. Reveal, and P. K. Holmgren, eds., Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 279 pp.

Benson, L. 1982. The cacti of the United States and Canada. Stanford, Ca.: Stanford Univer-sity Press. 1044 pp.

Billingsley, G. H., S. S. Priest, and T. J. Felger. 2004. Geologic map of Pipe Spring National Monument and the western Kaibab-Pai-ute Indian Reservation, Mohave County, Arizona. Pamphlet to accompany Scientifi c Investigations Map 2863. U.S. Geological

Survey, U.S. Department of the Interior.

Cronquist, A. 1994. Volume 5, Asterales. In Cronquist, A., A. H. Holmgren, N. H. Holmgren, J. L. Reveal, and P. K. Holmgren, eds., Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 496 pp.

Cronquist, A., A. H. Holmgren, N. H. Holmgren, and J. L. Reveal, eds. 1972. Volume 1, Geo-logical and botanical history of the region, its plant geography and a glossary: The vascular cryptogams and the gymnosperms. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 270 pp.

Cronquist, A., A. H. Holmgren, N. H. Hol-mgren, J. L. Reveal, and P. K. Holmgren, eds. 1977. Volume 6, The Monocotyledons. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 584 pp.

—— . 1984. Volume 4, Subclass Asteridae (except Asteraceae). Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 573 pp.

Cronquist, A., N. H. Holmgren, and P. K. Hol-mgren, eds. 1997. Volume 3, Part A, Subclass Rosidae (except Fabales). Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botani-cal Garden. 446 pp.

Dorn, R. D. 2001. Vascular plants of Wyoming. Third edition. Cheyenne, Wy.: Mountain West Publishing. 412 pp.

Evenden, A., M. Miller, M. Beer, E. Nance, S. Daw, A. Wight, M. Estenson, and L. Cudlip. 2002. Northern Colorado Plateau Vital Signs Network and Prototype Cluster, plan for natural resources monitoring: Phase I Report [two volumes]. National Park Ser-vice, Northern Colorado Plateau Network, Moab, Ut. 138 pp + app.

Flora of North America Editorial Committee. 1993. Pteridophytes and gymnosperms. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 475 pp.

—— . 1997. Magnoliophyta: Magnoliidae and Hamamelidae. Flora of North America north of Mexico, Volume 2. New York: Ox-


ford University Press. 590 pp.

—— . 2000. Magnoliophyta: Alismatidae, Are-cidae, Commelinidae (in part), and Zingi-beridae. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 352 pp.

—— . 2002a. Magnoliophyta: Commelinidae (in part): Cyperaceae. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 608 pp.

—— . 2002b. Magnoliophyta: Liliidae: Liliales and Orchidales. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 723 pp.

—— . 2003. Magnoliophyta: Caryophyllidae, part 1. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 559 pp.

—— . 2005. Magnoliophyta: Caryophyllidae, part 2. Flora of North America north of Mexico, Volume 2. New York: Oxford University Press. 656 pp.

—— . 2006a. Magnoliophyta: Asteridae, part 6: Asteraceae, part 1. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 579 pp.

—— . 2006b. Magnoliophyta: Asteridae, part 7: Asteraceae, part 2. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 666 pp.

—— . 2006c. Magnoliophyta: Asteridae, part 8: Asteraceae, part 3. Flora of North America north of Mexico, Volume 2. New York: Ox-ford University Press. 616 pp.

Heaton, L. 1935. Plants from Pipe Spring Na-tional Monument.

Holmgren, N. H., P. K. Holmgren, and A. Cron-quist. 2005. Volume 2, Part B Subclass Dille-niidae. Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 488 pp.

Kartesz, J. T. 2003. Synthesis of the North Ameri-can fl ora, Version 2.0. http://www.bonap.org/synth.html.

Kearney, T. H., and R. H. Peebles. 1960. Arizona fl ora. Second edition. Berkeley: University of California Press. 1085 pp. (with supplement by J. T. Howell, E. McClintock, and collabo-rators).

King, R. J. 1977. The fl ora of Pipe Spring Na-tional Monument. Report prepared for the National Park Service by the Soil Conserva-tion Service, Fredonia, Arizona.

Kurth, L. 1998. Pipe Spring National Monu-ment non-native plants survey report, 1997. Report prepared for Pipe Spring National Monument by Zion National Park. 3 pp.

Leach, N. 1999. Pipe Spring National Monu-ment: An ancient oasis on a storied frontier. Springdale, Ut.: Zion Natural History As-sociation.

Margules, C. R., and R. L. Pressey. 2000. Sys-tematic conservation planning. Nature 405:243–253.

McDougall, W. B. 1973. Seed plants of northern Arizona. Flagstaff : Museum of Northern Arizona. 594 pp.

McKoy, K. L. 2000. Cultures at a crossroads: An administrative history of Pipe Spring National Monument. Cultural Resources Selections No. 15, Intermountain Region. Denver, Co.: National Park Service.

National Park Service (NPS). 2000. Management Policies. Washington, D.C.: U.S. Department of Interior, National Park Service. Publica-tion # NPS D1416. 137 pp.

Northern Colorado Plateau Network (NCPN). in ed. Pipe Spring National Monument veg-etation classifi cation and mapping project.

Noss, R. F., and A. Y. Cooperrider. 1994. Saving nature’s legacy: Protecting and restoring biodiversity. Washington, D.C.: Island Press. 416 pp.

Parfi tt, B. D. 1997. Opuntia pinkavae (Cacta-ceae), a new species from Arizona and Utah. Rhodora 99:223–228.

Weber, W. A., and R. C. Wittmann. 2001. Colo-rado fl ora: Western Slope. Third edition. Boulder: University Press of Colorado. 488 pp.

Welsh, S. L., N. D. Atwood, S. Goodrich, and L. C. Higgins. 2003. A Utah fl ora. Third edition, revised. Provo, Ut.: Brigham Young Univer-sity. 912 pp.

Appendices 17

6 AppendicesThe following appendices contain annotated checklists of the vascular fl ora of Pipe Spring Na-tional Monument.

Appendix A lists all species that have been con-fi rmed for the fl ora with an authenticated vouch-er specimen (including extant and historical re-cords), or that have been reliably reported for the monument in the literature (Heaton 1935, King 1977, Alexander 1998b). Vouchers were exam-ined from the PISP herbarium and collections from Brigham Young University (BRY), the Uni-versity of Wyoming (RM), and the digital muse-um databases of the New York Botanical Garden (NY), Northern Arizona University, Arizona State University, University of Arizona, and the Desert Botanical Garden. Species entries in Appendix A are organized alphabetically by family and genus.

Species listed in Appendix B are known from the vicinity of Pipe Spring National Monument, but have not previously been reported or confi rmed for the monument; that is, these species may po-tentially occur within PISP, based on the prox-imity of the monument to other populations or the presence of suitable habitat. Species on this list are derived from a checklist of potential spe-cies for NCPN parks prepared by Dr. N. Duane Atwood, of Brigham Young University, in 2001, and Jason Alexander for the PISP NPSpecies da-tabase. Some species from Appendix C could be considered as potentially occurring in the monu-ment, but are not included here.

Appendix C lists species that have been previ-ously reported for Pipe Spring National Monu-ment from published and unpublished literature (Alexander 1998a, Heaton 1935, King 1977) or from herbarium specimens, but are now consid-ered erroneous because they were based on mis-identifi ed specimens (False Reports or FalsRep), questionable literature records, or because the monument is well outside the known or expected range of the species (Questionable or Ques?).

Appendix D contains the same confi rmed, his-torical, and reported taxa as in Appendix A, but organized by life form rather than by family in or-der to better meet the needs of NPS interpretive specialists.

The codes and references for each fi eld in the ap-propriate appendices are summarized below.

Family (Appendices A, B, C, and D)Family concepts and nomenclature follow A Utah Flora (Welsh et al. 2003). Alternative family names and treatments from other regional fl oras (Dorn 2001, Flora of North America Editorial Commit-tee 1993, 2002b, Holmgren et al. 2005, Kearney and Peebles 1960, McDougall 1973,Weber and Wittman 2001) are listed in parentheses.

Species name (Appendices A, B, C, and D)Nomenclature for scientifi c names and taxonom-ic concepts are derived from Welsh et al. (2003). Scientifi c names are listed in alphabetical rather than phylogenetic order.

Synonyms/Taxonomic notes (Appendices A, B, and C)Pertinent synonyms are included for species with alternative scientifi c names in other recent state and regional fl oras (Barkworth et al. 2003, 2007, Barneby 1989, Cronquist 1994, Cronquist et al. 1972, 1977, 1984, 1997, Dorn 2001, Flora of North America Editorial Committee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c, Holmgren et al. 2005, Kearney and Peebles 1960, McDougall 1973, and Weber and Wittmann 2001). No eff ort has been made to identify ad-ditional synonyms from monographs and fl oras preceding the early 1970s.

Common name (Appendices A, B, C, and D)Common names are derived primarily from Welsh et al. (2003) and the Intermountain Flora (Barneby 1989, Cronquist 1994, Cronquist et al. 1972, 1977, 1984, 1997, Holmgren et al. 2005).

Life form (Appendices A, B, C, and D)Life form is based on the stature and growth habit of typical members of a species at reproductive maturity. Annual Forbs (AnnF) are broad-leaved dicots or monocots with non-woody stems that complete their life cycle (mature, fl ower, and die) in one year. Annual Graminoids (AnnG) are lin-ear-leaved, grass-like monocots that complete their life cycle (mature, fl ower, and die) in one year. Ferns and fern-allies (Fern) are non-fl ow-ering vascular plants that reproduce by spores. Perennial Forbs (PerF) are broad-leaved dicots or monocots with non-woody stems (at least above


ground level) that live for multiple years. Perenni-al Graminoids (PerG) are linear-leaved, grass-like monocots that live for multiple years. Shrubs are woody perennials with one to many trunks aris-ing from near the base of the plant and are usually less than 3.5 m tall. Trees are woody perennials with a single stem or trunk and are typically over 3.5 m tall.

Range (Appendices A, B, and C)Range represents the distribution of a species within Arizona and the state’s contribution to the plant’s overall global range. Disjunct (Disj) taxa have their Arizona distribution separated from the main, contiguous portion of their range by a gap of more than 800 km (ca. 500 miles). Intro-duced (Intro) taxa are those that are not native to Arizona or North America but have become naturalized (breeding on their own without hu-man assistance). Local Endemics (LocEn) are taxa in which the entire global range is restricted to an area of less than 16,500 km2 (ca. 6,370 mi2, or 1° of latitude × 2° of longitude). Peripheral (Periph) taxa have a widespread global distribu-tion but occur at the margin of their contiguous range in Arizona and occupy less than 5% of the state’s area (usually only within a few miles of the state border). Regional Endemics (RegEn) are taxa with a global range of 16,500–250,000 km2 (an area about the size of the state of Wyoming). Sparse taxa occur widely across Arizona or North America but their range within Arizona is small and patchy, with populations restricted to special-ized or uncommon habitats. Widespread (Wide) taxa have global ranges exceeding 250,000 km2 and occur over at least 10% of the state.

Park status (Appendices A, C, and D)Park status identifi es whether a species is Pres-ent (Pres), Historical (Hist), or Reported (Rep) for Pipe Spring National Monument. Species are considered Present if a confi rmed voucher or documented observation has been made within monument boundaries since 1970. Historical species have been confi rmed with a voucher from the monument but have not been relocated since before 1970. Reported species are listed for the monument in the literature (Heaton 1935, King 1977, Alexander 1998b), but have not been cor-roborated with a voucher specimen. Species that may potentially occur in PISP are listed in Appen-dix B. Rejected taxa (listed in Appendix C) have either been falsely reported (FalsRep) in the lit-erature based on misidentifi ed herbarium speci-

mens or are considered Questionable (Ques?) because park reports are from well outside their known or expected range.

Pop. size (Appendix A)Population size refers to the abundance and fre-quency with which a species occurs within Pipe Spring National Monument. Abundance data are derived from the PISP NPSpecies database and personal observations. Abundant (Abun) taxa have large populations, have a broad ecological amplitude (often being dominant in one to sev-eral vegetation types), or occur extensively across the monument. Common (Com) taxa have large local populations but may be restricted to a single vegetation type or do not occur across the entire monument. Uncommon (Unc) taxa have low to medium-sized populations, typically have a nar-row ecological amplitude (or at least contribute relatively little to total vegetative cover), or have a small range across the monument. Rare taxa usually have low population numbers and are re-stricted to a single location within the monument. A “?” indicates that population size is unknown.

Source (Appendix A)Sources are the evidence for the assigned park status. In the case of Present and Historical taxa, sources are herbarium records (with collector, collection number, and repository) that were cor-roborated during this study. For reported taxa, the source is a published or unpublished refer-ence citing the species within the monument. Specimen repositories include the Pipe Spring National Monument herbarium (PISP), Brigham Young University (BRY), Utah State University (USU), New York Botanical Garden (NY), the University of Wyoming’s Rocky Mountain Her-barium (RM), Northern Arizona University, Ari-zona State University, University of Arizona, and Desert Botanical Garden.

Year doc. (Appendix A)Year documented indicates the fi rst year in which a species was confi rmed or reported to occur in Pipe Spring National Monument. This year does not necessarily correspond with the date of the voucher specimen cited under Source. “NA” indicates that the fi rst year is not known (used mostly for “reported” taxa).

Appendices 19

Comments (Appendices A, B, C, and D)This column includes additional comments on the status of a particular species in the monu-ment, the place of origin for introduced species, legal status, and other data.

Nativity (Appendix D)Nativity indicates whether a species is native to Arizona or introduced (Intro) based on McDou-gall (1973) and Kartesz (2003).

Flower time (Appendix D)Flower Time refers to the months in which a spe-cies is most likely to be in fl ower. NA indicates a species that is a non-fl owering fern or gymno-sperm.

Habitat (Appendix D)The distribution of species across generalized habitats in Pipe Spring National Monument is in-dicated by an “x” in the columns DesSh through Wet and is derived from the vegetation classifi ca-tion used by Alexander (1998b). DesSh = Desert shrub vegetation of fl ats and foothills below the monument’s Navajo Sandstone mesa dominated by greenish rabbitbrush (Chrysothamnus nauseo-sus var. consimilis), fourwing saltbush (Atriplex canescens), and sand sagebrush (Artemisia fi lifo-lia). PinJun = Pinyon-Juniper woodlands/rim-rock dominated by Utah juniper (Juniperus os-teosperma), Two-needle pinyon (Pinus edulis), turbinella live oak (Quercus turbinella), and Utah serviceberry (Amelanchier utahensis). Wet = Wet meadows associated with natural seeps and springs. Additional comments are included for species that are restricted to a particular soil/geo-logic type or are found mostly in disturbed sites.


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Pres

Unc

Fert

ig 2

3683

(P

ISP)

2007

Nat

ive

to S

W

USA

Ana

card

iace

aeRh

us a

rom

atic

a va

r. si

mpl

icif

olia

R. t

rilo

bata

var

. si

mpl

icif

olia

squa

wbu

shSh

rub

Wid

ePr

esU

ncPh

illip

s 95

-101

(N

AU

)19

95

Ana

card

iace

aeRh

us a

rom

atic

a va

r. tr

iloba

taR.

tri

loba

ta v

ar.

trilo

bata

squa

wbu

shSh

rub

Wid

eRe

p?

Hea

ton

(193

5)19

35

Apo

cyna

ceae

Am

soni

a jo

nesi

iJo

nes’

am

soni

aPe

rFRe

gEn

His

t?

Fann

ing

s.n.

(D

eser

t Bo

t G

arde

n)

1961

Apo

cyna

ceae

Am

soni

a to

men

tosa

va

r. st

enop

hylla

A. e

astw

oodi

ana

tom

ento

se

amso

nia

PerF

Spar

seH

ist

?H

eato

n s.

n.

(PIS

P)19

36PI

SP s

peci

men

is

the

“bre

vifo

lia”

form

Apo

cyna

ceae

Ap

ocy

nu

m

cann

abin

umIn

clud

es v

ars.

ca

nnab

inum

, gl

aber

rim

um, &

an

gust

ifol

ium

com

mon

do

gban

ePe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)19

98

Asc

lepi

adac

eae

Asc

lepi

as la

tifo

liabr

oadl

eaf

milk

wee

dPe

rFW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Asc

lepi

adac

eae

Asc

lep

ias

subv

erti

cilla

taA

. gal

ioid

esw

horl

ed

milk

wee

dPe

rFW

ide

Pres

Unc

Ale

xand

er

1225

(PIS

P)19

36

Appendices 21

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Bora

gina

ceae

Cry

pta

nth

a ci

rcum

scis

sacu

shio

n cr

ypta

nth

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Bora

gina

ceae

Cry

pta

nth

a co

nfer

tifl

ora

basi

n ye

llow

cr

ypta

nth

PerF

Wid

ePr

esU

ncN

elso

n 93

11

(RM

)19

37

Bora

gina

ceae

Cry

pta

nth

a cr

assi

sepa

la v

ar.

elac

hant

ha

thic

k-se

pale

d cr

ypta

nth

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Bora

gina

ceae

Cryp

tant

ha fl

ava

Ore

ocar

ya fl

ava

plat

eau

yello

w

cryp

tant

hPe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)N

A

Bora

gina

ceae

Cry

pta

nth

a fu

lvoc

anes

cens

Incl

udes

var

s.

fulv

ocan

esce

ns

& e

chin

oide

s,

Ore

ocar

ya

fulv

ocan

esce

ns

plat

eau

cryp

tant

hPe

rFRe

gEn

His

t?

Hea

ton

24

(PIS

P)19

37Ty

pe lo

calit

y of

va

r. ec

hino

ides

is

Par

ia C

anyo

n,

Gar

fi el

d Co

., U

T (J

ones

529

7p

POM

)

Bora

gina

ceae

Cryp

tant

ha g

raci

lissl

ende

r cr

ypta

nth

Ann

FW

ide

His

t?

Hea

ton

14

(PIS

P)19

37

Bora

gina

ceae

Cryp

tant

ha m

icra

ntha

redr

oot

cryp

tant

hA

nnF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

36Ty

pe lo

calit

y is

“S

outh

ern

Uta

h”

(Par

ry 1

64 )

Bora

gina

ceae

Cryp

tant

ha p

tero

cary

a va

r. cy

clop

tera

win

gnut

cr

ypta

nth

Ann

FW

ide

His

t?

Hea

ton

13

(PIS

P)19

37

Bora

gina

ceae

Cryp

tant

ha p

tero

cary

a va

r. pt

eroc

arya

win

gnut

cr

ypta

nth

Ann

FW

ide

Pres

Unc

Even

den

&

Loui

e PI

SP 1

6 (P

ISP)

2001

Bora

gina

ceae

Hel

iotr

opiu

m

conv

olvu

lace

umEu

plo

ca

conv

olvu

lace

a ss

p.

calif

orni

ca

bind

wee

d he

liotr

ope

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Bora

gina

ceae

Lapp

ula

occi

dent

alis

va

r. cu

pula

taL.

tex

ana,

L.

mar

gina

ta, L

. re

dow

skii

var.

cupu

lata

cups

eed

stic

ksee

dA

nnF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

14

(PIS

P)

1941

Bora

gina

ceae

Plag

iobo

thry

s jo

nesi

iJo

nes’

pop

corn

-fl

ower

Ann

FRe

gEn

Pres

Unc

Even

den

&

Loui

e PI

SP 1

5 (P

ISP)

2001

PISP

spe

cim

en

has

unus

ually

la

rge

fl ow

ers

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Cact

acea

eEc

hino

cere

us

enge

lman

nii v

ar.

vari

egat

us

E. e

ngel

man

nii v

ar.

decu

mbe

ns. V

ars.

not

re

cogn

ized

in F

lora

of

Nor

th A

mer

ica

(200

3)

Gle

n Ca

nyon

he

dgeh

og

cact

us

PerF

RegE

nRe

p?

