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AN ANNOTATED LIST OF NEARCTIC DONACIINAE (COLEOPTERA: CHRYSOMELIDAE): THE GENERIC CLASSIFICATION AND TYPE SPECIMENS OF THE NEW WORLD SPECIES. BY INGOLF S. ASKEVOLD Department of Entomology, University of Manitoba, Winnipeg, Manitoba, Canada R3T 2N21 INTRODUCTION During systematic revision of Nearctic donaciines, the majority of type specimens were examined. Some lectotype and neotype designations, made where necessary in the context of revisions of genera (Plateumaris Thomson and Neohaemonia Sz6kessy) or solution of specific taxonomic problems, were detailed elsewhere (Askevold 1987a,b, 1988, 1990, 1991). Some taxonomic problems remain despite relatively recent treatment by Marx (1957) of Donacia F. (sensu lato). I indicated earlier that Donacia is repre- sented in North America by 10 species in Donacia (s. str.), and 21 described species in Donacia (Donaciomima) Medvedev, but details were not given then (Askevold 1990). The purpose of the present paper is to detail remaining lectotype designations among Nearctic members of the genus Donacia F., to list the Nearctic members of Donaciinae, and to detail relevant tax- onomic references and information (i.e., synonymies, taxonomic changes since prior authors, and data about type specimens). Schaeffer (1919, 1925) indicated only if he based his species on a type and allotype, but not that there were also many paratypes; paratype information is included insofar as such specimens have been found. Species are listed within genera and subgenera alpha- betically rather than phylogenetically, because phylogenies have been reconstructed only for members of Plateumaris and Neo- haemonia (Askevold 1991, 1988). 1present address: Entomology-Biological Control, Division of Agricultural Sci- ences, Florida A & M University, Tallahassee, FL 32307-2001 Manuscript received June 7, 1991. 165

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AN ANNOTATED LIST OF NEARCTIC DONACIINAE(COLEOPTERA: CHRYSOMELIDAE): THE GENERIC

CLASSIFICATION AND TYPE SPECIMENS OF THE NEWWORLD SPECIES.

BY INGOLF S. ASKEVOLD

Department of Entomology, University of Manitoba, Winnipeg,Manitoba, Canada R3T 2N21

INTRODUCTION

During systematic revision of Nearctic donaciines, the majorityof type specimens were examined. Some lectotype and neotypedesignations, made where necessary in the context of revisions ofgenera (Plateumaris Thomson and Neohaemonia Sz6kessy) orsolution of specific taxonomic problems, were detailed elsewhere(Askevold 1987a,b, 1988, 1990, 1991). Some taxonomic problemsremain despite relatively recent treatment by Marx (1957) ofDonacia F. (sensu lato). I indicated earlier that Donacia is repre-sented in North America by 10 species in Donacia (s. str.), and 21described species in Donacia (Donaciomima) Medvedev, butdetails were not given then (Askevold 1990).

The purpose of the present paper is to detail remaining lectotypedesignations among Nearctic members of the genus Donacia F., tolist the Nearctic members of Donaciinae, and to detail relevant tax-onomic references and information (i.e., synonymies, taxonomicchanges since prior authors, and data about type specimens).Schaeffer (1919, 1925) indicated only if he based his species on atype and allotype, but not that there were also many paratypes;paratype information is included insofar as such specimens havebeen found. Species are listed within genera and subgenera alpha-betically rather than phylogenetically, because phylogenies havebeen reconstructed only for members of Plateumaris and Neo-haemonia (Askevold 1991, 1988).

1present address: Entomology-Biological Control, Division of Agricultural Sci-ences, Florida A & M University, Tallahassee, FL 32307-2001Manuscript received June 7, 1991.

165

166 Psyche [Vol. 98

ANNOTATED CATALOGUE OF NEW WORLD DONACIINAE

The following list includes all described extant New WorldDonaciinae, serving to summarize the many taxonomic changesthat have resulted from revision of North American Donaciinae.Details of historical use of species names is given where substan-tial changes in name use occurred [i.e., changes since Schaeffer(1925), Marx (1957) and Jolivet (1970)]. The taxonomic history ofsome names is complicated, often crossing generic boundaries;some species were synonymized with more than one taxon at once,based on composition of the original type series. I have judged thatwhat is important is current synonymy and whether or not that dif-fers from the most recent uses of names, and therefore have notpried deeply into tracing the varied uses of names since their origi-nal description.

Data of individual labels borne by type specimens, whererecorded, are stated verbatim and are separated by a slash ("/").The museums and private collections in which type specimens ofany kind are known to be preserved are identified by correspond-ing codens:

AMNH. Department of Entomology, American Museum of NaturalHistory, Central Park West at 79th St., New York, NY10024, U.S.A.

APHIS. Animal and Plant Health Inspection Service, USDA,Room 308, 40 South Gay St., Baltimore, MD 21202, U.S.A.