Ale

xand

er

(199

8b)

1998

Cact

acea

eEc

hino

cere

us

trig

loch

idia

tus

var.

mel

anac

anth

us

Incl

udes

var

. ine

rmis

. V

ars.

not

rec

ogni

zed

in F

lora

of

Nor

th

Am

eric

a (2

003)

clar

etcu

pPe

rFW

ide

Pres

Unc

Hea

ton

s.n.

(P

ISP)

1936

Cact

acea

eEc

hino

cere

us

trig

loch

idia

tus

var.

moj

aven

sis

Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

3)

Moh

ave

clar

etcu

pPe

rFW

ide

His

t?

Peeb

les

1469

6 (A

RIZ)

1940

Cact

acea

eO

punt

ia e

rina

cea

var.

aure

aO

. aur

ea, O

. bas

ilari

s va

r. au

rea

Pipe

Spr

ing

pric

klyp

ear

PerF

LocE

nPr

esCo

mM

acLe

od 4

22

(ASU

)19

30Ty

pe lo

calit

y is

1/

2 m

ile N

of

Pipe

Spr

ing,

M

ohav

e Co

., A

Z (M

cCab

e &

M

cCab

e s.

n. D

S)

Cact

acea

eO

pu

nti

a er

inac

ea v

ar.

utah

ensi

sV

ar. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

3);

cons

ider

ed a

few

-sp

ined

var

iant

of

O.

poly

acan

tha

Uta

h pr

ickl

ypea

rPe

rFW

ide

Pres

Com

Mac

Leod

428

(A

SU)

1971

Cact

acea

eO

punt

ia f

ragi

lisO

. fra

gilis

var

. br

achy

arth

abr

ittl

e pr

ickl

ypea

rPe

rFW

ide

Rep

?K

ing

(197

7),

Ale

xand

er

(199

8b)

1977

Cact

acea

eO

punt

ia p

haea

cant

ha

var.

maj

orV

ars.

not

rec

ogni

zed

in F

lora

of

Nor

th

Am

eric

a (2

003)

larg

e pr

ickl

ypea

rPe

rFW

ide

Pres

Unc

Gan

z 17

7 (A

SU)

1965

Als

o ci

ted

by

Ale

xand

er

(199

8b)

Cact

acea

eO

punt

ia p

inka

vae

O. b

asila

ris

var.

woo

dbur

yi,

cons

ider

ed p

art

of O

. m

acro

rhiz

a in

Uta

h

Flor

a (2

003)

Pink

ava’

s pr

ickl

ypea

rPe

rFLo

cEn

Pres

Unc

Leht

o s.

n.

(ASU

)19

40Ty

pe lo

calit

y is

N

W o

f Bu

lrus

h Ca

nyon

S o

f Pi

pe

Spri

ng, M

ohav

e Co

., A

Z (P

arfi

t 28

74 A

SU)

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 23

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Cact

acea

eO

punt

ia w

hipp

lei v

ar.

whi

pple

iCy

lindr

opun

tia

whi

pple

iW

hipp

le’s

chol

laSh

rub

RegE

nPr

esU

ncK

ing

(197

7),

Ale

xand

er

(199

8b)

1977

Phot

ogra

phed

(b

ut n

ot

vouc

here

d) b

y W

. Fe

rtig

, 200

7

Cact

acea

eSc

lero

cact

us w

hipp

lei

var.

rose

usS.

par

vifl

oru

s,

S. w

hipp

lei v

ar.

inte

rmed

ius.

Incl

udes

S.

con

tort

us &

S.

terr

ae-c

anyo

nae

smal

lfl ow

er

fi sh

hook

cac

tus

PerF

Peri

phPr

esU

ncEv

ende

n &

Lo

uie

PISP

26

(PIS

P)

1998

Type

of

S.

inte

rmed

ius

is 9

m

i SW

of

Pipe

Sp

ring

, Moh

ave

Co.,

AZ

(Pee

bles

&

Par

ker

1471

2 CA

S)

Capp

arac

eae

(Cle

omac

eae)

Cleo

me

lute

aye

llow

bee

plan

tA

nnF

Wid

ePr

esU

ncA

lexa

nder

11

87 (P

ISP)

1935

Chen

opod

iace

aeA

trip

lex

arge

ntea

var

. ar

gent

eaA

. arg

ente

asi

lver

ora

chA

nnF

Wid

ePr

esU

ncA

lexa

nder

12

36 (P

ISP)

2001

Chen

opod

iace

aeA

trip

lex

cane

scen

s va

r. ca

nesc

ens

A. c

anes

cens

var

. oc

cide

ntal

isfo

urw

ing

salt

bush

Shru

bW

ide

Pres

Com

Ale

xand

er

1212

(PIS

P)19

66

Chen

opod

iace

aeCh

enop

odiu

m a

lbum

va

r. al

bum

C. a

lbum

lam

bsqu

arte

rA

nnF

Intr

oPr

esU

ncA

lexa

nder

12

61 (P

ISP)

1998

Nat

ive

to E

uras

ia

Chen

opod

iace

aeC

hen

op

od

ium

fr

emon

tii v

ar.

frem

onti

i

C. f

rem

onti

iFr

emon

t’s

goos

efoo

tA

nnF

Wid

ePr

esU

ncA

lexa

nder

11

92 (P

ISP)

2001

Chen

opod

iace

aeC

hen

op

od

ium

le

ptop

hyllu

mna

rrow

leaf

go

osef

oot

Ann

FW

ide

Rep

?H

eato

n (1

935)

1935

Chen

opod

iace

aeK

rasc

heni

nnik

ovia

la

nata

var

. sub

spin

osa

Euro

tia

lana

ta,

Cera

toid

es la

nata

var

. su

bspi

nosa

. Var

s. n

ot

reco

gniz

ed in

Flo

ra

of N

orth

Am

eric

a (2

003)

.

shru

bby

win

terf

atSh

rub

Wid

eH

ist

?Pe

eble

s 13

079

(ARI

Z)19

36Ty

pe lo

calit

y is

ne

ar S

t. G

eorg

e,

Was

hing

ton

Co.,

UT

(Goo

ddin

g 81

0 N

Y)

Chen

opod

iace

aeSa

lsol

a tr

agus

S. k

ali,

S. ib

eric

a, S

. pe

stif

er, S

. aus

tral

isRu

ssia

n th

istl

eA

nnF

Intr

oPr

esU

ncFe

rtig

236

76

(PIS

P)19

35N

ativ

e to

Asi

a

Chen

opod

iace

ae

(Sar

coba

tace

ae)

Sarc

ob

atu

s ve

rmic

ulat

usgr

ease

woo

dSh

rub

Wid

ePr

esU

ncA

lexa

nder

12

26 (P

ISP)

1935

Chen

opod

iace

aeSu

aeda

tor

reya

na v

ar.

torr

eyan

aS.

moq

uini

i, S.

nig

ra,

S. f

ruti

cosa

Torr

ey’s

seep

wee

dA

nnF

Wid

ePr

esU

ncA

lexa

nder

12

52 (P

ISP)

2001

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Com

posi

tae

(Ast

erac

eae)

Am

bro

sia

acan

thic

arpa

Fran

seri

a ac

anth

icar

pabu

r ra

gwee

dA

nnF

Wid

ePr

esU

ncA

lexa

nder

11

85 (P

ISP)

1935

Com

posi

tae

(Ast

erac

eae)

Am

bro

sia

arte

mis

iifol

iaA

. art

emis

iifol

ia v

ar.

elat

ior

com

mon

ra

gwee

dA

nnF

Wid

ePr

esU

ncA

lexa

nder

11

86 (P

ISP)

1977

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia b

igel

ovii

Bige

low

’s sa

gebr

ush

Shru

bW

ide

Pres

Unc

Ale

xand

er

(199

8b)

1977

Obs

erve

d in

fi el

d by

J. C

oles

and

W

. Fer

tig,

200

7

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia c

ampe

stri

s va

r. sc

oule

rian

aA

. cam

pest

ris

ssp.

pac

ifi ca

, A.

paci

fi ca

, Olig

ospo

rus

cam

pest

ris

ssp.

pa

cifi

cus,

O. p

acifi

cus

sand

w

orm

woo

dPe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)19

98O

bser

ved

in fi

eld

by J

. Col

es, 2

007

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia d

racu

ncul

us

ssp.

gla

uca

Olig

osp

oru

s d

racu

ncu

lus

ssp.

gla

ucus

, A.

drac

uncu

loid

es. V

ar.

not

reco

gniz

ed in

Cr

onqu

ist

(199

4)

and

Flor

a of

Nor

th

Am

eric

a (2

006)

tarr

agon

PerF

Wid

eRe

p?

Kin

g (1

977)

1977

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia fi

lifol

iaO

ligos

poru

s fi

lifol

ius

sand

sag

ebru

sh

Shru

bW

ide

Pres

Unc

Ale

xand

er

1257

(PIS

P)19

77

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia lu

dovi

cian

a va

r. al

bula

A. l

udov

icia

na s

sp.

albu

laLo

uisi

ana

wor

mw

ood

PerF

Wid

ePr

esU

ncFe

rtig

236

81

(PIS

P)19

77

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia t

ride

ntat

a va

r. tr

iden

tata

Seri

ph

idiu

m

trid

enta

tum

ssp

. tr

iden

tatu

m

basi

n bi

g sa

gebr

ush

Shru

bW

ide

Pres

Unc

Fert

ig 2

3674

(P

ISP)

2007

Com

posi

tae

(Ast

erac

eae)

Ast

er h

espe

rius

A. c

oeru

lesc

ens,

A

. lan

ceol

atus

va

r. h

esp

eriu

s,

Sym

ph

yotr

ich

um

la

nceo

latu

m v

ar.

hesp

eriu

m

Sisk

iyou

ast

erPe

rFW

ide

Pres

Rare

Ale

xand

er

1240

(PIS

P)20

01

Com

posi

tae

(Ast

erac

eae)

Ch

aen

acti

s ca

rpho

clin

iaC.

car

phoc

linia

var

. ca

rpho

clin

iaG

ray’

s ch

aena

ctis

Ann

FW

ide

Pres

Unc

Loui

e &

Ev

ende

n PI

SP-

10 (P

ISP)

1937

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 25

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Com

posi

tae

(Ast

erac

eae)

Ch

aeto

pap

pa

eric

oide

sLe

ucel

ene

eric

oide

s,

Ast

er a

reno

sus

rose

-hea

thPe

rFW

ide

Pres

Unc

Ale

xand

er

1207

(PIS

P)19

39

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

cons

imili

s

Eric

amer

ia n

ause

osa

var.

cons

imili

s,

E. n

ause

osa

var.

oreo

phila

, C.

naus

eosu

s va

r. or

eoph

ilus

gree

nish

ra

bbit

brus

hSh

rub

Wid

ePr

esCo

mA

lexa

nder

12

19 (P

ISP)

1998

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

gnap

halo

des

C. n

ause

osus

va

r. ho

lole

ucus

, Er

icam

eria

nau

seos

a va

r. ho

lole

uca

gray

stem

ra

bbit

brus

hSh

rub

Wid

ePr

esU

ncFe

rtig

236

65

(PIS

P)20

07

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

grav

eole

ns

Eric

amer

ia n

ause

osa

var.

grav

eole

ns,

E. n

ause

osa

var.

glab

rata

glab

rate

ra

bbit

brus

hSh

rub

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

junc

eus

Eric

amer

ia n

ause

osa

var.

junc

earu

sh

rabb

itbr

ush

Shru

bRe

gEn

Pres

Unc

Fert

ig 2

3678

(P

ISP)

2007

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s vi

scid

ifl or

us v

ar.

pube

rulu

s

Chry

soth

amnu

s vi

scid

ifl or

us s

sp.

pube

rulu

s

pube

rule

nt

rabb

itbr

ush

Shru

bW

ide

Pres

Unc

Ale

xand

er

1193

(PIS

P)20

01

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s vi

scid

ifl or

us v

ar.

sten

ophy

llus

Incl

uded

in v

ar.

visc

idifl

orus

by

som

e au

thor

s. V

ar. n

ot

reco

gniz

ed in

Flo

ra

of N

orth

Am

eric

a (2

006)

.

slen

derl

eaf

rabb

itbr

ush

Shru

bW

ide

Pres

Com

Ale

xand

er

1247

(PIS

P)19

98

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s vi

scid

ifl or

us v

ar.

visc

idifl

orus

gree

n ra

bbit

brus

hSh

rub

Wid

eRe

p?

Kin

g (1

977)

1977

Com

posi

tae

(Ast

erac

eae)

Cony

za c

anad

ensi

s va

r. gl

abra

taEr

iger

on c

anad

ensi

s,

Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

6)

Cana

dian

ho

rsew

eed

Ann

FW

ide

Rep

?H

eato

n (1

935)

1935

Com

posi

tae

(Ast

erac

eae)

Erig

eron

div

erge

ns

var.

dive

rgen

sE.

div

erge

nssp

read

ing

dais

yPe

rFW

ide

Pres

Unc

Hea

ton

s.n.

(P

ISP)

1936

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Com

posi

tae

(Ast

erac

eae)

Erig

eron

pum

ilus

var.

conc

innu

sE.

con

cinn

us v

ar.

conc

innu

s. In

clud

es

var.

subg

labe

r

Nav

ajo

fl ea

bane

PerF

Peri

phRe

p?

Ale

xand

er

(199

8b)

1998

Com

posi

tae

(Ast

erac

eae)

Gai

llard

ia a

rizo

nica

Ari

zona

bl

anke

tfl o

wer

Ann

FRe

gEn

Rep

?A

lexa

nder

(1

998b

)19

98

Com

posi

tae

(Ast

erac

eae)

Gai

llard

ia p

inna

tifi

daH

opi

blan

ketfl

ow

erPe

rFW

ide

Pres

Unc

Ale

xand

er

1208

(PIS

P)20

01

Com

posi

tae

(Ast

erac

eae)

Gri

ndel

ia s

quar

rosa

va

r. se

rrul

ata

Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

6)

curl

ycup

gu

mw

eed

PerF

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Com

posi

tae

(Ast

erac

eae)

Gu

tier

rezi

a m

icro

ceph

ala

Incl

udes

G. l

ucid

ath

read

sn

akew

eed

Shru

bW

ide

Pres

Unc

Ale

xand

er

1198

(PIS

P)19

35

Com

posi

tae

(Ast

erac

eae)

Gut

ierr

ezia

sar

othr

aebr

oom

sn

akew

eed

Shru

bW

ide

Pres

Com

Ale

xand

er

1258

(PIS

P)20

01

Com

posi

tae

(Ast

erac

eae)

Hel

iant

hus

annu

us

ssp.

lent

icul

aris

H. a

ridu

s; s

sp. n

ot

reco

gniz

ed in

Flo

ra

of N

orth

Am

eric

a (2

006)

com

mon

su

nfl o

wer

Ann

FW

ide

Pres

Unc

Ale

xand

er

1197

(PIS

P)19

35

Com

posi

tae

(Ast

erac

eae)

Hym

enop

appu

s fi

lifol

ius

var.

cine

reus

H. l

ugen

s, H

. fi

lifol

ius

var.

luge

ns, H

. fi li

foliu

s va

r. pa

ucifl

oru

s,

H. fi

lifo

lius

var.

meg

acep

halu

s,

H. fi

lifo

lius

var.

tom

ento

sus

com

mon

hy

alin

eher

bPe

rFW

ide

Pres

Unc

Ale

xand

er

1202

(PIS

P)19

98

Com

posi

tae

(Ast

erac

eae)

Iva

axill

aris

I.

axill

aris

var

. ax

illar

is. V

ars.

not

re

cogn

ized

in F

lora

of

Nor

th A

mer

ica

(200

6).

pove

rtyw

eed

PerF

Wid

ePr

esU

ncA

lexa

nder

12

17 (P

ISP)

1998

Com

posi

tae

(Ast

erac

eae)

Lact

uca

serr

iola

pric

kly

lett

uce

Ann

FIn

tro

Pres

Unc

Fert

ig 2

3690

(P

ISP)

1935

Nat

ive

to E

urop

e

Com

posi

tae

(Ast

erac

eae)

Layi

a gl

andu

losa

tidy

tips

Ann

FW

ide

Pres

Unc

Loui

e &

Ev

ende

n PI

SP 8

(P

ISP)

2001

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 27

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Com

posi

tae

(Ast

erac

eae)

Mac

haer

anth

era

cane

scen

s va

r. ca

nesc

ens

Ast

er c

anes

cens

, D

iete

ria

cane

scen

s va

r. ca

nesc

ens

hoar

y as

ter

PerF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

31

(PIS

P)

1936

Com

posi

tae

(Ast

erac

eae)

Mac

haer

anth

era

tana

ceti

folia

Ast

er t

anac

etif

oliu

sta

nsyl

eaf

aste

rA

nnF

Wid

ePr

esU

ncA

lexa

nder

12

15 (P

ISP)

1998

Com

posi

tae

(Ast

erac

eae)

Mal

acot

hrix

gla

brat

aM

. cal

ifor

nica

var

. gl

abra

tafi

lifor

m d

eser

t-da

ndel

ion

Ann

FW

ide

Pres

Unc

Even

den

&

Loui

e PI

SP 2

7 (P

ISP)

2001

Com

posi

tae

(Ast

erac

eae)

Mal

aco

thri

x so

ncho

ides

sow

this

tle

dese

rt-

dand

elio

n

Ann

FW

ide

His

t?

Hea

ton

3 (P

ISP)

1937

Com

posi

tae

(Ast

erac

eae)

On

op

ord

um

ac

anth

ium

O. a

cant

hium

var

. ac

anth

ium

Scot

ch t

hist

lePe

rFIn

tro

Rep

?A

lexa

nder

(1

998b

)19

98A

Z st

ate

proh

ibit

ed

and

rest

rict

ed

noxi

ous

wee

d.

Nat

ive

to E

uras

ia

Com

posi

tae

(Ast

erac

eae)

Psilo

stro

phe

spar

sifl

ora

gree

nste

m

pape

rfl o

wer

PerF

RegE

nH

ist

?H

eato

n s.

n.

(PIS

P)19

36Ty

pe lo

calit

y is

“S

outh

ern

Uta

h”

(Tho

mps

on s

.n.

GH

)

Com

posi

tae

(Ast

erac

eae)

Sene

cio

doug

lasi

i var

. lo

ngilo

bus

S. lo

ngilo

bus,

S.

fl ac

cidu

s va

r. fl

acci

dus,

S. fl

acc

idu

s va

r. do

ugla

sii

Dou

glas

’ gr

ound

sel

PerF

Wid

ePr

esU

ncW

hite

head

s.n

. (P

ISP)

1935

Com

posi

tae

(Ast

erac

eae)

Solid

ago

sp.

gold

enro

dPe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)19

98Id

enti

fi ca

tion

ne

eded

Com

posi

tae

(Ast

erac

eae)

Sonc

hus

aspe

rsp

inyl

eaf

sow

this

tle

Ann

FIn

tro

Pres

Unc

Even

den

&

Loui

e PI

SP 3

5 (P

ISP)

1997

Nat

ive

to E

urop

e

Com

posi

tae

(Ast

erac

eae)

Sonc

hus

oler

aceu

sco

mm

on

sow

this

tle

Ann

FIn

tro

Rep

?A

lexa

nder

(1

998b

)19

98N

ativ

e to

Eur

ope

Com

posi

tae

(Ast

erac

eae)

Step

hano

mer

ia e

xigu

aS.

exi

gua

ssp.

exi

gua

whi

te-p

lum

e w

ire

lett

uce

Ann

FW

ide

Pres

Unc

Ale

xand

er

1199

(PIS

P)20

01

Com

posi

tae

(Ast

erac

eae)

Step

hano

mer

ia

pauc

ifl or

abr

own-

plum

e w

ire

lett

uce

PerF

Wid

ePr

esU

ncFe

rtig

236

68

(PIS

P)19

98

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Com

posi

tae

(Ast

erac

eae)

Thel

esp

erm

a su

bnud

um v

ar.

subn

udum

Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

6)

scap

ose

gree

nthr

ead

PerF

Wid

ePr

esU

ncA

lexa

nder

12

04 (P

ISP)

1937

Type

loca

lity

is

“Sou

ther

n U

tah”

(P

arry

109

GH

)

Com

posi

tae

(Ast

erac

eae)

Tow

nsen

dia

inca

naho

ary

tow

nsen

dia

PerF

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

19

(PIS

P)

2001

Com

posi

tae

(Ast

erac

eae)

Trag

opog

on d

ubiu

sT.

dub

ius

var.

maj

orye

llow

sal

sify

PerF

Intr

oPr

esU

ncEv

ende

n &

Lo

uie

PISP

28

(PIS

P)

1997

Nat

ive

to E

urop

e

Com

posi

tae

(Ast

erac

eae)

Ver

besi

na e

ncel

ioid

es

var.

exau

ricu

lata

Xim

enes

ia

ence

lioid

esgo

lden

cr

ownb

eard

Ann

FW

ide

Pres

Unc

Ale

xand

er

1188

(PIS

P)19

35

Com

posi

tae

(Ast

erac

eae)

Xan

thiu

m s

trum

ariu

m

var.

cana

dens

eX

. pen

sylv

anic

um,

X. s

acch

arat

um.

Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

6).

roug

h co

ckle

bur

Ann

FW

ide

Rep

?H

eato

n (1

935)

&

Kin

g (1

977)

1935

Conv

olvu

lace

aeCo

nvol

vulu

s ar

vens

isfi

eld

bind

wee

dPe

rFIn

tro

His

t?

Park

er 6

238

(ARI

Z)19

46A

Z st

ate

proh

ibit

ed

and

regu

late

d no

xiou

s w

eed.

N

ativ

e to

Eur

asia

Cruc

ifer

ae

(Bra

ssic

acea

e)A

rabi

s pe

renn

ans

var.

pere

nnan

sBo

eche

ra p

eren

nans

pere

nnia

l ro

ckcr

ess

PerF

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Cruc

ifer

ae

(Bra

ssic

acea

e)Ca

psel

la b

ursa

-pas

tori

ssh

ephe

rd’s-

purs

eA

nnF

Intr

oH

ist

?H

eato

n 17

(P

ISP)

1937

Nat

ive

to E

urop

e

Cruc

ifer

ae

(Bra

ssic

acea

e)D

escu

rain

ia p

inna

ta

var.

glab

raw

este

rn

tans

ymus

tard

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1935

Cruc

ifer

ae

(Bra

ssic

acea

e)D

escu

rain

ia p

inna

ta

var.

inte

rmed

iaIn

clud

ed in

D.

pinn

ata

var.

osm

iaru

m b

y so

me

auth

ors

wes

tern

ta

nsym

usta

rdA

nnF

Peri

phH

ist

?H

eato

n 19

41-1

(P

ISP)

1939

Cruc

ifer

ae

(Bra

ssic

acea

e)D

escu

rain

ia p

inna

ta

var.

osm

iaru

mD

. pin

nata

ssp

. ha

licto

rum

wes

tern

ta

nsym

usta

rdA

nnF

Wid

ePr

esU

ncH

eato

n 19

41-2

(P

ISP)

1941

Cruc

ifer

ae

(Bra

ssic

acea

e)D

escu

rain

ia s

ophi

afl

ixw

eed

Ann

FIn

tro

Pres

Unc

Nel

son

9304

(R

M)

1971

Nat

ive

to E

urop

e

Cruc

ifer

ae

(Bra

ssic

acea

e)D

ithy

rea

wis

lizen

iD

imor

phoc

arpa

w

isliz

eni

spec

tacl

epod

Ann

FW

ide

Pres

Unc

Hea

ton

s.n.

(P

ISP)

1936

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 29

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Cruc

ifer

ae

(Bra

ssic

acea

e)Er

ysim

um r

epan

dum

spre

adin

g w

allfl

ower

Ann

FIn

tro

His

t?

Hea

ton

s.n.

(P

ISP)

1939

Nat

ive

to E

urop

e

Cruc

ifer

ae

(Bra

ssic

acea

e)Le

pidi

um d

ensi

fl o

rum

va

r. ra

mos

umIn

clud

es v

ar.

mac

roca

rpum

prai

rie

pepp

erw

ort

Ann

FPe

riph

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Cruc

ifer

ae

(Bra

ssic

acea

e)Le

pidi

um m

onta

num

va

r. jo

nesi

iJo

nes’

pe

pper

wor

tPe

rFW

ide

Pres

Com

Hea

ton

s.n.

(P

ISP)

1937

Type

loca

lity

is S

t. G

eorg

e,

Was

hing

ton

Co.,

UT

(Jon

es 1

636

NY

)

Cruc

ifer

ae

(Bra

ssic

acea

e)Le

pidi

um m

onta

num

va

r. st

ella

eL.

mon

tanu

m v

ar.

cine

reum

Stel

la’s

pepp

erw

ort

PerF

LocE

nH

ist

?H

eato

n 19

41-

17 (P

ISP)

1941

Type

loca

lity

is S

E of

Can

nonv

ille,

K

ane

Co.,

UT

(Rev

eal &

Rev

eal

4454

BRY

)

Cruc

ifer

ae

(Bra

ssic

acea

e)Le

pidi

um p

erfo

liatu

mcl

aspi

ng

pepp

erw

ort

Ann

FIn

tro

His

t?

Leht

o 71

72

(ARI

Z)19

66N

ativ

e to

Eur

ope

Cruc

ifer

ae

(Bra

ssic

acea

e)N

astu

rtiu

m o

ffi c

inal

eRo

ripp

a na

stur

tium

-aq

uati

cum

wat

ercr

ess

PerF

Intr

oPr

esU

ncFe

rtig

236

92

(PIS

P)19

97N

ativ

e to

Eur

ope

Cruc

ifer

ae

(Bra

ssic

acea

e)Si

sym

briu

m a

ltis

sim

umtu

mbl

e m

usta

rdA

nnF

Intr

oPr

esU

ncH

eato

n 19

41-

11 (P

ISP)

1941

Nat

ive

to E

urop

e

Cruc

ifer

ae

(Bra

ssic

acea

e)St

anle

ya p

inna

taS.

pin

nata

var

. pi

nnat

apr

ince

’s-pl

ume

PerF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

3

(PIS

P)

1939

Cruc

ifer

ae

(Bra

ssic

acea

e)St

rept

anth

ella

lo

ngir

ostr

islo

ng-b

eak

fi dd

le m

usta

rdA

nnF

Wid

eH

ist

?H

eato

n 18

(P

ISP)

1937

Cucu

rbit

acea

eCu

curb

ita

foet

idis

sim

abu

ffal

o go

urd

PerF

Wid

ePr

esU

ncA

lexa

nder

12

18 (P

ISP)

1935

Cupr

essa

ceae

Jun

iper

us

oste

ospe

rma

Sabi

na o

steo

sper

ma

Uta

h ju

nipe

rSh

rub

Wid

ePr

esU

ncA

lexa

nder

12

46 (P

ISP)

1935

Cupr

essa

ceae

Juni

peru

s sc

opul

orum

Sabi

na s

copu

loru

mRo

cky

Mou

ntai

n ju

nipe

r

Tree

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Cype

race

aeCa

rex

nebr

asce

nsis

C. n

ebra

sken

sis

Neb

rask

a se

dge

PerG

Wid

ePr

esRa

reA

lexa

nder

12

44 (P

ISP)

1998

Cype

race

aeCa

rex

prae

grac

ilis

blac

kcre

eper

se

dge

PerG

Wid

ePr

esRa

reH

eato

n s.

n.

(PIS

P)19

38

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Cype

race

aeCy

peru

s sp

.fl

at-s

edge

Ann

FIn

tro

Rep

?A

lexa

nder

(1

998b

)19

98Id

enti

fi ca

tion

ne

eded

Cype

race

aeEl

eoch

aris

par

ishi

iPa

rish

’s sp

iker

ush

PerG

Wid

ePr

esRa

reA

lexa

nder

12

42 (P

ISP)

1998

Cype

race

aeSc

irpu

s pu

ngen

s va

r. lo

ngis

pica

tus

Scri

pus

pung

ens

var.

poly

phyl

lus,

Sc

irpu

s am

eric

anus

va

r. lo

ngis

pica

tus,

Sc

hoen

ople

ctus

pu

ngen

s va

r. lo

ngis

pica

tus,

Sc

hoen

ople

ctus

pu

ngen

s va

r. po

lyph

yllu

s

thre

e-sq

uare

bu

lrus

hPe

rGW

ide

Pres

Rare

Ale

xand

er

1243

(PIS

P)19

98

Elae

agna

ceae

Shep

her

dia

ro

tund

ifol

iaro

undl

eaf

buff

alob

erry

Shru

bRe

gEn

Pres

Unc

Fert

ig 2

3667

(P

ISP)

1977

Type

loca

lity

is

uppe

r va

lley

of

the

Vir

gin,

Kan

e Co

., U

T (S

iler

s.n.

IS

C)

Ephe

drac

eae

Ephe

dra

torr

eyan

aTo

rrey

’s ep

hedr

aSh

rub

Wid

ePr

esRa

reFe

rtig

236

69

(PIS

P)20

07

Ephe

drac

eae

Ephe

dra

viri

dis

var.

viri

dis

E. v

irid

isgr

een

ephe

dra

Shru

bW

ide

Pres

Unc

Ale

xand

er

1201

(PIS

P)19

77

Ephe

drac

eae

Ephe

dra

viri

dis

var.

visc

ida

E. c

utle

riCu

tler

’s ep

hedr

aSh

rub

RegE

nPr

esU

ncG

anz

s.n.

(A

RIZ)

1965

Obs

erve

d ve

geta

tive

ly b

y Fe

rtig

, 200

7

Euph

orbi

acea

eC

ham

aesy

ce

albo

mar

gina

taEu

ph

orb

ia

albo

mar

gina

tara

ttle

snak

e-w

eed

PerF

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Euph

orbi

acea

eCh

amae

syce

fen

dler

iEu

phor

bia

fend

leri

Fend

ler’s

spu

rge

PerF

Wid

ePr

esU

ncA

lexa

nder

12

03 (P

ISP)

1939

Euph

orbi

acea

eC

ham

aesy

ce

glyp

tosp

erm

aEu

ph

orb

ia

glyp

tosp

erm

ari

dge-

seed

ed

spur

geA

nnF

Wid

ePr

esCo

mH

eato

n s.

n.

(PIS

P)19

36

Euph

orbi

acea

eCh

amae

syce

pro

stra

taEu

phor

bia

pros

trat

a,

E. c

ham

aesy

cepr

ostr

ate

spur

geA

nnF

Intr

oPr

esU

ncFe

rtig

236

91

(PIS

P)20

07N

ativ

e to

eas

tern

N

orth

Am

eric

a

Faga

ceae

Que

rcus

tur

bine

llatu

rbin

ella

live

-oa

kSh

rub

Wid

ePr

esU

ncA

lexa

nder

12

49 (P

ISP)

1935

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 31

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Fum

aria

ceae

Cory

dalis

aur

ea v

ar.

occi

dent

alis

C. c

urvi

siliq

ua s

sp.

occi

dent

alis

gold

en

cory

dalis

PerF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

36

Gen

tian

acea

eSw

erti

a al

bom

argi

nata

Fras

era

albo

mar

gina

taw

hite

-mar

gine

d sw

erti

aPe

rFW

ide

His

t?

Peeb

les

1308

0 (A

RIZ)

1936

Type

loca

lity

is

near

St.

Geo

rge,

W

ashi

ngto

n Co

., U

T (P

alm

er s

.n.

US)

Gen

tian

acea

eSw

erti

a ut

ahen

sis

Fras

era

pani

cula

taU

tah

swer

tia

PerF

RegE

nRe

p?

Ale

xand

er

(199

8b)

1998

Type

loca

lity

is

Buck

skin

Mtn

s,

Kai

bab

Plat

eau

“on

the

sout

hern

ed

ge o

f U

tah”

, K

ane

Co.,

UT

(Jon

es s

.n. P

OM

)

Ger

ania

ceae

Erod

ium

cic

utar

ium

stor

k’s-

bill

Ann

FIn

tro

Pres

Com

Ale

xand

er

1213

(PIS

P)19

97N

ativ

e to

Eur

ope

Gra

min

eae

(Poa

ceae

)A

grop

yron

cri

stat

umA

. cri

stat

um s

sp.

cris

tatu

m, A

. cr

ista

tum

ssp

. pe

ctin

atum

, A.

pect

inat

um, A

. si

biri

cum

, A. f

ragi

le,

A. m

ongo

licum

, A.

dese

rtor

um

cres

ted

whe

atgr

ass

PerG

Intr

oRe

p?

Kin

g (1

977)

1977

Nat

ive

to E

uras

ia

Gra

min

eae

(Poa

ceae

)A

gros

tis

stol

onif

era

A. a

lba,

A. g

igan

tea

redt

opPe

rGIn

tro

Pres

Unc

Ale

xand

er

1241

(PIS

P)19

98N

ativ

e to

Eur

asia

an

d A

fric

a

Gra

min

eae

(Poa

ceae

)A

rist

ida

purp

urea

A. g

lauc

a, A

. pu

rpur

ea v

ar.

fend

leri

ana,

A.

fend

leri

ana,

A.

purp

urea

var

. lo

ngis

eta

purp

le t

hree

-aw

nPe

rGW

ide

Pres

Com

Ale

xand

er

1210

(PIS

P)19

41

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

barb

ata

Chon

dros

um

barb

atum

. Inc

lude

s va

rs. b

arba

ta &

ro

thro

ckii

sixw

eeks

gra

ma

Ann

GW

ide

Pres

Unc

Fert

ig 2

3671

(P

ISP)

1935

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Gra

min

eae

(Poa

ceae

)B

ou

telo

ua

curt

ipen

dula

Incl

udes

var

s.

curt

ipen

dula

&

caes

pito

sa

side

oats

gra

ma

PerG

Wid

eRe

p?

Kin

g (1

977)

1977

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

erio

poda

Chon

dros

um

erio

podu

mbl

ack

gram

aPe

rGW

ide

Pres

Unc

Ale

xand

er

1250

(PIS

P)19

77

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

grac

ilis

Chon

dros

um g

raci

lebl

ue g

ram

aPe

rGW

ide

Pres

Unc

Purc

hase

272

(A

RIZ)

1936

Gra

min

eae

(Poa

ceae

)Br

omus

dia

ndru

sB.

rig

idus

, Ani

sant

ha

dian

dra

ripg

ut b

rom

eA

nnG

Intr

oPr

esU

ncLo

uie

&

Even

den

PISP

22

(PIS

P)

1977

Nat

ive

to E

uras

ia

Gra

min

eae

(Poa

ceae

)Br

omus

iner

mis

var

. in

erm

isBr

omop

sis

iner

mis

, Br

omus

iner

mis

smoo

th b

rom

ePe

rGIn

tro

Rep

?K

ing

(197

7)19

77N

ativ

e to

Eur

asia

Gra

min

eae

(Poa

ceae

)Br

omus

rub

ens

red

brom

eA

nnG

Intr

oPr

esCo

mLo

uie

&

Even

den

PISP

34

(PIS

P)

1998

Nat

ive

to E

uras

ia

Gra

min

eae

(Poa

ceae

)Br

omus

tec

toru

mA

nisa

ntha

tec

toru

mch

eatg

rass

Ann

GIn

tro

Pres

Com

Hea

ton

1941

-14

(PIS

P)19

38N

ativ

e to

Eur

asia

Gra

min

eae

(Poa

ceae

)Ce

nchr

us lo

ngis

pinu

sC.

pau

cifl

orus

fi el

d sa

ndbu

rA

nnG

Wid

eRe

p?

Kur

th (1

998)

1997

Gra

min

eae

(Poa

ceae

)D

isti

chlis

spi

cata

D

. spi

cata

var

. str

icta

, D

. str

icta

dese

rt s

altg

rass

PerG

Wid

eRe

p?

Kin

g (1

977)

, A

lexa

nder

(1

998b

)

1977

Gra

min

eae

(Poa

ceae

)Ec

hino

chlo

a cr

us-g

alli

Incl

udes

E.

mic

rost

achy

a,

E. m

uric

ata

var.

mic

rost

achy

a

barn

yard

-gra

ssA

nnG

Intr

oRe

p?

Ale

xand

er

(199

8b)

1998

Nat

ive

to E

uras

ia

Gra

min

eae

(Poa

ceae

)El

ymus

elo

ngat

us

E. e

long

atus

var

. po

ntic

us, A

grop

yron

el

onga

tum

, Th

inop

yrum

po

ntic

um

tall

whe

atgr

ass

PerG

Intr

oRe

p?

Kur

th (1

998)

1997

Nat

ive

to E

uras

ia

Gra

min

eae

(Poa

ceae

)El

ymus

ely

moi

des

Sita

nion

hys

trix

, E.

ely

moi

des

var.

elym

oide

s

squi

rrel

tail

PerG

Wid

eRe

p?

Kin

g (1

977)

, A

lexa

nder

(1

998b

)

1977

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 33

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Gra

min

eae

(Poa

ceae

)El

ymus

his

pidu

s A

grop

yron

in

term

ediu

m,

Elyt

rigi

a in

term

edia

, E.

his

pidu

s va

r. hi

spid

us, T

hino

pyru

m

inte

rmed

ium

inte

rmed

iate

w

heat

gras

sPe

rGIn

tro

Rep

?K

ing

(197

7)19

77N

ativ

e to

Eur

asia

Gra

min

eae

(Poa

ceae

)El

ymus

sm

ithi

iA

grop

yron

sm

ithi

i, Pa

scop

yrum

sm

ithi

i; in

clud

es v

ars.

sm

ith

ii &

pal

mer

i

wes

tern

w

heat

gras

sPe

rGW

ide

Rep

?K

ing

(197

7),

Ale

xand

er

(199

8b)

1977

Gra

min

eae

(Poa

ceae

)El

ymus

tra

chyc

aulu

s E.

tra

chyc

aulu

s va

r. tr

achy

caul

us,

E. a

lask

anus

ss

p. la

tigl

umis

, A

grop

yron

can

inum

, A

. sub

secu

ndum

, A

. tra

chyc

aulu

m, A

. la

tigl

ume

slen

der

whe

atgr

ass

PerG

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

38

Gra

min

eae

(Poa

ceae

)Er

agro

stis

cili

anen

sis

stin

kgra

ssA

nnG

Intr

oRe

p?

Kin

g (1

977)

1977

Nat

ive

to E

uras

ia

Gra

min

eae

(Poa

ceae

)Er

agro

stis

mex

ican

aE.

orc

utti

ana

Mex

ican

lo

vegr

ass

Ann

GIn

tro

Rep

?A

lexa

nder

(1

998b

)19

98N

ativ

e to

w

este

rn N

orth

A

mer

ica

Gra

min

eae

(Poa

ceae

)Er

agro

stis

pec

tina

cea

E. d

iffu

satu

fted

love

gras

sA

nnG

Wid

ePr

esU

ncFe

rtig

236

82

(PIS

P)19

77

Gra

min

eae

(Poa

ceae

)Er

ion

euro

n

pulc

hellu

mD

asyo

chlo

a pu

lche

lla,

Trid

ens

pulc

hellu

sfl

uffg

rass

PerG

Wid

eRe

p?