BMNH. Department of Entomology, British Museum of NaturalHistory, Cromwell Road, London SW7 5BD, England.

CARR. J.L. and A.F. Carr, 24 Dalrymple Green, N.W. Calgary,Alta. Canada, T3A 1Y2.

CAS. Department of Entomology, California Academy of Sci-ences, Golden Gate Park, San Francisco, CA 94118, U.S.A.

CDAS. California Department of Food and Agriculture, CaliforniaState collection of Arthropods, Insect Taxonomy Laboratory,1220 N Street, Sacramento, CA 95814, U.S.A.

CLCH. C. Chantal, B.P. 2072, St. Nicolas-est, P.Q., Canada, G0S3L0.

CMP. Section of Entomology, Carnegie Museum of Natural His-tory, 4400 Forbes Ave., Pittsburgh, PA 15213, U.S.A.

1991 Askevold 167

CNC. Biosyntematics Research Centre, Agriculture Canada,Ottawa, Ont., Canada K1A 0C6.

CUIC. Department of Entomology, Comstock Hall, Cornell Uni-versity, Ithaca, NY 14853, U.S.A.

DEFW. Department of Entomology, Fisheries and Wildlife, Uni-versity of Minnesota, 219 Hodson Hall, 1980 Folwell Ave.,St. Paul, MN 55108, U.S.A.

EGRC. E.G. Riley, Department of Entomology, Texas A & M Uni-versity, College Station, TX 77843-2475, U.S.A.

FMNH. Field Museum of Natural History, Roosevelt Road at LakeShore Drive, Chicago, IL 60605, U.S.A.

FSCA. Florida State Collection of Arthropods, Division of PlantIndustry, Florida Department of Agriculture, 1911 S.W. 34thSt., P.O. Box 1269, Gainesville, FL 32602, U.S.A.

INHS. Department of Entomology, Illinois Natural History Survey,Natural Resources Bldg., Urbana, IL 61801, U.S.A.

ISAC. I.S. Askevold, Entomology-Biological Control, Division ofAgricultural Sciences, Florida A & M University, Tallahas-see, FL 32307, U.S.A.

JBWM. J.B. Wallis Museum, Department of Entomology, Univer-sity of Manitoba, Winnipeg, Man. R3T 2N2, Canada.

LEM. Collection Provancher, D6partement de biologie, Universit6Laval, Qu6bec, P.Q., G1K 7P4, Canada.

LSU. Department of Entomology, Louisiana State University,Baton Rouge, LA 70803, U.S.A.

MCZ. Department of Entomology, Museum of Comparative Zool-ogy, Harvard University, Cambridge, MA 02138, U.S.A.

MLU. Section Biowissenschaften, WB Zoologie, Martin-Luther-Universitit Halle-Wittenberg, Domplatz 4, 4010 Halle(Saale) Germany.

NMDC. N.M. Downie, 505 Lingle Terrace, Lafayette, IN 47901,U.S.A.

OSUC. Systematic Entomology Laboratory, Department of Ento-mology, Oregon State University, Corvallis, OR 97331,U.S.A.

PUR. Department of Entomology, Entomology Hall, Purdue Uni-versity, West Lafayette, IN 47907, U.S.A.

ROM. Department of Entomology, Royal Ontario Museum, 100Queen’s Park, Toronto, Ont. M5S 2C6, Canada.

168 Psyche [Vol. 98

SCSU. Department of Biological Sciences, St. Cloud State Univer-sity, St. Cloud, MN 56301, U.S.A.

SMCL. S.M. Clark, Pest Identification Laboratory, West VirginiaDepartment of Agriculture, Charleston, WV 25305, U.S.A.

TAMU. Department of Entomology, Texas A & M University,College Station, TX 77843-2516, U.S.A.

UAE. Department of Entomology, University of Alberta, Edmon-ton, Alta. T6G 2E3, Canada.

UMHF. Universitetets Zoologiska Museum, Entomologiska Avdel-ningen, N. Jirnvigsgatan 13, SF-00100 Helsingfors 10, Fin-land.

UMMA. Museum of Zoology, Insect Division, The University ofMichigan, Ann Arbor, MI 48109, U.S.A.

USNM. National Museum of Natural History, Smithsonian Institu-tion NHB-105, Washington, D.C. 20560, U.S.A.

UVDZ. Department of Zoology, Marsh Life Science Bldg., Uni-versity of Vermont, Burlington, VT 05401, U.S.A.

ZMHB. Zoologisches Museum, Museum ftir Naturkunde der Hum-bolt-Universitit zu Berlin, Invalidenstr. 43, 1040 Berlin, Ger-many.

PLATEUMARINI ASKEVOLD, 1990

Plateumaris Thomson (1859:154). Revision of genus by Askevold(1991). Rather than making numerous, repetitive statementsabout taxonomic details for this genus, users are referred toAskevold (1991) for details about type specimens. Unless other-wise stated, lectotype and neotype designations were made byAskevold (1991); a detailed account of the historical use ofnames is given there also, and not repeated here.