Hea

ton

(193

5)19

35

Gra

min

eae

(Poa

ceae

)Fe

stuc

a ar

undi

nace

aSc

hedo

noru

s p

ho

enix

, S.

arun

dina

ceus

, Lol

ium

ar

undi

nace

um

tall

fesc

uePe

rGIn

tro

Rep

?K

ing

(197

7),

Ale

xand

er

(199

8b)

1977

Nat

ive

to E

urop

e

Gra

min

eae

(Poa

ceae

)H

ilari

a ja

mes

iiPl

eura

phis

jam

esii

galle

taPe

rGW

ide

Pres

Unc

Ale

xand

er

1209

(PIS

P)19

36

Gra

min

eae

(Poa

ceae

)H

orde

um ju

batu

mCr

ites

ion

juba

tum

foxt

ail b

arle

yPe

rGW

ide

Rep

?K

ing

(197

7)19

77

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Gra

min

eae

(Poa

ceae

)H

orde

um m

urin

umH

. lep

orin

um,

H. m

urin

um s

sp.

lepo

rinu

m, C

rite

sion

gl

aucu

m

rabb

it b

arle

yA

nnG

Intr

oPr

esU

ncH

eato

n 19

41-3

(P

ISP)

1938

Gra

min

eae

(Poa

ceae

)M

uhle

nber

gia

aspe

rifo

liasc

ratc

hgra

ssPe

rGW

ide

Pres

Unc

Ale

xand

er

1223

(PIS

P)19

77

Gra

min

eae

(Poa

ceae

)M

uhle

nber

gia

port

eri

bush

muh

lyPe

rGW

ide

Pres

Unc

Fert

ig 2

3680

(P

ISP)

1977

Gra

min

eae

(Poa

ceae

)M

unro

a sq

uarr

osa

Mon

roa

squa

rros

afa

lse

buff

alog

rass

Ann

GW

ide

Pres

Rare

Fert

ig 2

3670

(P

ISP)

2007

Gra

min

eae

(Poa

ceae

)Po

a fe

ndle

rian

aP.

long

iligu

la, P

. fe

ndle

rian

a ss

p.

long

iligu

la

mut

tong

rass

PerG

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Gra

min

eae

(Poa

ceae

)Po

a pr

aten

sis

Poa

agas

size

nsis

Ken

tuck

y bl

uegr

ass

PerG

Intr

oRe

p?

Kin

g (1

977)

, A

lexa

nder

(1

998b

)

1977

Nat

ive

to E

urop

e

Gra

min

eae

(Poa

ceae

)Po

lyp

og

on

m

onsp

elie

nsis

rabb

itsf

oot-

gras

sA

nnG

Intr

oPr

esU

ncFe

rtig

236

86

(PIS

P)19

98N

ativ

e to

Eur

asia

an

d A

fric

a

Gra

min

eae

(Poa

ceae

)Po

lyp

og

on

se

miv

erti

cilla

tus

Ag

rost

is

sem

iver

tici

llata

, P.

viri

dis

wat

er

poly

pogo

nPe

rGIn

tro

Rep

?A

lexa

nder

(1

998b

)19

98N

ativ

e to

Eur

asia

an

d A

fric

a

Gra

min

eae

(Poa

ceae

)Sp

orob

olus

air

oide

s va

r. ai

roid

esS.

air

oide

sal

kali

saca

ton

PerG

Wid

eRe

p?

Kin

g (1

977)

, A

lexa

nder

(1

998b

)

1977

Gra

min

eae

(Poa

ceae

)Sp

orob

olus

con

trac

tus

spik

e dr

opse

edPe

rGW

ide

Pres

Unc

Ale

xand

er

1221

(PIS

P)19

36

Gra

min

eae

(Poa

ceae

)Sp

oro

bo

lus

cryp

tand

rus

sand

dro

psee

dPe

rGW

ide

Pres

Unc

Purc

hase

271

(A

RIZ)

1936

Gra

min

eae

(Poa

ceae

)Sp

orob

olus

fl ex

uosu

sm

esa

drop

seed

PerG

Wid

ePr

esU

ncA

lexa

nder

12

20 (P

ISP)

1977

Gra

min

eae

(Poa

ceae

)St

ipa

com

ata

var.

com

ata

Hes

pero

stip

a co

mat

a va

r. co

mat

ane

edle

-and

-th

read

PerG

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

28

(PIS

P)

1977

Gra

min

eae

(Poa

ceae

)St

ipa

hym

enoi

des

Ory

zops

is

hym

eno

ides

, A

chn

ath

eru

m

hym

enoi

des

Indi

an r

iceg

rass

PerG

Wid

ePr

esU

ncK

ing

(197

7)19

77O

bser

ved

vege

tati

vely

by

Fert

ig, 2

007

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 35

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Gra

min

eae

(Poa

ceae

)St

ipa

spec

iosa

Ach

nath

erum

sp

ecio

sum

, Jar

ava

spec

iosu

m

dese

rt

need

legr

ass

PerG

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

23

(PIS

P)

1977

Hyd

roph

ylla

ceae

Phac

elia

cre

nula

ta v

ar.

corr

ug

ata

P. c

orru

gata

corr

ugat

e ph

acel

iaA

nnF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

37

Hyd

roph

ylla

ceae

Phac

elia

cre

nula

ta v

ar.

cren

ulat

aP.

cre

nula

tacr

enul

ate

phac

elia

Ann

FW

ide

Pres

Unc

Loui

e &

Ev

ende

n PI

SP 6

(P

ISP)

2001

Hyd

roph

ylla

ceae

Phac

elia

ives

iana

Ives

’ pha

celia

Ann

FW

ide

Pres

Unc

Loui

e &

Ev

ende

n PI

SP

12 (P

ISP)

1937

Hyd

roph

ylla

ceae

Phac

elia

pul

chel

la v

ar.

good

ding

iiG

oodd

ing’

s pr

etty

pha

celia

Ann

FRe

gEn

His

t?

Hea

ton

7 (P

ISP)

1937

Irid

acea

eSi

syri

nchi

um

dem

issu

mbl

ue-e

yed

gras

sPe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)19

98

Junc

acea

eJu

ncus

arc

ticu

sJ.

bal

ticu

s va

r. m

onta

nus,

J.

arct

icus

var

. bal

ticu

s,

J. a

rcti

cus

ssp.

at

er; i

nclu

des

var.

mex

ican

us

Balt

ic r

ush

PerG

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

33 (P

ISP)

1998

Junc

acea

eJu

ncus

ens

ifol

ius

var.

brun

nesc

ens

J. s

axim

onta

nus

form

a br

unne

scen

s.

Incl

uded

in J

. en

sifo

lius

var.

mon

tanu

s by

som

e au

thor

s

Trac

y’s

rush

PerG

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

38

Junc

acea

eJu

ncus

tor

reyi

Torr

ey’s

rush

PerG

Wid

ePr

esU

ncFe

rtig

236

93

(PIS

P)19

98

Labi

atae

(L

amia

ceae

)M

arru

bium

vul

gare

com

mon

ho

reho

und

PerF

Intr

oPr

esU

ncA

lexa

nder

12

35 (P

ISP)

1935

Nat

ive

to E

uras

ia

Labi

atae

(L

amia

ceae

)M

enth

a sp

icat

asp

earm

int

PerF

Intr

oPr

esRa

reA

lexa

nder

11

90 (P

ISP)

1998

Nat

ive

to E

urop

e

Labi

atae

(L

amia

ceae

)Sa

lvia

dor

rii s

sp. d

orri

iS.

car

nosa

Dor

r’s s

age

Shru

bW

ide

Pres

Unc

Hea

ton

1 (P

ISP)

1937

Legu

min

osae

(F

abac

eae)

Ast

raga

lus

amph

ioxy

s va

r. ve

sper

tinu

sSh

eldo

n’s

milk

vetc

hPe

rFRe

gEn

His

t?

Hea

ton

s.n.

(P

ISP)

1934

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Legu

min

osae

(F

abac

eae)

Ast

raga

lus

cera

mic

us

var.

cera

mic

uspa

inte

d m

ilkve

tch

PerF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

21

(PIS

P)

1998

Legu

min

osae

(F

abac

eae)

Ast

raga

lus

prae

long

us

var.

prae

long

usst

inki

ng

milk

vetc

hPe

rFW

ide

Rep

?A

lexa

nder

&

Dec

ker

1444

(P

ISP)

1998

Legu

min

osae

(F

abac

eae)

Gly

cyrr

hiza

lepi

dota

G

. lep

idot

a va

r. le

pido

taN

utta

ll’s

licor

ice

PerF

Wid

eRe

p?

Ale

xand

er

(199

8b)

1998

Legu

min

osae

(F

abac

eae)

Lupi

nus

pusi

llus

var.

pusi

llus

L. p

usill

us s

sp.

pusi

llus

dwar

f lu

pine

Ann

FW

ide

His

t?

Hea

ton

2 (P

ISP)

1937

Legu

min

osae

(F

abac

eae)

Med

icag

o sa

tiva

M. s

ativ

a ss

p. s

ativ

aal

falf

aPe

rFIn

tro

Rep

?K

urth

(199

8)19

97N

ativ

e to

Eur

ope

Legu

min

osae

(F

abac

eae)

Mel

ilotu

s al

baM

. alb

us, i

nclu

ded

in M

. offi

cin

alis

by

som

e au

thor

s

whi

te

swee

tclo

ver

Ann

FIn

tro

Rep

?K

ing

(197

7),

Ale

xand

er

(199

8b)

1977

Nat

ive

to E

urop

e

Legu

min

osae

(F

abac

eae)

Mel

ilotu

s of

fi ci

nalis

yello

w

swee

tclo

ver

PerF

Intr

oRe

p?

Ale

xand

er

(199

8b)

1998

Nat

ive

to E

urop

e

Legu

min

osae

(F

abac

eae)

Pete

ria

thom

pson

iae

Thom

pson

’s pe

teri

aPe

rFRe

gEn

Pres

Rare

Ale

xand

er

1251

(PIS

P)20

01Ty

pe lo

calit

y is

K

anab

, Kan

e Co

., U

T (T

hom

pson

s.

n. G

H)

Legu

min

osae

(F

abac

eae)

Robi

nia

pseu

doac

acia

blac

k lo

cust

Tree

Intr

oPr

esU

ncA

lexa

nder

12

56 (P

ISP)

1998

Nat

ive

to e

aste

rn

Nor

th A

mer

ica

Lilia

ceae

(T

hem

idac

eae)

And

rost

ephi

um

brev

ifl or

umBr

odia

ea p

ayso

nii

pink

fun

nel-l

ilyPe

rFW

ide

His

t?

Peeb

les

1307

8 (U

niv

Ari

zona

)19

36Ty

pe lo

calit

y is

nea

r K

anab

, K

ane

Co.,

UT

(Tho

mps

on s

.n.)

Lilia

ceae

(C

aloc

hort

acea

e)Ca

loch

ortu

s fl

exuo

sus

sinu

ous

mar

ipos

aPe

rFW

ide

Pres

Unc

Loui

e &

Ev

ende

n PI

SP 1

(P

ISP)

1935

Type

loca

lity

is n

ear

Kan

ab,

Kan

e Co

., U

T (T

hom

pson

s.n

. G

H)

Lina

ceae

Linu

m a

rist

atum

Mes

yniu

m a

rist

atum

broo

mfl

axA

nnF

Wid

ePr

esU

ncFe

rtig

236

75

(PIS

P)19

98

Loas

acea

eM

entz

elia

alb

icau

lisM

. obs

cura

, Acr

olas

ia

albi

caul

us, A

. gra

cilis

whi

te-s

tem

bl

azin

gsta

rA

nnF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

17

(PIS

P)

1937

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 37

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Loas

acea

eM

entz

elia

inte

gra

M. m

ulti

fl or

a va

r. in

tegr

a, M

. pum

ila

var.

inte

gra

Vir

gin

stic

klea

fPe

rFW

ide

His

t?

Leht

o 70

37

(ARI

Z)19

66Ty

pe lo

calit

y is

Roc

kvill

e,

Was

hing

ton

Co.,

UT

(Jon

es 6

082c

)

Loas

acea

eM

entz

elia

mul

tifl

ora

M. p

umila

var

. m

ulti

fl o

ra, M

. m

ulti

fl or

a va

r. m

ulti

fl or

a, N

utta

llia

mul

tifl

ora

dese

rt s

tick

leaf

PerF

Wid

ePr

esU

ncA

lexa

nder

12

14 (P

ISP)

1998

Mal

vace

aeA

ltha

ea r

osea

Alc

ea r

osea

holly

hock

PerF

Intr

oRe

p?

Kur

th (1

998)

1997

Nat

ive

to C

hina

Mal

vace

aeM

alva

neg

lect

aco

mm

on

mal

low

Ann

FIn

tro

Pres

Com

Ale

xand

er

1231

(PIS

P)19

97N

ativ

e to

Eur

asia

Mal

vace

aeSp

haer

alce

a am

bigu

a va

r. am

bigu

aM

ohav

e gl

obem

allo

wPe

rFW

ide

Pres

Unc

Ale

xand

er

1255

(PIS

P)19

35

Mal

vace

aeSp

haer

alce

a pa

rvif

olia

smal

l-lea

f gl

obem

allo

wPe

rFW

ide

His

t?

Whi

tehe

ad s

.n.

(PIS

P)19

36

Nyc

tagi

nace

aeA

bron

ia f

ragr

ans

A. f

ragr

ans

var.

ellip

tica

, A. n

ana

var.

harr

isii,

incl

udes

A.

ellip

tica

frag

rant

san

d-ve

rben

aPe

rFW

ide

Pres

Unc

Hea

ton

20

(PIS

P)19

37O

bser

ved

vege

tati

vely

by

Fert

ig, 2

007

Nyc

tagi

nace

aeM

irab

ilis

mul

tifl

ora

M. m

ulti

fl o

ra

var.

mul

tifl

ora

, Q

uam

oclid

ion

mul

tifl

orum

show

y fo

ur-

o’cl

ock

PerF

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

18

(PIS

P)

1935

Ole

acea

eM

enod

ora

scab

raro

ugh

men

odor

aPe

rFW

ide

Pres

Unc

Ale

xand

er

1200

(PIS

P)19

98

Ona

grac

eae

Cam

isso

nia

ch

amae

neri

oide

sO

eno

ther

a ch

amae

neri

oide

ssl

ende

r-po

d ca

mis

soni

aA

nnF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

24

(PIS

P)

2001

Ona

grac

eae

Cam

isso

nia

parr

yiO

enot

hera

par

ryi

Parr

y’s

cam

isso

nia

Ann

FRe

gEn

His

t?

Hea

ton

25

(PIS

P)19

36Ty

pe lo

calit

y is

ne

ar S

t. G

eorg

e,

Was

hing

ton

Co.,

UT

(Par

ry 7

2 G

H)

Ona

grac

eae

Cam

isso

nia

parv

ula

Oen

othe

ra c

onto

rta

var.

fl ex

uosa

tiny

cam

isso

nia

Ann

FPe

riph

His

t?

Hea

ton

s.n.

(P

ISP)

1936

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Ona

grac

eae

Epilo

bium

cili

atum

E. c

iliat

um v

ar.

cilia

tum

, E. w

atso

nii,

E. a

deno

caul

on

nort

hern

w

illow

-her

b Pe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)19

98

Ona

grac

eae

Gau

ra c

occi

nea

scar

let

gaur

aPe

rFW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1939

Ona

grac

eae

Gau

ra p

arvi

fl or

aG

. mol

lissm

all-fl

ow

ered

ga

ura

Ann

FW

ide

Pres

Unc

Ale

xand

er

1224

(PIS

P)19

98

Ona

grac

eae

Oen

othe

ra a

lbic

aulis

whi

te-s

tem

ev

enin

g pr

imro

se

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Ona

grac

eae

Oen

othe

ra c

aesp

itos

a va

r. na

vajo

ensi

sPa

ria

even

ing

prim

rose

PerF

RegE

nPr

esU

ncEv

ende

n &

Lo

uie

PISP

20

(PIS

P)

1998

Type

loca

lity

is P

aria

Riv

er,

Kan

e Co

., U

T (C

ronq

uist

102

02

MO

)

Ona

grac

eae

Oen

othe

ra c

alif

orni

caCa

lifor

nia

even

ing

prim

rose

PerF

Wid

ePr

esU

ncA

lexa

nder

12

05 (P

ISP)

2001

Ona

grac

eae

Oen

othe

ra p

allid

a va

r. pa

llida

Incl

udes

var

. ru

ncin

ata

pale

eve

ning

pr

imro

sePe

rFW

ide

Pres

Unc

Ale

xand

er

1206

(PIS

P)19

36

Ona

grac

eae

Oen

othe

ra p

rim

iver

is

var.

bufo

nus

Oen

othe

ra p

rim

iver

isea

rly

even

ing

prim

rose

Ann

FRe

gEn

Rep

?A

lexa

nder

(1

998b

)N

A

Oro

banc

hace

aeO

rob

anch

e lu

dovi

cian

aO

. lud

ovic

iana

var

. lu

dovi

cian

a, O

. co

oper

i; in

clud

es O

. m

ulti

fl or

a fr

om U

tah

Fl

ora

(199

3) a

nd v

ars.

co

oper

i & a

rane

osa

Coop

er’s

broo

mra

pePe

rFW

ide

Pres

Unc

Whi

tehe

ad s

.n.

(PIS

P)19

35

Papa

vera

ceae

Esch

scho

lzia

m

inut

ifl or

apy

gmy

gold

en-

popp

yA

nnF

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

5

(PIS

P)

1935

Pina

ceae

Pinu

s ed

ulis

two-

need

le

piny

on

Tree

Wid

ePr

esU

ncA

lexa

nder

12

48 (P

ISP)

1935

Plan

tagi

nace

aePl

anta

go la

nceo

lata

Engl

ish

plan

tain

PerF

Intr

oRe

p?

Ale

xand

er

(199

8b)

1998

Nat

ive

to E

uras

ia

Plan

tagi

nace

aePl

anta

go m

ajor

Incl

udes

var

. pa

chyp

hylla

com

mon

pl

anta

inPe

rFIn

tro

Pres

Rare

Ale

xand

er

1189

(PIS

P)19

35M

ostl

y na

tive

to

Euro

pe

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 39

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Pole

mon

iace

aeEr

iast

rum

ere

mic

umM

ohav

e er

iast

rum

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1941

Pole

mon

iace

aeG

ilia

inco

nspi

cua

Incl

udes

var

s.

inco

nspi

ca &

sin

uat

a,

G. o

phth

alm

oide

s, G

. si

nuat

a, G

ilia

clok

eyi

shy

gilia

Ann

FW

ide

Pres

Unc

Even

den

&

Loui

e PI

SP 1

3 (P

ISP)

1936

If r

ecog

nize

d,

plan

ts f

rom

PIS

P re

pres

ent

vars

. in

cons

picu

a an

d si

nuat

a

Pole

mon

iace

aeG

ilia

lept

omer

ia v

ar.

lept

omer

iaA

licie

lla le

ptom

eria

com

mon

gili

aA

nnF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

37

Pole

mon

iace

aeIp

omop

sis

long

ifl or

aG

ilia

long

ifl or

alo

ng-fl

ow

ered

gi

liaPe

rFW

ide

Pres

Unc

Ale

xand

er

1211

(PIS

P)19

36

Pole

mon

iace

aeIp

omop

sis

poly

clad

onG

ilia

poly

clad

onsp

read

ing

gilia

Ann

FW

ide

His

t?

Hea

ton

30

(PIS

P)19

37

Pole

mon

iace

aeLa

nglo

isia

set

osis

sim

aM

ohav

e la

nglo

isia

Ann

FW

ide

His

t?

Hea

ton

21

(PIS

P)19

37

Poly

gala

ceae

Poly

gala

sub

spin

osa

P. s

ubsp

inos

a va

r. su

bspi

nosa

cush

ion

milk

wor

tPe

rFW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1939

Poly

gona

ceae

Cent

rost

egia

thu

rber

iCh

oriz

anth

e th

urbe

riTh

urbe

r’s

spin

efl o

wer

Ann

FW

ide

His

t?

Hea

ton

19

(PIS

P)19

37

Poly

gona

ceae

Erio

gonu

m c

ernu

um

var.

cern

uum

Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

A

mer

ica

(200

5)

nodd

ing

wild

bu

ckw

heat

Ann

FW

ide

His

t?

Hea

ton

s.n.

(P

ISP)

1936

Poly

gona

ceae

Erio

go

nu

m

cory

mbo

sum

var

. au

reum

E. a

ureu

m, i

nclu

des

var.

glut

inos

umgo

lden

wild

bu

ckw

heat

Shru

bRe

gEn

Rep

?K

ing

(197

7),

Ale

xand

er

(199

8b)

1977

Type

loca

lity

is S

t. G

eorg

e,

Was

hing

ton

Co.,

UT

(Jon

es

6091

PO

M).

Cons

ider

ed a

lo

cal e

ndem

ic

rest

rict

ed t

o th

e Sh

ivw

its

area

of

Was

hing

ton

Co.

in F

lora

of

Nor

th

Am

eric

a (2

005)

.

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Poly

gona

ceae

Erio

go

nu

m

cory

mbo

sum

var

. co

rym

bosu

m

Incl

udes

var

s.

velu

tinu

m,

diva

rica

tum

, er

ectu

m, &

dav

idse

an

d E.

lanc

ifol

ium

cris

plea

f w

ild

buck

whe

atSh

rub

Wid

ePr

esU

ncA

lexa

nder

12

28 (P

ISP)

2001

Poly

gona

ceae

Erio

gonu

m d

efl e

xum

E. d

efl e

xum

var

. de

fl ex

umsk

elet

onw

eed

wild

buc

kwhe

atA

nnF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

36

Poly

gona

ceae

Erio

gonu

m in

fl at

um

va

r. in

fl at

umE.

infl

atum

dese

rt t

rum

pet

PerF

Wid

ePr

esU

ncEv

ende

n &

Lo

uie

PISP

36

(PIS

P)

1935

Poly

gona

ceae

Erio

go

nu

m

mic

roth

ecum

var

. si

mps

onii

E. s

imps

onii,

E.

mic

roth

ecum

var

. fo

liosu

m; i

nclu

des

E.

phoe

nice

um

slen

der

wild

bu

ckw

heat

PerF

Wid

ePr

esU

ncA

lexa

nder

12

27 (P

ISP)

1977

Poly

gona

ceae

Erio

go

nu

m

palm

eria

num

Palm

er’s

wild

bu

ckw

heat

Ann

FW

ide

Pres

Unc

Loui

e &

Ev

ende

n PI

SP 7

(P

ISP)

2001

Poly

gona

ceae

Poly

go

nu

m

amph

ibiu

mP.

am

phib

ium

va

r. em

ersu

m, P

. co

ccin

eum

, Per

sica

ria

cocc

inea

, Per

sica

ria

amph

ibia

wat

er

smar

twee

dPe

rFW

ide

Rep

?H

eato

n (1

935)

1935

Poly

gona

ceae

Poly

gonu

m a

vicu

lare

P. a

rena

stru

m,

P. a

vicu

lare

ssp

. av

icul

are

yard

kno

twee

dA

nnF

Intr

oPr

esU

ncA

lexa

nder

12

30 (P

ISP)

1998

Nat

ive

to E

uras

ia.

PISP

spe

cim

en

is “

aren

astr

um”

phas

e.

Poly

gona

ceae

Rum

ex c

risp

uscu

rly

dock

PerF

Intr

oRe

p?

Kur

th (1

998)

1997

Nat

ive

to E

uras

ia

Poly

gona

ceae

Rum

ex h

ymen

osep

alus

cana

igre

doc

kPe

rFW

ide

Pres

Unc

Fert

ig 2

3672

(P

ISP)

1998

Port

ulac

acea

ePo

rtul

aca

oler

acea

com

mon

pu

rsla

neA

nnF

Intr

oPr

esCo

mFe

rtig

236

63

(PIS

P)19

98A

Z st

ate

proh

ibit

ed

and

regu

late

d no

xiou

s w

eed.

N

ativ

e to

eas

tern

N

orth

Am

eric

a

Ranu

ncul

acea

e (H

elle

bora

ceae

)D

elph

iniu

m a

nder

soni

i va

r. sc

apos

umD

. sca

posu

mpa

le la

rksp

urPe

rFW

ide

His

t?

Peeb

les

1307

7 (A

RIZ)

1936

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 41

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Rosa

ceae

Am

elan

chie

r ut

ahen

sis

Uta

h se

rvic

eber

rySh

rub

Wid

ePr

esU

ncN

elso

n 93

08

(RM

)19

71Ty

pe lo

calit

y is

Toq

uerv

ille,

W

ashi

ngto

n Co

., U

T (J

ones

171

6 G

H)

Rosa

ceae

Fallu

gia

para

doxa

Apa

che

plum

eSh

rub

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

38

(PIS

P)

1936

Rosa

ceae

Pote

ntill

a gr

acili

s va

r. el

mer

iP.

pec

tini

sect

aco

mb-

leaf

ci

nque

foil

PerF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

39

Rosa

ceae

Prun

us a

mer

ican

aA

mer

ican

plu

mSh

rub

Intr

oPr

esU

ncFe

rtig

236

89

(PIS

P)19

98N

ativ

e to

cen

tral

N

orth

Am

eric

a

Rosa

ceae

Purs

hia

mex

ican

a va

r. st

ansb

urya

naP.

sta

nsbu

rian

a,

Cow

ania

mex

ican

acl

iff-

rose

Shru

bW

ide

Pres

Unc

Hea

ton

s.n.

(P

ISP)

1936

Rosa

ceae

Rosa

woo

dsii

R. a

rizo

nica

, R.

woo

dsii

var.

ultr

amon

tana

. In

clud

es R

. man

ca.

Woo

ds’ r

ose

Shru

bW

ide

Pres

Unc

Ale

xand

er

1239

(PIS

P)19

77Ci

ted

by K

ing

(197

7) a

s R

. ar

izon

ica

Salic

acea

ePo

pulu

s al

baw

hite

pop

lar

Tree

Intr

oPr

esU

ncA

lexa

nder

12

53 (P

ISP)

1997

Nat

ive

to E

uras

ia

Salic

acea

ePo

pulu

s fr

emon

tii v

ar.

frem

onti

iP.

fre

mon

tii,

P.

delt

oide

s va

r. fr

emon

tii

Frem

ont

cott

onw

ood

Tree

Wid

ePr

esU

ncA

lexa

nder

12

60 (P

ISP)

1998

Salic

acea

ePo

pulu

s ni

gra

Lom

bard

y po

plar

Tree

Intr

oRe

p?

Ale

xand

er

(199

8b)

1998

Nat

ive

to E

uras

ia

Salic

acea

ePo

pulu

s x

cana

dens

isH

ybri

d be

twee

n P.

de

ltoi

des

x P.

nig

raCa

rolin

a po

plar

Tree

Intr

oPr

esU

ncFe

rtig

236

87

(PIS

P)20

07

Salic

acea

ePo

pulu

s x

jack

iiH

ybri

d be

twee

n P.

bal

sam

ifer

a x

P.

delt

oide

s

balm

of

Gile

adTr

eeIn

tro

Pres

Unc

Nel

son

s.n.

(R

M)

1973

Salic

acea

eSa

lix e

xigu

a va

r. st

enop

hylla

S. e

xigu

a va

r. ex

igua

coyo

te w

illow

Shru

bW

ide

Pres

Unc

Ale

xand

er

1259

(PIS

P)19

98

Saxi

frag

acea

e (G

ross

ular

iace

ae)

Ribe

s au

reum

R.

aur

eum

var

. au

reum

gold

en c

urra

ntSh

rub

Wid

ePr

esU

ncFe

rtig

236

85

(PIS

P)20

07

Saxi

frag

acea

e (G

ross

ular

iace

ae)

Ribe

s sa

tivu

mR.

rub

rum

gard

en c

urra

ntSh

rub

Intr

oRe

p?

Ale

xand

er

(199

8b)

1998

Nat

ive

to E

urop

e

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Scro

phul

aria

ceae

Cast

illej

a ch

rom

osa

C. a

pple

gate

i ss

p. m

arti

nii,

C.

angu

stif

olia

var

. du

bia

dese

rt

pain

tbru

shPe

rFW

ide

Pres

Unc

Even

den

&

Loui

e 29

(PIS

P)20

01

Scro

phul

aria

ceae

Cast

illej

a lin

ariif

olia

C. li

nari

aefo

liaW

yom

ing

pain

tbru

shPe

rFW

ide

Pres

Unc

Ale

xand

er

1195

(PIS

P)19

98

Scro

phul

aria

ceae

Cord

ylan

thus

pa

rvifl

orus

smal

lfl ow

er

bird

’s-be

akA

nnF

Wid

ePr

esU

ncA

lexa

nder

12

29 (P

ISP)

1935

Scro

phul

aria

ceae

Pens

tem

on la

evis

smoo

th

pens

tem

onPe

rFRe

gEn

Rep

?A

lexa

nder

(1

998b

)19

98Ty

pe lo

calit

y is

Spr

ingd

ale,

W

ashi

ngto

n Co

., U

T (J

ones

525

0 in

pa

rt U

S)

Scro

phul

aria

ceae

Pens

tem

on p

alm

eri

var.

egla

ndul

osus

Palm

er’s

pens

tem

onPe

rFRe

gEn

Pres

Unc

Fert

ig 2

3666

(P

ISP)

1998

Type

loca

lity

is

2.5

mile

s N

of

Kan

ab, K

ane

Co.,

UT

(Mag

uire

, M

agui

re, &

Pi

rani

an 1

2279

U

TC)

Scro

phul

aria

ceae

Pens

tem

on u

tahe

nsis

Uta

h pe

nste

mon

PerF

Wid

eH

ist

?H

eato

n 11

(P

ISP)

1937

Sim

arou

bace

aeA

ilant

hus

alti

ssim

atr

ee-o

f-he

aven

Tree

Intr

oPr

esU

ncFe

rtig

236

84

(PIS

P)19

35N

ativ

e to

Chi

na

Sola

nace

aeCh

amae

sara

cha

coro

nopu

sfa

lse

nigh

tsha

dePe

rFW

ide

Pres

Rare

Fert

ig 2

3695

(P

ISP)

2007

Sola

nace

aeD

atur

a w

righ

tii

Dat

ura

met

eloi

des

ange

l’s t

rum

pet

Ann

FW

ide

Pres

Unc

Hea

ton

(193

5)19

35

Sola

nace

aeLy

cium

pal

lidum

pale

wol

fber

rySh

rub

Wid

ePr

esU

ncFe

rtig

236

73

(PIS

P)19

77

Sola

nace

aeN

ico

tian

a tr

igon

ophy

llaN

. obt

usif

olia

; in

clud

es v

ars.

tr

igon

ophy

lla &

pa

lmer

i

dese

rt t

obac

coA

nnF

Wid

eH

ist

?H

eato

n s.

n.

(PIS

P)19

39

Sola

nace

aePh

ysal

is c

rass

ifol

iath

ick-

leav

ed

grou

nd-c

herr

yPe

rFW

ide

Rep

?A

lexa

nder

(1

998b

)19

98

Sola

nace

aePh

ysal

is h

eder

ifol

ia

var.

palm

eri

P. h

eder

aefo

lia v

ar.

palm

eri

Palm

er’s

grou

nd-c

herr

yPe

rFRe

gEn

Rep

?H

eato

n (1

935)

1935

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.

Appendices 43

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

onna

me

Life

fo

rmRa

nge

Park

st

atus

Pop.

si

zeSo

urce

Year

do

c.Co

mm

ents

Sola

nace

aeSo

lanu

m s

arra

choi

des

S. v

illos

um, S

. ph

ysal

ifol

ium

var

. ni

tidi

bacc

atum

grou

nd-c

herr

y ni

ghts

hade

Ann

FIn

tro

Rep

?A

lexa

nder

(1

998b

)N

AN

ativ

e to

Sou

th

Am

eric

a

Tam

aric

acea

eTa

mar

ix c

hine

nsis

T. p

enta

ndra

, T.

ram

osis

sim

afi

ve-s

tam

en

tam

aris

kSh

rub

Intr

oRe

p?

Kin

g (1

977)

, K

urth

(199

8)19

77N

ativ

e to

Eur

asia

Ulm

acea

eU

lmus

pum

ilaSi

beri

an e

lmTr

eeIn

tro

Pres

Unc

Nel

son

s.n.

(R

M)

1973

Nat

ive

to A

sia

Um

belli

fera

e (A

piac

eae)

Beru

la e

rect

a va

r. in

cisa

B. e

rect

acu

tlea

f w

ater

-pa

rsni

pPe

rFW

ide

Pres

Rare

Ale

xand

er

1245

(PIS

P)19

98

Um

belli

fera

e (A

piac

eae)

Cym

op

teru

s m

ulti

nerv

atus

purp

le-n

erve

d sp

ring

-par

sley

PerF

Wid

ePr

esU

ncLo

uie

&

Even

den

PISP

-2

(PIS

P)

1937

Ver

bena

ceae

Ver

bena

bra

ctea

tapr

ostr

ate

verv

ain

PerF

Wid

ePr

esU

ncA

lexa

nder

12

16 (P

ISP)

1997

Vis

cace

aePh

orad

endr

on

juni

peri

num

juni

per

mis

tlet

oePe

rFW

ide

Pres

Unc

Fert

ig 2

3677

(P

ISP)

1998

Typi

cally

par

asit

ic

on J

un

iper

us

oste

ospe

rma

Zygo

phyl

lace

aeLa

rrea

tri

dent

ata

creo

sote

bus

hSh

rub

Wid

ePr

esRa

reA

lexa

nder

12

37 (P

ISP)

2001

Zygo

phyl

lace

aeTr

ibul

us t

erre

stri

spu

nctu

revi

neA

nnF

Intr

oPr

esU

ncA

lexa

nder

12

33 (P

ISP)

1997

AZ

stat

e pr

ohib

ited

an

d re

gula

ted

noxi

ous

wee

d.

Nat

ive

to E

uras

ia.

App

endi

x A

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed v

ascu

lar

plan

t ta

xa o

f Pi

pe S

prin

g N

atio

nal M

onum

ent,

org

aniz

ed b

y fa

mily

and

sci

enti

fi c

nam

e w

ith

taxo

nom

ic

and

biog

eogr

aphi

c no

tes,

con

t.


App

endi

x B.

Pot

enti

al (u

ncon

fi rm

ed) t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent.

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

on n

ame

Life

fo

rmRa

nge

Com

men

ts

Am

aran

thac

eae

Am

aran

thus

alb

ustu

mbl

e pi

gwee

dA

nnF

Intr

oN

ativ

e to

tro

pica

l Am

eric

a

Am

aran

thac

eae

Am

aran

thu

s re

trofl

exu

sre

droo

t pi

gwee

dA

nnF

Intr

oN

ativ

e to

Cen

tral

Am

eric

a

Asc

lepi

adac

eae

Asc

lepi

as s

peci

osa

show

y m

ilkw

eed

PerF

Wid

e

Bora

gina

ceae

Cryp

tant

ha h

umili

sO

reoc

arya

hum

ilis

dwar

f cr

ypta

nth

PerF

Peri

ph

Bora

gina

ceae

Cryp

tant

ha r

ecur

vata

recu

rved

cry

ptan

thA

nnF

Wid

e

Bora

gina

ceae

Cyno

glos

sum

offi

cin

ale

com

mon

hou

nd’s-

tong

uePe

rFIn

tro

Nat

ive

to E

uras

ia

Bora

gina

ceae

Lith

ospe

rmum

inci

sum

show

y st

ones

eed

PerF

Wid

e

Cact

acea

ePe

dioc

actu

s si

leri

Uta

hia

sile

riSi

ler’s

pin

cush

ion

cact

usPe

rFLo

cEn

Type

loca

lity

is P

ipe

Spri

ng,

Moh

ave

Co.,

AZ

(Sile

r s.

n.

MO

). U

SFW

S: T

hrea

tene

d

Capp

arac

eae

(Cle

omac

eae)

Cleo

me

serr

ulat

aRo

cky

Mou

ntai

n be

epla

ntA

bbF

Wid

e

Chen

opod

iace

aeA

trip

lex

conf

erti

folia

shad

scal

eSh

rub

Wid

e

Chen

opod

iace

aeBa

ssia

sco

pari

aK

ochi

a sc

opar

ia s

sp. s

cop

aria

, B.

sie

vers

iana

sum

mer

-cyp

ress

Ann

FIn

tro

Nat

ive

to E

uras

ia

Chen

opod

iace

aeC

hen

op

od

ium

de

sicc

atum

Som

e au

thor

s in

clud

e C

. pr

ater

icol

ade

sert

goo

sefo

otA

nnF

Peri

ph

Chen

opod

iace

aeCh

enop

odiu

m g

lauc

um

var.

salin

umC.

sal

inum

oakl

eaf

goos

efoo

tA

nnF

Wid

e

Chen

opod

iace

aeH

alog

eton

glo

mer

atus

halo

geto

nA

nnF

Intr

oN

ativ

e to

Eur

asia

Chen

opod

iace

aeK

rasc

heni

nnik

ovia

la

nata

var

. lan

ata

Cera

toid

es la

nata

, Eur

otia

la

nata

. Var

s. n

ot r

ecog

nize

d in

Flo

ra o

f N

orth

Am

eric

a (2

003)

.

win

terf

atSh

rub

Wid

e

Chen

opod

iace

aeSa

lsol

a pa

ulse

nii

barb

wir

e Ru

ssia

n th

istl

eA

nnF

Intr

oN

ativ

e to

Eur

asia

Com

posi

tae

(Ast

erac

eae)

Aca

mpt

opap

pus

spha

eroc

epha

lus

A. s

phae

roce

phal

us v

ar.

spha

eroc

epha

lus

gold

enhe

adSh

rub

Wid

e

Com

posi

tae

(Ast

erac

eae)

Arc

tium

min

usbu

rdoc

kPe

rFIn

tro

Nat

ive

to E

uras

ia

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia s

pine

scen

sPi

crot

ham

nus

dese

rtor

umbu

dsag

eSh

rub

Peri

ph

Com

posi

tae

(Ast

erac

eae)

Chae

nact

is d

ougl

asii

Incl

udes

var

s. d

ougl

asii

&

mon

tana

hoar

y du

sty-

mai

den

PerF

Wid

e

Com

posi

tae

(Ast

erac

eae)

Chae

nact

is f

rem

onti

iFr

emon

t’s d

usty

-mai

den

Ann

FW

ide

Com

posi

tae

(Ast

erac

eae)

Chae

nact

is s

tevi

oide

sst

evia

dus

ty-m

aide

nA

nnF

Wid

e

Appendices 45

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

on n

ame

Life

fo

rmRa

nge

Com

men

ts

Com

posi

tae

(Ast

erac

eae)

Gly

ptop

leur

a se

tulo

sase

tose

cru

stw

eed

Ann

FW

ide

Type

loca

lity

is n

ear

St.

Geo

rge,

Was

hing

ton

Co.,

UT

(Pal

mer

6 G

H)

Com

posi

tae

(Ast

erac

eae)

Hel

iant

hus

peti

olar

is

ssp.

fal

lax

Incl

udes

var

. pet

iola

ris

prai

rie

sunfl

ow

erA

nnF

Wid

e

Com

posi

tae

(Ast

erac

eae)

Tara

xacu

m o

ffi c

inal

eco

mm

on d

ande

lion

PerF

Intr

oN

ativ

e to

Eur

asia

Cruc

ifer

ae (B

rass

icac

eae)

Chor

ispo

ra t

enel

labl

ue m

usta

rdA

nnF

Intr

oN

ativ

e to

Asi

a

Cruc

ifer

ae (B

rass

icac

eae)

Lepi

dium

vir

gini

cum

va

r. pu

besc

ens

poor

-man

’s pe

pper

wor

tA

nnF

Wid

e

Cruc

ifer

ae (B

rass

icac

eae)

Mal

colm

ia a

fric

ana

Afr

ican

mus

tard

Ann

FIn

tro

Nat

ive

to A

fric

a

Cruc

ifer

ae (B

rass

icac

eae)

Scho

enoc

ram

be

linif

olia

Sisy

mbr

ium

lini

foliu

mfl

ax-le

aved

pla

insm

usta

rdPe

rFW

ide

Gra

min

eae

(Poa

ceae

)Er

ione

uron

pilo

sum

hair

y tr

iden

sPe

rGW

ide

Gra

min

eae

(Poa

ceae

)Fe

stuc

a oc

tofl

ora

Vul

pia

octo

fl or

a; in

clud

es

vars

. gla

uca,

hir

tella

, &

octo

fl or

a

sixw

eeks

fes

cue

Ann

GW

ide

Gra

min

eae

(Poa

ceae

)Tr

itic

um a

esti

vum

whe

atA

nnG

Intr

oN

ativ

e to

Eur

asia

Legu

min

osae

(Fab

acea

e)A

stra

galu

s m

ollis

sim

us

var.

thom

pson

iae

woo

lly m

ilkve

tch

PerF

Wid

eTy

pe lo

calit

y is

Kan

ab,

Kan

e Co

., U

T (T

hom

pson

s.

n. G

H)

Legu

min

osae

(Fab

acea

e)Lu

pinu

s ar

gent

eus

var.

arge

nteu

ssi

lver

y lu

pine

PerF

Wid

e

Nyc

tagi

nace

aeTr

ipte

roca

lyx

mic

rant

hus

Abr

onia

mic

rant

ha, T

. pe

dunc

ulat

ussm

all-fl

ow

er s

andp

uffs

Ann

FW

ide

Ona

grac

eae

Oen

oth

era

coro

nopi

folia

hair

y-th

roat

eve

ning

-pr

imro

sePe

rFW

ide

Plan

tagi

nace

aePl

anta

go p

atag

onic

aP.

pat

agon

ica

var.

pata

goni

ca, P

. pur

shii;

in

clud

es v

ars.

gna

phal

oide

s,

spin

ulos

a, &

bre

visc

apa

woo

lly p

lant

ain

Ann

FW

ide

Pole

mon

iace

aeLi

nant

hus

dich

otom

usev

enin

g-sn

owA

nnF

Wid

e

Pole

mon

iace

aePh

lox

long

ifol

iaIn

clud

es v

ars.

long

ifol

ia &

st

ansb

uryi

long

-leaf

phl

oxPe

rFW

ide

Poly

gona

ceae

Erio

go

nu

m

cory

mbo

sum

var

. th

omps

onia

e

E. t

hom

pson

iae

var.

thom

pson

iae

Thom

pson

’s w

ild

buck

whe

atSh

rub

RegE

nTy

pe lo

calit

y is

nea

r K

anab

, K

ane

Co.,

UT

(Tho

mps

on

s.n.

GH

). U

SFW

S: 3

C

App

endi

x B.

Pot

enti

al (u

ncon

fi rm

ed) t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent,

con

t.


Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

on n

ame

Life

fo

rmRa

nge

Com

men

ts

Poly

gona

ceae

Poly

go

nu

m

lapa

thif

oliu

mPe

rsic

aria

lapa

thif

olia

will

ow-w

eed

Ann

FIn

tro

Nat

ive

to E

uras

ia

Ranu

ncul

acea

eRa

nunc

ulus

inam

oenu

s R.

inam

oenu

s va

r. in

amoe

nus.

Incl

udes

var

. al

peop

hilu

s.

drab

but

terc

upPe

rFW

ide

Ranu

ncul

acea

eRa

nunc

ulus

tes

ticu

latu

sCe

rato

ceph

ala

test

icul

ata,

C.

orth

ocer

asbu

r bu

tter

cup

Ann

FIn

tro

Nat

ive

to E

uras

ia

Sant

alac

eae

Com

andr

a um

bella

ta

var.

palli

daC.

pal

lida

bast

ard

toad

fl ax

PerF

Wid

e

Scro

phul

aria

ceae

Cast

illej

a sc

abri

da v

ar.

scab

rida

East

woo

d’s

pain

tbru

shPe

rFRe

gEn

Sola

nace

aeLy

cium

and

erso

nii

And

erso

n’s

wol

fber

rySh

rub

Wid

e

App

endi

x B.

Pot

enti

al (u

ncon

fi rm

ed) t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent,

con

t.

Appendices 47

App

endi

x C.

Rej

ecte

d (f

alse

ly r

epor

ted

or q

uest

iona

ble)

tax

a of

Pip

e Sp

ring

Nat

iona

l Mon

umen

t.

Fam

ilySp

ecie

s na

me

Syno

nym

s/Ta

xono

mic

not

esCo

mm

on n

ame

Life

fo

rmRa

nge

Park

st

atus

Com

men

ts

Aga

vace

ae

(Lili

acea

e)Yu

cca

angu

stis

sim

aY.

ang

usti

ssim

a va

r. an

gust

issi

ma

narr

owle

aved

yuc

caSh

rub

RegE

nQ

ues?

Repo

rted

by

Kin

g (1

977)

, but

pro

babl

y ba

sed

on Y

. ka

nabe

nsis

.

Bora

gina

ceae

Cry

pta

nth

a ba

rbig

era

bear

ded

cryp

tant

haA

nnF

Wid

eFa

lsRe

pRe

port

fro

m P

ISP

herb

ariu

m b

ased

on

a m

isid

enti

fi ed

spe

cim

en o

f Pl

agio

both

rys

jone

sii.

Type

lo

calit

y is

St.

Geo

rge,

Was

hing

ton

Co.,

UT

(Par

ry 1

71

GH

).

Cact

acea

eO

punt

ia b

asila

ris

O. b

asila

ris

var.

basi

lari

sbe

aver

tail

cact

usPe

rFW

ide

Que

s?Re

port

ed b

y K

ing

(197

7), b

ut p

roba

bly

base

d on

O.

erin

acea

var

. aur

ea.

Chen

opod

iace

aeA

trip

lex

gard

neri

va

r. cu

neat

aA

. cun

eata

valle

y sa

ltbu

shSh

rub

RegE

nQ

ues?

Repo

rted

by

Hea

ton

(193

5), b

ut p

roba

bly

base

d on

A

. can

esce

ns.

Com

posi

tae

(Ast

erac

eae)

Mad

ia g

lom

erat

am

ount

ain

tarw

eeed

Ann

FPe

riph

Fals

Rep

Repo

rt f

rom

Ale

xand

er (1

998a

) bas

ed o

n a

mis

iden

tifi

ed s

peci

men

of

Cord

ylan

thus

par

vifl

orus

.

Com

posi

tae

(Ast

erac

eae)

Sonc

hus

arve

nsis

spin

y-le

af s

owth

istl

eA

nnF

Intr

oFa

lsRe

pRe

port

fro

m P

ISP

herb

ariu

m b

ased

on

a m

isid

enti

fi ed

spe

cim

en o

f S.

asp

er. N

ativ

e to

Eur

ope.

A

Z st

ate

proh

ibit

ed n

oxio

us w

eed.

Faga

ceae

Que

rcus

dum

osa

coas

tal s

age

scru

b oa

kSh

rub

RegE

nQ

ues?

Repo

rted

by

Hea

ton

(193

5), b

ut p

roba

bly

base

d on

Q

. tur

bine

lla. Q

. dum

osa

occu

rs in

Cal

ifor

nia

but

is

not

know

n fr

om A

rizo

na.

Hyd

roph

ylla

ceae

Phac

elia

am

bigu

aP.

cre

nula

ta v

ar.

ambi

gua

Jone

s’ p

hace

liaA

nnF

Wid

eFa

lsRe

pRe

port

fro

m P

ISP

herb

ariu

m b

ased

on

a m

isid

enti

fi ed

spe

cim

en o

f P.

cre

nula

ta v

ar. c

renu

lata

.

Loas

acea

eM

entz

elia

nit

ens

M. n

iten

s va

r. jo

nesi

icu

rvep

od s

tick

leaf

Ann

FW

ide

Fals

Rep

Repo

rt f

rom

PIS

P he

rbar

ium

bas

ed o

n a

mis

iden

tifi

ed s

peci

men

of

M. a

lbic

aulis

.

Loas

acea

eM

entz

elia

pu

mila

var

. pu

mila

M. p

umila

Wyo

min

g st

ickl

eaf

PerF

RegE

nFa

lsRe

pRe

port

fro

m A

lexa

nder

(199

8a) b

ased

on

a m

isid

enti

fi ed

spe

cim

en o

f M

. mul

tifl

ora.

Not

kno

wn

from

Ari

zona

Mal

vace

aeSp

haer

alce

a la

xaca

liche

glo

be m

allo

wPe

rFW

ide

Que

s?Re

port

ed b

y K

ing

(197

7), b

ut p

roba

bly

base

d on

S.

ambi

gua.

Ona

grac

eae

Oen

oth

era

cesp

itos

a va

r. m

argi

nata

long

-tub

e ev

enin

g-pr

imro

sePe

rFW

ide

Fals

Rep

Repo

rt f

rom

PIS

P he

rbar

ium

bas

ed o

n a

mis

iden

tifi

ed s

peci

men

of

O. c

espi

tosa

var

. na

vajo

ensi

s.

Pole

mon

iace

aeG

ilia

scop

ulor

umro

ck g

ilia

Ann

FW

ide

Fals

Rep

Repo

rt f

rom

PIS

P he

rbar

ium

bas

ed o

n a

mis

iden

tifi

ed s

peci

men

of

G. i

ncon

spic

ua.T

ype

loca

lity

is S

t. G

eorg

e, W

ashi

ngto

n Co

., U

T (J

ones

16

59 P

OM

).

Scro

phul

aria

ceae

Cord

ylan

thus

ne

vini

iN

evin

’s bi

rd’s-

beak

Ann

FRe

gEn

Que

s?Re

port

ed b

y H

eato

n (1

935)

, but

pro

babl

y ba

sed

on

C. p

arvi

fl or

us.

Um

belli

fera

e (A

piac

eae)

Cym

op

teru

s pu

rpur

eus

C. p

urpu

reus

var

. pu

rpur

eus

Colo

rado

Pla

teau

sp

ring

-par

sley

PerF

RegE

nFa

lsRe

pRe

port

fro

m P

ISP

herb

ariu

m b

ased

on

a m

isid

enti

fi ed

spe

cim

en o

f C.

mul

tine

rvat

us.


App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Tree

s

Cupr

essa

ceae

Juni

peru

s sc

opul

orum

Rock

y M

ount

ain

juni

per

Nat

ive

Rep

NA

xx

Legu

min

osae

(Fab

acea

e)Ro

bini

a ps

eudo

acac

iabl

ack

locu

stIn

tro

Pres

May

–Jun

Cult

ivat

ed

Pina

ceae

Pinu

s ed

ulis

two-

need

le p

inyo

n N

ativ

ePr

esN

Ax

Salic

acea

ePo

pulu

s al

baw

hite

pop

lar

Intr

oPr

esM

ar–J

unCu

ltiv

ated

Salic

acea

ePo

pulu

s fr

emon

tii v

ar.

frem

onti

iFr

emon

t co

tton

woo

dN

ativ

ePr

esM

ar–J

unx

Salic

acea

ePo

pulu

s ni

gra

Lom

bard

y po

plar

Intr

oRe

pA

pr–M

ayCu

ltiv

ated

Salic

acea

ePo

pulu

s x

cana

dens

isCa

rolin

a po

plar

Intr

oPr

esA

pr–M

ayCu

ltiv

ated

Salic

acea

ePo

pulu

s x

jack

iiba

lm o

f G

ilead

Intr

oPr

esA

pr–M

ayCu

ltiv

ated

Sim

arou

bace

aeA

ilant

hus

alti

ssim

atr

ee-o

f-he

aven

Intr

oPr

esJu

n–Ju

lD

istu

rbed

are

as

Ulm

acea

eU

lmus

pum

ilaSi

beri

an e

lmIn

tro

Pres

Mar

–Apr

xD

istu

rbed

are

as

Shru

bs

Aga

vace

ae (L

iliac

eae)

Yucc

a ba

ccat

ada

til y

ucca

Nat

ive

Pres

Apr

–Jun

xx

Aga

vace

ae (L

iliac

eae)

Yucc

a ka

nabe

nsis

Kan

ab y

ucca

Nat

ive

Pres

May

–Jul

xx

Ana

card

iace

aeRh

us a

rom

atic

a va

r. si

mpl

icif

olia

squa

wbu

shN

ativ

ePr

esM

ay–J

ulx

Ana

card

iace

aeRh

us a

rom

atic

a va

r. tr

iloba

tasq

uaw

bush

Nat

ive

Rep

May

–Jul

x

Cact

acea

eO

punt

ia w

hipp

lei v

ar.

whi

pple

iW

hipp

le’s

chol

laN

ativ

ePr

esM

ay–J

ulx

Chen

opod

iace

aeA

trip

lex

cane

scen

s va

r. ca

nesc

ens

four

win

g sa

ltbu

shN

ativ

ePr

esM

ay–S

epx

x

Chen

opod

iace

aeK

rasc

heni

nnik

ovia

lana

ta v

ar.

subs

pino

sash

rubb

y w

inte

rfat

Nat

ive

His

tM

ay–J

ulx

Chen

opod

iace

ae

(Sar

coba

tace

ae)

Sarc

obat

us v

erm

icul

atus

grea

sew

ood

Nat

ive

Pres

May

–Aug

x

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia b

igel

ovii

Bige

low

’s sa

gebr

ush

Nat

ive

Pres

Aug

–Oct

xx

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia fi

lifol

iasa

nd s

ageb

rush

N

ativ

ePr

esA

ug–O

ctx

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia t

ride

ntat

a va

r. tr

iden

tata

basi

n bi

g sa

gebr

ush

Nat

ive

Pres

Jul–

Oct

x

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

cons

imili

sgr

eeni

sh r

abbi

tbru

shN

ativ

ePr

esA

ug–O

ctx

Appendices 49

App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Shru

bs, c

ont.

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

gnap

halo

des

gray

stem

rab

bitb

rush

Nat

ive

Pres

Aug

–Oct

x

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

grav

eole

nsgl

abra

te r

abbi

tbru

shN

ativ

eRe

pA

ug–O

ctx

xx

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s na

useo

sus

var.

junc

eus

rush

rab

bitb

rush

Nat

ive

Pres

Aug

–Oct

x

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s vi

scid

ifl o

rus

var.

pube

rulu

spu

beru

lent

rab

bitb

rush

Nat

ive

Pres

Jul–

Sep

x

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s vi

scid

ifl o

rus

var.

sten

ophy

llus

slen

derl

eaf

rabb

itbr

ush

Nat

ive

Pres

Jul–

Sep

xx

Com

posi

tae

(Ast

erac

eae)

Chry

soth

amnu

s vi

scid

ifl o

rus

var.

visc

idifl

orus

gree

n ra

bbit

brus

hN

ativ

eRe

pJu

l–Se

px

Com

posi

tae

(Ast

erac

eae)

Gut

ierr

ezia

mic

roce

phal

ath

read

sna

kew

eed

Nat

ive

Pres

Jul–

Oct

xx

Com

posi

tae

(Ast

erac

eae)

Gut

ierr

ezia

sar

othr

aebr

oom

sna

kew

eed

Nat

ive

Pres

Jul–

Oct

xx

Cupr

essa

ceae

Juni

peru

s os

teos

perm

aU

tah

juni

per

Nat

ive

Pres

NA

xx

Elae

agna

ceae

Shep

herd

ia r

otun

difo

liaro

undl

eaf

buff

alob

erry

Nat

ive

Pres

Mar

–May

xx

Ephe

drac

eae

Ephe

dra

torr

eyan

aTo

rrey

’s ep

hedr

aN

ativ

ePr

esN

Ax

Ephe

drac

eae

Ephe

dra

viri

dis

var.

viri

dis

gree

n ep

hedr

aN

ativ

ePr

esN

Ax

Ephe

drac

eae

Ephe

dra

viri

dis

var.

visc

ida

Cutl

er’s

ephe

dra

Nat

ive

Pres

NA

x

Faga

ceae

Que

rcus

tur

bine

llatu

rbin

ella

live

-oak

Nat

ive

Pres

x

Labi

atae

(Lam

iace

ae)

Salv

ia d

orri

i ssp

. dor

rii

Dor

r’s s

age

Nat

ive

Pres

May

–Jun

x

Poly

gona

ceae

Erio

gonu

m c

orym

bosu

m v

ar.

aure

umgo

lden

wild

buc

kwhe

atN

ativ

eRe

pA

ug–O

ctx

Poly

gona

ceae

Erio

gonu

m c

orym

bosu

m v

ar.

cory

mbo

sum

cris

plea

f w

ild b

uckw

heat

Nat

ive

Pres

Jul–

Oct

Rosa

ceae

Am

elan

chie

r ut

ahen

sis

Uta

h se

rvic

eber

ryN

ativ

ePr

esM

ay–J

unx

Rosa

ceae

Fallu

gia

para

doxa

Apa

che

plum

eN

ativ

ePr

esA

pr–A

ugx

Rosa

ceae

Prun

us a

mer

ican

aA

mer

ican

plu

mIn

tro

Pres

Cult

ivat

ed

Rosa

ceae

Purs

hia

mex

ican

a va

r. st

ansb

urya

nacl

iff-

rose

Nat

ive

Pres

May

–Jun

xx

Rosa

ceae

Rosa

woo

dsii

Woo

ds’ r

ose

Nat

ive

Pres

May

–Jul

xx

Salic

acea

eSa

lix e

xigu

a va

r. st

enop

hylla

coyo

te w

illow

Nat

ive

Pres

Apr

–Aug

x


App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Shru

bs, c

ont.

Saxi

frag

acea

e (G

ross

ular

iace

ae)

Ribe

s au

reum

go

lden

cur

rant

Nat

ive

Pres

Mar

–Jul

Dis

turb

ed a

reas

Saxi

frag

acea

e (G

ross

ular

iace

ae)

Ribe

s sa

tivu

mga

rden

cur

rant

Intr

oRe

pM

ayCu

ltiv

ated

Sola

nace

aeLy

cium

pal

lidum

pale

wol

fber

ryN

ativ

ePr

esM

ar–M

ayx

x

Tam

aric

acea

eTa

mar

ix c

hine

nsis

fi ve

-sta

men

tam

aris

kIn

tro

Rep

Apr

–Sep

x

Zygo

phyl

lace

aeLa

rrea

tri

dent

ata

creo

sote

bus

hN

ativ

ePr

esA

pr–J

unx

Pere

nnia

l For

bs

Apo

cyna

ceae

Am

soni

a jo

nesi

iJo

nes’

am

soni

aN

ativ

eH

ist

Apr

–Jun

xx

Apo

cyna

ceae

Am

soni

a to

men

tosa

var

. st

enop

hylla

tom

ento

se a

mso

nia

Nat

ive

His

tA

pr–M

ayx

Apo

cyna

ceae

Apo

cynu

m c

anna

binu

mco

mm

on d

ogba

neN

ativ

eRe

pJu

n–A

ugx

Asc

lepi

adac

eae

Asc

lepi

as la

tifo

liabr

oadl

eaf

milk

wee

dN

ativ

eH

ist

Jun–

Aug

xx

Asc

lepi

adac

eae

Asc

lepi

as s

ubve

rtic

illat

aw

horl

ed m

ilkw

eed

Nat

ive

Pres

May

–Sep

xD

istu

rbed

are

as

Bora

gina

ceae

Cryp

tant

ha c

onfe

rtifl

ora

basi

n ye

llow

cry

ptan

thN

ativ

ePr

esA

pr–J

unx

x

Bora

gina

ceae

Cryp

tant

ha fl

ava

plat

eau

yello

w c

rypt

anth

Nat

ive

Rep

Apr

–Jun

x

Bora

gina

ceae

Cryp

tant

ha f

ulvo

cane

scen

spl

atea

u cr

ypta

nth

Nat

ive

His

tM

ay–J

unx

Cact

acea

eEc

hino

cere

us e

ngel

man

nii v

ar.

vari

egat

usG

len

Cany

on h

edge

hog

cact

usN

ativ

eRe

pM

ar–J

ulx

Cact

acea

eEc

hino

cere

us t

rigl

ochi

diat

us

var.

mel

anac

anth

uscl

aret

cup

Nat

ive

Pres

Apr

–Jun

x

Cact

acea

eEc

hino

cere

us t

rigl

ochi

diat

us

var.

moj

aven

sis

Moh

ave

clar

etcu

pN

ativ

eH

ist

Apr

–Jun

xx

Cact

acea

eO

punt

ia e

rina

cea

var.

aure

aPi

pe S

prin

g pr

ickl

ypea

rN

ativ

ePr

esM

ay–J

unx

x

Cact

acea

eO

punt

ia e

rina

cea

var.

utah

ensi

sU

tah

pric

klyp

ear

Nat

ive

Pres

May

–Jun

xx

Cact

acea

eO

punt

ia f

ragi

lisbr

ittl

e pr

ickl

y-pe

arN

ativ

eRe

pJu

n–Ju

lx

x

Cact

acea

eO

punt

ia p

haea

cant

ha v

ar.

maj

orla

rge

pric

klyp

ear

Nat

ive

Pres

Apr

–Jul

xx

Cact

acea

eO

punt

ia p

inka

vae

Pink

ava’

s pr

ickl

ypea

rN

ativ

ePr

esM

ay–J

unx

Cact

acea

eSc

lero

cact

us w

hipp

lei v

ar.

rose

ussm

all-fl

ow

er fi

shho

ok

cact

usN

ativ

ePr

esA

pr–M

ayx

x

Appendices 51

App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Pere

nnia

l For

bs, c

ont.

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia c

ampe

stri

s va

r. sc

oule

rian

asa

nd w

orm

woo

dN

ativ

eRe

pJu

l–Se

px

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia d

racu

ncul

us s

sp.

glau

cata

rrag

onN

ativ

eRe

pJu

l–O

ctx

Com

posi

tae

(Ast

erac

eae)

Art

emis

ia lu

dovi

cian

a va

r. al

bula

Loui

sian

a w

orm

woo

dN

ativ

ePr

esJu

l–O

ctx

x

Com

posi

tae

(Ast

erac

eae)

Ast

er h

espe

rius

Sisk

iyou

ast

erN

ativ

ePr

esJu

l–Se

px

Com

posi

tae

(Ast

erac

eae)

Chae

topa

ppa

eric

oide

sro

se-h

eath

Nat

ive

Pres

May

–Sep

xx

Com

posi

tae

(Ast

erac

eae)

Erig

eron

div

erge

ns v

ar.

dive

rgen

ssp

read

ing

dais

yN

ativ

ePr

esM

ay–S

epx

Com

posi

tae

(Ast

erac

eae)

Erig

eron

pum

ilus

var.

conc

innu

sN

avaj

o fl

eaba

neN

ativ

eRe

pM

ay–A

ugx

Com

posi

tae

(Ast

erac

eae)

Gai

llard

ia p

inna

tifi

daH

opi b

lank

etfl

ower

Nat

ive

Pres

Apr

–Aug

xx

Com

posi

tae

(Ast

erac

eae)

Gri

ndel

ia s

quar

rosa

var

. se

rrul

ata

curl

ycup

gum

wee

dN

ativ

eRe

pJu

l–Se

pD

istu

rbed

are

as

Com

posi

tae

(Ast

erac

eae)

Hym

enop

appu

s fi

lifol

ius

var.

cine

reus

com

mon

hya

lineh

erb

Nat

ive

Pres

May

–Oct

x

Com

posi

tae

(Ast

erac

eae)

Iva

axill

aris

po

vert

ywee

dN

ativ

ePr

esM

ay–S

epx

Com

posi

tae

(Ast

erac

eae)

Mac

haer

anth

era

cane

scen

s va

r. ca

nesc

ens

hoar

y as

ter

Nat

ive

Pres

May

–Sep

xx

x

Com

posi

tae

(Ast

erac

eae)

Ono

pord

um a

cant

hium

Scot

ch t

hist

leIn

tro

Rep

Jun–

Sep

x

Com

posi

tae

(Ast

erac

eae)

Psilo

stro

phe

spar

sifl

ora

gree

nste

m p

aper

fl ow

erN

ativ

eH

ist

May

–Sep

Com

posi

tae

(Ast

erac

eae)

Sene

cio

doug

lasi

i var

. lo

ngilo

bus

Dou

glas

’ gro

unds

elN

ativ

ePr

esM

ay–O

ctx

Com

posi

tae

(Ast

erac

eae)

Solid

ago

sp.

gold

enro

dN

ativ

eRe

pJu

n–O

ctx

Com

posi

tae

(Ast

erac

eae)

Step

hano

mer

ia p

auci

fl or

abr

own-

plum

e w

ire

lett

uce

Nat

ive

Pres

May

–Oct

xx

Com

posi

tae

(Ast

erac

eae)

Thel

espe

rma

subn

udum

var

. su

bnud

umsc

apos

e gr

eent

hrea

dN

ativ

ePr

esM

ay–A

ugx

x

Com

posi

tae

(Ast

erac

eae)

Tow

nsen

dia

inca

naho

ary

tow

nsen

dia

Nat

ive

Pres

Apr

–Jul

xx

Com

posi

tae

(Ast

erac

eae)

Trag

opog

on d

ubiu

sye

llow

sal

sify

Intr

oPr

esM

ay–J

ulx

Conv

olvu

lace

aeCo

nvol

vulu

s ar

vens

isfi

eld

bind

wee

dIn

tro

His

tJu

n–A

ugD

istu

rbed

are

as

Cruc

ifer

ae (B

rass

icac

eae)

Ara

bis

pere

nnan

s va

r. pe

renn

ans

pere

nnia

l roc

kcre

ssN

ativ

eRe

pM

ar–J

ulx


App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Pere

nnia

l For

bs, c

ont.

Cruc

ifer

ae (B

rass

icac

eae)

Lepi

dium

mon

tanu

m v

ar.

jone

sii

Jone

s’ p

eppe

rwor

tN

ativ

ePr

esA

pr–S

epx

x

Cruc

ifer

ae (B

rass

icac

eae)

Lepi

dium

mon

tanu

m v

ar.

stel

lae

Stel

la’s

pepp

erw

ort

Nat

ive

His

tM

ay–J

unx

Cruc

ifer

ae (B

rass

icac

eae)

Nas

turt

ium

offi

cin

ale

wat

ercr

ess

Intr

oPr

esM

ay–S

epx

Cruc

ifer

ae (B

rass

icac

eae)

Stan

leya

pin

nata

prin

ce’s-

plum

eN

ativ

ePr

esA

pr–S

epx

x

Cucu

rbit

acea

eCu

curb

ita

foet

idis

sim

abu

ffal

o go

urd

Nat

ive

Pres

May

–Aug

xx

Dis

turb

ed a

reas

Euph

orbi

acea

eCh

amae

syce

alb

omar

gina

tara

ttle

snak

e-w

eed

Nat

ive

Rep

Apr

–Sep

x

Euph

orbi

acea

eCh

amae

syce

fen

dler

iFe

ndle

r’s s

purg

eN

ativ

ePr

esM

ay–S

epx

x

Fum

aria

ceae

Cory

dalis

aur

ea v

ar.

occi

dent

alis

gold

en c

oryd

alis

Nat

ive

His

tx

x

Gen

tian

acea

eSw

erti

a al

bom

argi

nata

whi

te-m

argi

ned

swer

tia

Nat

ive

His

tM

ay–J

ulx

x

Gen

tian

acea

eSw

erti

a ut

ahen

sis

Uta

h sw

erti

aN

ativ

eRe

pJu

n–Ju

lx

x

Irid

acea

eSi

syri

nchi

um d

emis

sum

blue

-eye

d gr

ass

Nat

ive

Rep

Apr

–Aug

x

Labi

atae

(Lam

iace

ae)

Mar

rubi

um v

ulga

reco

mm

on h

oreh

ound

Intr

oPr

esM

ay–O

ctx

xD

istu

rbed

are

as

Labi

atae

(Lam

iace

ae)

Men

tha

spic

ata

spea

rmin

tIn

tro

Pres

Jun–

Aug

x

Legu

min

osae

(Fab

acea

e)A

stra

galu

s am

phio

xys

var.

vesp

erti

nus

Shel

don’

s m

ilkve

tch

Nat

ive

His

tM

ar–J

unx

x

Legu

min

osae

(Fab

acea

e)A

stra

galu

s ce

ram

icus

var

. ce

ram

icus

pain

ted

milk

vetc

hN

ativ

ePr

esA

pr–J

unx

Legu

min

osae

(Fab

acea

e)A

stra

galu

s pr

aelo

ngus

var

. pr

aelo

ngus

stin

king

milk

vetc

hN

ativ

eRe

pA

pr–J

ulx

x

Legu

min

osae

(Fab

acea

e)G

lycy

rrhi

za le

pido

ta

Nut

tall’

s lic

oric

eN

ativ

eRe

pM

ay–J

ulx

Legu

min

osae

(Fab

acea

e)M

edic

ago

sati

vaal

falf

aIn

tro

Rep

May

–Sep

Dis

turb

ed a

reas

Legu

min

osae

(Fab

acea

e)M

elilo

tus

offi

cina

lisye

llow

sw

eetc

love

rIn

tro

Rep

May

–Oct

xD

istu

rbed

are

as

Legu

min

osae

(Fab

acea

e)Pe

teri

a th

omps

onia

eTh

omps

on’s

pete

ria

Nat

ive

Pres

Apr

–Jun

x

Lilia

ceae

(The

mid

acea

e)A

ndro

step

hium

bre

vifl

orum

pink

fun

nel-l

ilyN

ativ

eH

ist

Apr

–May

Lilia

ceae

(C

aloc

hort

acea

e)Ca

loch

ortu

s fl

exuo

sus

sinu

ous

mar

ipos

aN

ativ

ePr

esA

pr–J

unx

Loas

acea

eM

entz

elia

inte

gra

Vir

gin

stic

klea

fN

ativ

eH

ist

Apr

–Oct

x

Loas

acea

eM

entz

elia

mul

tifl

ora

dese

rt s

tick

leaf

Nat

ive

Pres

Apr

–Sep

x

Mal

vace

aeA

ltha

ea r

osea

holly

hock

Intr

oRe

pJu

n–O

ctCu

ltiv

ated

Appendices 53

App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Pere

nnia

l For

bs, c

ont.

Mal

vace

aeSp

haer

alce

a am

bigu

a va

r. am

bigu

aM

ohav

e gl

obem

allo

wN

ativ

ePr

esM

ar–S

epx

x

Mal

vace

aeSp

haer

alce

a pa

rvif

olia

smal

l-lea

f gl

obem

allo

wN

ativ

eH

ist

Mar

–Oct

xx

Nyc

tagi

nace

aeA

bron

ia f

ragr

ans

frag

rant

san

d-ve

rben

aN

ativ

ePr

esA

pr–S

ep

Nyc

tagi

nace

aeM

irab

ilis

mul

tifl

ora

sh

owy

four

-o’c

lock

Nat

ive

Pres

Apr

–Jul

x

Ole

acea

eM

enod

ora

scab

raro

ugh

men

odor

aN

ativ

ePr

esA

pr–A

ugx

Ona

grac

eae

Epilo

bium

cili

atum

nort

hern

will

ow-h

erb

Nat

ive

Rep

Jun–

Aug

x

Ona

grac

eae

Gau

ra c

occi

nea

scar

let

gaur

aN

ativ

eH

ist

May

–Jul

xx

Ona

grac

eae

Oen

othe

ra c

aesp

itos

a va

r. na

vajo

ensi

sPa

ria

even

ing

prim

rose

Nat

ive

Pres

Apr

–Jun

x

Ona

grac

eae

Oen

othe

ra c

alif

orni

caCa

lifor

nia

even

ing

prim

rose

Nat

ive

Pres

May

–Jul

x

Ona

grac

eae

Oen

othe

ra p

allid

a va

r. pa

llida

pale

eve

ning

pri

mro

seN

ativ

ePr

esM

ay–J

ulx

Oro

banc

hace

aeO

roba

nche

ludo

vici

ana

Coop

er’s

broo

mra

peN

ativ

ePr

esM

ay–J

ulx

Plan

tagi

nace

aePl

anta

go la

nceo

lata

Engl

ish

plan

tain

Intr

oRe

pM

ay–A

ugD

istu

rbed

are

as

Plan

tagi

nace

aePl

anta

go m

ajor

com

mon

pla

ntai

nIn

tro

Pres

May

–Aug

xD

istu

rbed

are

as

Pole

mon

iace

aeIp

omop

sis

long

ifl or

alo

ng-fl

ow

ered

gili

aN

ativ

ePr

esM

ay–A

ugx

Poly

gala

ceae

Poly

gala

sub

spin

osa

cush

ion

milk

wor

tN

ativ

eH

ist

May

–Jun

x

Poly

gona

ceae

Erio

gonu

m in

fl at

um v

ar.

infl

atum

dese

rt t

rum

pet

Nat

ive

Pres

Mar

–Oct

xx

Poly

gona

ceae

Erio

gonu

m m

icro

thec

um v

ar.

sim

pson

iisl

ende

r w

ild b

uckw

heat

Nat

ive

Pres

Jun–

Oct

x

Poly

gona

ceae

Poly

gonu

m a

mph

ibiu

mw

ater

sm

artw

eed

Nat

ive

Rep

Jun–

Sep

x

Poly

gona

ceae

Rum

ex c

risp

uscu

rly

dock

Intr

oRe

pM

ay–S

epx

Poly

gona

ceae

Rum

ex h

ymen

osep

alus

cana

igre

doc

kN

ativ

ePr

esA

pr–J

unx

Ranu

ncul

acea

e (H

elle

bora

ceae

)D

elph

iniu

m a

nder

soni

i var

. sc

apos

umpa

le la

rksp

urN

ativ

eH

ist

Apr

–Jun

xx

Rosa

ceae

Pote

ntill

a gr

acili

s va

r. el

mer

ico

mb-

leaf

cin

quef

oil

Nat

ive

His

tJu

n–A

ugx

Scro

phul

aria

ceae

Cast

illej

a ch

rom

osa

dese

rt p

aint

brus

hN

ativ

ePr

esA

pr–J

unx

x

Scro

phul

aria

ceae

Cast

illej

a lin

ariif

olia

Wyo

min

g pa

intb

rush

Nat

ive

Pres

Jul–

Aug

xx

Scro

phul

aria

ceae

Pens

tem

on la

evis

smoo

th p

enst

emon

Nat

ive

Rep

May

–Jun

x

Scro

phul

aria

ceae

Pens

tem

on p

alm

eri v

ar.

egla

ndul

osus

Palm

er’s

pens

tem

onN

ativ

ePr

esM

ay–A

ugx

x


App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Pere

nnia

l For

bs, c

ont.

Scro

phul

aria

ceae

Pens

tem

on u

tahe

nsis

Uta

h pe

nste

mon

Nat

ive

His

tA

pr–J

unx

Sola

nace

aeCh

amae

sara

cha

coro

nopu

sfa

lse

nigh

tsha

deN

ativ

ePr

esA

pr–J

ulx

Sola

nace

aePh

ysal

is c

rass

ifol

iath

ick-

leav

ed g

roun

d-ch

erry

Nat

ive

Rep

Mar

–May

x

Sola

nace

aePh

ysal

is h

eder

ifol

ia v

ar.

palm

eri

Palm

er’s

grou

nd-c

herr

yN

ativ

eRe

pM

ay–O

ctx

Um

belli

fera

e (A

piac

eae)

Beru

la e

rect

a va

r. in

cisa

cutl

eaf

wat

er-p

arsn

ipN

ativ

ePr

esJu

n–Se

px

Um

belli

fera

e (A

piac

eae)

Cym

opte

rus

mul

tine

rvat

uspu

rple

-ner

ved

spri

ng-

pars

ley

Nat

ive

Pres

Mar

–Apr

x

Ver

bena

ceae

Ver

bena

bra

ctea

tapr

ostr

ate

verv

ain

Nat

ive

Pres

May

–Sep

xD

istu

rbed

are

as

Vis

cace

aePh

orad

endr

on ju

nipe

rinu

mju

nipe

r m

istl

etoe

Nat

ive

Pres

Jul–

Sep

x

Ann

ual F

orbs

Am

aran

thac

eae

Am

aran

thus

blit

oide

spr

ostr

ate

pigw

eed

Nat

ive

Pres

Jun–

Sep

Dis

turb

ed a

reas

Am

aran

thac

eae

Am

aran

thus

hyp

ocho

ndri

acus

grai

n am

aran

thIn

tro

Pres

Jun–

Oct

Cult

ivat

ed

Bora

gina

ceae

Cryp

tant

ha c

ircu

msc

issa

cush

ion

cryp

tant

hN

ativ

eH

ist

Apr

–Jul

x

Bora

gina

ceae

Cryp

tant

ha c

rass

isep

ala

var.

elac

hant

hath

ick-

sepa

led

cryp

tant

hN

ativ

eH

ist

Apr

–Jun

xx

Bora

gina

ceae

Cryp

tant

ha g

raci

lissl

ende

r cr

ypta

nth

Nat

ive

His

tA

pr–J

ulx

x

Bora

gina

ceae

Cryp

tant

ha m

icra

ntha

redr

oot

cryp

tant

hN

ativ

eH

ist

Mar

–May

x

Bora

gina

ceae

Cryp

tant

ha p

tero

cary

a va

r. cy

clop

tera

win

gnut

cry

ptan

thN

ativ

eH

ist

Apr

–Jul

x

Bora

gina

ceae

Cryp

tant

ha p

tero

cary

a va

r. pt

eroc

arya

win

gnut

cry

ptan

thN

ativ

ePr

esA

pr–J

ulx

x

Bora

gina

ceae

Hel

iotr

opiu

m c

onvo

lvul

aceu

mbi

ndw

eed

helio

trop

eN

ativ

eH

ist

May

–Sep

x

Bora

gina

ceae

Lapp

ula

occi

dent

alis

var

. cu

pula

tacu

psee

d st

icks

eed

Nat

ive

Pres

Apr

–Jun

Dis

turb

ed a

reas

Bora

gina

ceae

Plag

iobo

thry

s jo

nesi

iJo

nes’

pop

corn

-fl o

wer

Nat

ive

Pres

Apr

–May

x

Capp

arac

eae

(Cle

omac

eae)

Cleo

me

lute

aye

llow

bee

plan

tN

ativ

ePr

esA

pr–S

epx

Dis

turb

ed a

reas

Chen

opod

iace

aeA

trip

lex

arge

ntea

var

. ar

gent

easi

lver

ora

chN

ativ

ePr

esJu

n–Se

pD

istu

rbed

are

as

Chen

opod

iace

aeCh

enop

odiu

m a

lbum

var

. al

bum

lam

bsqu

arte

rIn

tro

Pres

Jun–

Sep

Dis

turb

ed a

reas

Appendices 55

App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Ann

ual F

orbs

, con

t.

Chen

opod

iace

aeCh

enop

odiu

m f

rem

onti

i var

. fr

emon

tii

Frem

ont’s

goo

sefo

otN

ativ

ePr

esA

ug–O

ctx

Chen

opod

iace

aeCh

enop

odiu

m le

ptop

hyllu

mna

rrow

leaf

goo

sefo

otN

ativ

eRe

pJu

l–Se

px

Chen

opod

iace

aeSa

lsol

a tr

agus

Russ

ian

this

tle

Intr

oPr

esJu

n–Se

px

xD

istu

rbed

are

as

Chen

opod

iace

aeSu

aeda

tor

reya

na v

ar.

torr

eyan

aTo

rrey

’s se

epw

eed

Nat

ive

Pres

Jun–

Sep

x

Com

posi

tae

(Ast

erac

eae)

Am

bros

ia a

cant

hica

rpa

bur

ragw

eed

Nat

ive

Pres

Jul–

Oct

x

Com

posi

tae

(Ast

erac

eae)

Am

bros

ia a

rtem

isiif

olia

com

mon

rag

wee

dN

ativ

ePr

esJu

l–O

ctx

Dis

turb

ed a

reas

Com

posi

tae

(Ast

erac

eae)

Chae

nact

is c

arph

oclin

iaG

ray’

s ch

aena

ctis

Nat

ive

Pres

Apr

–Jun

x

Com

posi

tae

(Ast

erac

eae)

Cony

za c

anad

ensi

s va

r. gl

abra

taCa

nadi

an h

orse

wee

dN

ativ

eRe

pJu

l–Se

px

Com

posi

tae

(Ast

erac

eae)

Gai

llard

ia a

rizo

nica

Ari

zona

bla

nket

fl ow

erN

ativ

eRe

pA

pr–M

ayx

Com

posi

tae

(Ast

erac

eae)

Hel

iant

hus

annu

us s

sp.

lent

icul

aris

com

mon

sun

fl ow

erN

ativ

ePr

esJu

n–Se

px

Com

posi

tae

(Ast

erac

eae)

Lact

uca

serr

iola

pric

kly

lett

uce

Intr

oPr

esJu

l–Se

px

Dis

turb

ed a

reas

Com

posi

tae

(Ast

erac

eae)

Layi

a gl

andu

losa

tidy

tips

Nat

ive

Pres

Apr

–Jul

xx

Com

posi

tae

(Ast

erac

eae)

Mac

haer

anth

era

tana

ceti

folia

tans

ylea

f as

ter

Nat

ive

Pres

May

–Sep

xD

istu

rbed

are

as

Com

posi

tae

(Ast

erac

eae)

Mal

acot

hrix

gla

brat

afi

lifor

m d

eser

t-da

ndel

ion

Nat

ive

Pres

Apr

–Jul

x

Com

posi

tae

(Ast

erac

eae)

Mal

acot

hrix

son

choi

des

sow

this

tle

dese

rt-

dand

elio

nN

ativ

eH

ist

Apr

–Jun

x

Com

posi

tae

(Ast

erac

eae)

Sonc

hus

aspe

rsp

iny-

leaf

sow

-thi

stle

Intr

oPr

esJu

l–O

ctx

Com

posi

tae

(Ast

erac

eae)

Sonc

hus

oler

aceu

sco

mm

on s

owth

istl

eIn

tro

Rep

Jul–

Oct

x

Com

posi

tae

(Ast

erac

eae)

Step

hano

mer

ia e

xigu

aw

hite

-plu

me

wir

e le

ttuc

eN

ativ

ePr

esM

ay–S

ep

Com

posi

tae

(Ast

erac

eae)

Ver

besi

na e

ncel

ioid

es v

ar.

exau

ricu

lata

gold

en c

row

nbea

rdN

ativ

ePr

esM

ay–O

ctx

Dis

turb

ed a

reas

Com

posi

tae

(Ast

erac

eae)

Xan

thiu

m s

trum

ariu

m v

ar.

cana

dens

ero

ugh

cock

lebu

rN

ativ

eRe

pA

pr–O

ctx

Dis

turb

ed a

reas

Cruc

ifer

ae (B

rass

icac

eae)

Caps

ella

bur

sa-p

asto

ris

shep

herd

’s-pu

rse

Intr

oH

ist

Apr

–Sep

Dis

turb

ed a

reas

Cruc

ifer

ae (B

rass

icac

eae)

Des

cura

inia

pin

nata

var

. gl

abra

wes

tern

tan

sym

usta

rdN

ativ

eH

ist

Mar

–Aug

xx

Cruc

ifer

ae (B

rass

icac

eae)

Des

cura

inia

pin

nata

var

. in

term

edia

wes

tern

tan

sym

usta

rdN

ativ

eH

ist

Mar

–Aug

xx


App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Ann

ual F

orbs

, con

t.

Cruc

ifer

ae (B

rass

icac

eae)

Des

cura

inia

pin

nata

var

. os

mia

rum

wes

tern

tan

sym

usta

rdN

ativ

ePr

esM

ar–A

ugx

x

Cruc

ifer

ae (B

rass

icac

eae)

Des

cura

inia

sop

hia

fl ix

wee

dIn

tro

Pres

Mar

–Aug

xD

istu

rbed

are

as

Cruc

ifer

ae (B

rass

icac

eae)

Dit

hyre

a w

isliz

eni

spec

tacl

epod

Nat

ive

Pres

Apr

–Sep

x

Cruc

ifer

ae (B

rass

icac

eae)

Erys

imum

rep

andu

msp

read

ing

wal

lfl ow

erIn

tro

His

tA

pr–J

un

Cruc

ifer

ae (B

rass

icac

eae)

Lepi

dium

den

sifl

orum

var

. ra

mos

umpr

airi

e pe

pper

wor

tN

ativ

eH

ist

May

–Jul

x

Cruc

ifer

ae (B

rass

icac

eae)

Lepi

dium

per

folia

tum

clas

ping

pep

perw

ort

Intr

oH

ist

May

–Jul

Dis

turb

ed a

reas

Cruc

ifer

ae (B

rass

icac

eae)

Sisy

mbr

ium

alt

issi

mum

tum

ble

mus

tard

Intr

oPr

esM

ay–A

ugx

xD

istu

rbed

are

as

Cruc

ifer

ae (B

rass

icac

eae)

Stre

ptan

thel

la lo

ngir

ostr

islo

ng-b

eak

fi dd

lem

usta

rdN

ativ

eH

ist

Mar

–Jun

xx

Cype

race

aeCy

peru

s sp

.fl

at-s

edge

Intr

oRe

pJu

n–Se

pD

istu

rbed

are

as

Euph

orbi

acea

eCh

amae

syce

gly

ptos

perm

ari

dge-

seed

ed s

purg

eN

ativ

ePr

esJu

n–A

ugx

Dis

turb

ed a

reas

Euph

orbi

acea

eCh

amae

syce

pro

stra

tapr

ostr

ate

spur

geIn

tro

Pres

May

–Sep

Dis

turb

ed a

reas

Ger

ania

ceae

Erod

ium

cic

utar

ium

stor

k’s-

bill

Intr

oPr

esFe

b–O

ctx

xD

istu

rbed

are

as

Hyd

roph

ylla

ceae

Phac

elia

cre

nula

ta v

ar.

corr

ug

ata

corr

ugat

e ph

acel

iaN

ativ

eH

ist

May

–Jun

x

Hyd

roph

ylla

ceae

Phac

elia

cre

nula

ta v

ar.

cren

ulat

acr

enul

ate

phac

elia

Nat

ive

Pres

May

–Jun

x

Hyd

roph

ylla

ceae

Phac

elia

ives

iana

Ives

’ pha

celia

Nat

ive

Pres

Apr

–Jun

xx

Hyd

roph

ylla

ceae

Phac

elia

pul

chel

la v

ar.

good

ding

iiG

oodd

ing’

s pr

etty

ph

acel

iaN

ativ

eH

ist

Apr

–May

x

Legu

min

osae

(Fab

acea

e)Lu

pinu

s pu

sillu

s va

r. pu

sillu

sdw

arf

lupi

neN

ativ

eH

ist

Apr

–Jul

x

Legu

min

osae

(Fab

acea

e)M

elilo

tus

alba

whi

te s

wee

tclo

ver

Intr

oRe

pM

ay–O

ctx

xx

Dis

turb

ed a

reas

Lina

ceae

Linu

m a

rist

atum

broo

mfl

axN

ativ

ePr

esA

pr–J

unx

Loas

acea

eM

entz

elia

alb

icau

lisw

hite

-ste

m b

lazi

ngst

arN

ativ

ePr

esA

pr–A

ugx

Mal

vace

aeM

alva

neg

lect

aco

mm

on m

allo

wIn

tro

Pres

Apr

–Oct

Dis

turb

ed a

reas

Ona

grac

eae

Cam

isso

nia

cham

aene

rioi

des

slen

der-

pod

cam

isso

nia

Nat

ive

Pres

Apr

–Jun

x

Ona

grac

eae

Cam

isso

nia

parr

yiPa

rry’

s ca

mis

soni

aN

ativ

eH

ist

May

–Jun

x

Ona

grac

eae

Cam

isso

nia

parv

ula

tiny

cam

isso

nia

Nat

ive

His

tM

ay–J

unx

x

Ona

grac

eae

Gau

ra p

arvi

fl or

asm

all-fl

ow

ered

gau

raN

ativ

ePr

esM

ay–A

ugD

istu

rbed

are

as

Ona

grac

eae

Oen

othe

ra a

lbic

aulis

whi

te-s

tem

eve

ning

pr

imro

seN

ativ

eH

ist

Mar

–Jun

x

Appendices 57

App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Ann

ual F

orbs

, con

t.

Ona

grac

eae

Oen

othe

ra p

rim

iver

is v

ar.

bufo

nus

earl

y ev

enin

g pr

imro

seN

ativ

eRe

pM

ar–M

ayx

Papa

vera

ceae

Esch

scho

lzia

min

utifl

ora

pygm

y go

lden

-pop

pyN

ativ

ePr

esFa

b–M

ayx

Sand

y so

il

Pole

mon

iace

aeEr

iast

rum

ere

mic

umM

ohav

e er

iast

rum

Nat

ive

His

tA

pr–J

un

Pole

mon

iace

aeG

ilia

inco

nspi

cua

shy

gilia

Nat

ive

Pres

Apr

–Jul

xx

Pole

mon

iace

aeG

ilia

lept

omer

ia v

ar.

lept

omer

iaco

mm

on g

ilia

Nat

ive

His

tM

ay–J

unx

x

Pole

mon

iace

aeIp

omop

sis

poly

clad

onsp

read

ing

gilia

Nat

ive

His

tA

pr–J

unx

Pole

mon

iace

aeLa

nglo

isia

set

osis

sim

aM

ohav

e la

nglo

isia

Nat

ive

His

tM

ar–M

ayx

Poly

gona

ceae

Cent

rost

egia

thu

rber

iTh

urbe

r’s s

pine

fl ow

erN

ativ

eH

ist

Mar

–Jul

x

Poly

gona

ceae

Erio

gonu

m c

ernu

um v

ar.

cern

uum

nodd

ing

wild

buc

kwhe

atN

ativ

eH

ist

Apr

–Oct

xx

Poly

gona

ceae

Erio

gonu

m d

efl e

xum

skel

eton

wee

d w

ild

buck

whe

atN

ativ

eH

ist

Apr

–Oct

x

Poly

gona

ceae

Erio

gonu

m p

alm

eria

num

Palm

er’s

wild

buc

kwhe

atN

ativ

ePr

esM

ar–O

ctx

Poly

gona

ceae

Poly

gonu

m a

vicu

lare

yard

kno

twee

dIn

tro

Pres

May

–Oct

Dis

turb

ed a

reas

Port

ulac

acea

ePo

rtul

aca

oler

acea

com

mon

pur

slan

eIn

tro

Pres

Jun–

Sep

xD

istu

rbed

are

as

Scro

phul

aria

ceae

Cord

ylan

thus

par

vifl

orus

smal

lfl ow

er b

ird’

s-be

akN

ativ

ePr

esA

ug–S

epx

Sola

nace

aeD

atur

a w

righ

tii

ange

l’s t

rum

pet

Nat

ive

Pres

Apr

–Oct

xD

istu

rbed

are

as

Sola

nace

aeN

icot

iana

tri

gono

phyl

lade

sert

tob

acco

Nat

ive

His

tM

ar–O

ctx

Sola

nace

aeSo

lanu

m s

arra

choi

des

grou

nd-c

herr

y ni

ghts

hade

Intr

oRe

pM

ay–O

ctD

istu

rbed

are

as

Zygo

phyl

lace

aeTr

ibul

us t

erre

stri

spu

nctu

revi

neIn

tro

Pres

May

–Sep

Dis

turb

ed a

reas

Pere

nnia

l Gra

min

oids

Cype

race

aeCa

rex

nebr

asce

nsis

Neb

rask

a se

dge

Nat

ive

Pres

x

Cype

race

aeCa

rex

prae

grac

ilis

blac

kcre

eper

sed

geN

ativ

ePr

esM

ay–A

ugx

Cype

race

aeEl

eoch

aris

par

ishi

iPa

rish

’s sp

iker

ush

Nat

ive

Pres

Apr

–Sep

x

Cype

race

aeSc

irpu

s pu

ngen

s va

r. lo

ngis

pica

tus

thre

e-sq

uare

bul

rush

Nat

ive

Pres

May

–Aug

x

Gra

min

eae

(Poa

ceae

)A

grop

yron

cri

stat

umcr

este

d w

heat

gras

sIn

tro

Rep

Jun–

Aug

xCu

ltiv

ated

Gra

min

eae

(Poa

ceae

)A

gros

tis

stol

onif

era

redt

opIn

tro

Pres

Jun–

Sep

x

Gra

min

eae

(Poa

ceae

)A

rist

ida

purp

urea

purp

le t

hree

-aw

nN

ativ

ePr

esM

ar–S

epx

xD

istu

rbed

are

as

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

curt

ipen

dula

side

oats

gra

ma

Nat

ive

Rep

Jul–

Sep

x


App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Pere

nnia

l Gra

min

oids

, con

t.

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

erio

poda

blac

k gr

ama

Nat

ive

Pres

Jun–

Aug

x

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

grac

ilis

blue

gra

ma

Nat

ive

Pres

Jul–

Sep

xx

Gra

min

eae

(Poa

ceae

)Br

omus

iner

mis

var

. ine

rmis

smoo

th b

rom

eIn

tro

Rep

Jun–

Aug

x

Gra

min

eae

(Poa

ceae

)D

isti

chlis

spi

cata

de

sert

sal

tgra

ssN

ativ

eRe

pJu

n–Se

px

Gra

min

eae

(Poa

ceae

)El

ymus

elo

ngat

us

tall

whe

atgr

ass

Intr

oRe

pJu

n–A

ugx

Gra

min

eae

(Poa

ceae

)El

ymus

ely

moi

des

squi

rrel

tail

Nat

ive

Rep

May

–Aug

xx

Gra

min

eae

(Poa

ceae

)El

ymus

his

pidu

s in

term

edia

te w

heat

gras

sIn

tro

Rep

Jun–

Aug

Dis

turb

ed a

reas

Gra

min

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ceae

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ymus

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ithi

iw

este

rn w

heat

gras

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eRe

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min

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(Poa

ceae

)El

ymus

tra

chyc

aulu

s sl

ende

r w

heat

gras

sN

ativ

eH

ist

Jun–

Aug

x

Gra

min

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(Poa

ceae

)Er

ione

uron

pul

chel

lum

fl uf

fgra

ssN

ativ

eRe

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x

Gra

min

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ceae

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stuc

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undi

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ata

ll fe

scue

Intr

oRe

pM

ay–J

ulx

Gra

min

eae

(Poa

ceae

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ilari

a ja

mes

iiga

lleta

Nat

ive

Pres

May

–Aug

xx

Gra

min

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(Poa

ceae

)H

orde

um ju

batu

mfo

xtai

l bar

ley

Nat

ive

Rep

Jun–

Aug

x

Gra

min

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(Poa

ceae

)M

uhle

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gia

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liasc

ratc

hgra

ssN

ativ

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l–A

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min

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nber

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port

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lyN

ativ

ePr

esJu

l–O

ctx

Gra

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)Po

a fe

ndle

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utto

ngra

ssN

ativ

eRe

pM

ay–A

ugx

x

Gra

min

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a pr

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sis

Ken

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ass

Intr

oRe

pM

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ceae

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iver

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sw

ater

pol

ypog

onIn

tro

Rep

Jul–

Aug

xD

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rbed

are

as

Gra

min

eae

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ceae

)Sp

orob

olus

air

oide

s va

r. ai

roid

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kali

saca

ton

Nat

ive

Rep

Jun–

Aug

xx

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min

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ceae

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orob

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trac

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spik

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opse

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x

Gra

min

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min

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esa

drop

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Nat

ive

Pres

Jun–

Sep

xx

Gra

min

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ceae

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ipa

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min

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ceae

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Indi

an r

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rass

Nat

ive

Pres

May

–Jul

xx

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ceae

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ipa

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rt n

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tor

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Torr

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Nat

ive

Pres

Jun–

Aug

x

Appendices 59

App

endi

x D

. Con

fi rm

ed, h

isto

rica

l, an

d re

port

ed t

axa

of P

ipe

Spri

ng N

atio

nal M

onum

ent

orga

nize

d by

life

for

m a

nd c

omm

on n

ame

wit

h ec

olog

ical

not

es, c

ont.

Fam

ilySp

ecie

s na

me

Com

mon

nam

eN

ativ

ity

Park

st

atus

Flow

er

tim

eD

esSh

Pin

Jun

Wet

Com

men

ts

Ann

ual G

ram

inoi

ds

Gra

min

eae

(Poa

ceae

)Bo

utel

oua

barb

ata

sixw

eeks

gra

ma

Nat

ive

Pres

Aug

–Oct

x

Gra

min

eae

(Poa

ceae

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omus

dia

ndru

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bro

me

Intr

oPr

esA

pr–J

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as

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omus

rub

ens

red

brom

eIn

tro

Pres

Mar

–Jun

x

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min

eae

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ceae

)Br

omus

tec

toru

mch

eatg

rass

Intr

oPr

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ay–J

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xx

Dis

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ed a

reas

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min

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ceae

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nchr

us lo

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eld

sand

bur

Nat

ive

Rep

Jul–

Sep

xD

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as

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sIn

tro

Rep

Aug

–Sep

Dis

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ed a

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min

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ceae

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agro

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sis

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kgra

ssIn

tro

Rep

Jul–

Sep

Dis

turb

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ceae

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agro

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ican

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an lo

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ass

Intr

oRe

pJu

l–O

ctD

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rbed

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as

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min

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ceae

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pec

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tuft

ed lo

vegr

ass

Nat

ive

Pres

Jul–

Aug

Dis

turb

ed a

reas

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min

eae

(Poa

ceae

)H

orde

um m

urin

umra

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bar

ley

Intr

oPr

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ed a

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min

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ceae

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osa

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e bu

ffal

ogra

ssN

ativ

ePr

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px

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min

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ceae

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lypo

gon

mon

spel

iens

isra

bbit

sfoo

tgra

ssIn

tro

Pres

Jun–

Aug

xD

istu

rbed

are

as

The Department of the Interior protects and manages the nation’s natural resources and cultural heritage; provides scientifi c and other information about those resources; and honors its special responsibilities to American Indians, Alaska Natives, and affi liated Island Communities.

NPS D-222, October 2008

Natural Resource Program Center 1201 Oak Ridge Drive, Suite 150 Fort Collins, Colorado 80525

www.nature.nps.gov

National Park ServiceU.S. Department of the Interior

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