P. aurifera (LeConte, 1851:313). Lectotype ( MCZ #4247. Vali-dated, and removed from synonymy with P. germari byAskevold (1991).

P. balli Askevold (1991). Holotype c, CNC #19497, and 23paratypes, ISAC, USNM, AMNH, CNC, APHIS, UAE, UVDZ,CAS and SMCL.

P. diversa (Schaeffer, 1925:143). Holotype’9, USNM #42397 andallotype c, USNM.

P. dubia (Schaeffer, 1925:152). Holotype c), USNM #4240-C, allo-type 9 USNM #42400 and 4 paratypes, USNM.

1991] Askevold 169

Donacia idola Hatch (1938:110). Holotype c, USNM type#103893, allotype 90SUC and 45 paratypes, OSUC. Syn-onymy with P. dubia by Askevold (1991).

P. flavipes (Kirby, 1837:223). Holotype 9, BMNH. This name wasused incorrectly by previous authors; the taxon previouslyknown as P. flavipes is now called P. shoemakeri (Schaef-fer). The taxon formerly called P. wallisi is now correctlycalled P. flavipes.

Donacia wallisi (Schaeffer, 1925:147). Holotype ( USNM#42399, allotype 9 USNM #42399, and 13 paratypes, USNM.Synonymy with P. flavipes by Askevold (1991).

P. frosti (Schaeffer, 1925). Plateumaris frosti was formerly placedas a subspecies of P. emarginata (now P. nitida).

Donacia emarginata var. frosti Schaeffer (1925:136). Holotypec, USNM #42395, allotype 9, USNM #42395, 28 paratypes,USNM, and 3 paratypes, MCZ.

P. fulvipes (Lacordaire, 1845:192). Holotype c, BMNH.P. germari (Mannerheim, 1843:306). Lectotype 9, MCZ #4250,

and 1 syntype, UMHF.Donacia flavipennis Mannerheim (1843"306). Lectotype c,MCZ #4251, and 1 syntype, UMHF. Specimens from westernCanada possessing flavous elytra and/or pronotum weregiven varietal status by Schaeffer (1925); Askevold (1991)gave no taxonomic status to specimens showing such varia-tion. Synonymy with P. germari by Askevold (1991).

Donacia dives LeConte (1851:314). Holotype 9, MCZ #4245.Donacia serricauda Schaeffer (1919:318). Holotype 9, USNM

#42396.P. metallica (Ahrens, 1810:33). Neotype cy, CNC #19493.

Donaciafemoralis Kirby (1837:223). Lectotype 9, BMNH.Donacia cataractae Newman (1838:391). Lectotype c, BMNH.Donacia parva Lacordaire (1845:190), Lectotype c, BMNH.Donacia indica Melsheimer (1847:158) (not Clark, 1866:1).

Lectotype 9, MCZ #33363.Donacia nana Melsheimer (1847:160). Neotype c, MCZ

#32953.Donacia gentilis LeConte (1851:314). Lectotype (y, MCZ

#4249, and two paralectotypes MCZ #4249.

170 Psyche [Vol. 98

P. neomexicana (Schaeffer, 1925:154). Holotype or, USNM#42401-1, allotype 9 USNM #42401, three paratypes,USNM, and two paratypes, MCZ #27227 [Fall colin.].

Donacia longicollis Schaeffer (1925:156). Holotype cr, USNM#4248-2, allotype USNM #42402 and 22 paratypes, USNM.Synonymized with P. neomexicana by Askevold (1991).

Donacia vermiculata Schaeffer (1925:158). Holotype 9, USNM.Synonymized with P. neomexicana given by Askevold(1991).

P. nitida (Germar, 1811:31). Lectotype 9 MLU. This name wasused incorrectly by Schaeffer (1925); the species misidenti-fled by Schaeffer as P. nitida is now called P. schaefferi. Thetaxon to which the name P. nitida applies was called P.emarginata by Schaeffer (1925) and other authors.

Donacia emarginata Kirby (1837:224). Lectotype or, BMNH.Synonymized with P. nitida by Askevold (1991).

Donacia binodosa LeConte (1851:316) (cf. Suffrian, 1872).Unjustified emendation or nomen nudum. Synonymy with P.nitida by Askevold (1991).

Donacia juncina Couper (1864:88). Lectotype 9, LEM, desig-nated by Becker (1974). Synonymized with P. nitida byAskevold (1991).

Donacia emarginata var. pacifica Schaeffer (1925:135). Holo-type c, USNM #42394, allotype 9, USNM #42394, and 9paratypes, USNM. Synonymized with P. nitida by Askevold(1991).

P. notmani (Schaeffer, 1925:132). Holotype cr, USNM #42393,allotype 9 #42393, and five paratypes, USNM, 1 paratype MCZ#27226 [Fall Colln.].

P. pusilla (Say, 1827:293). Neotype c, MCZ type #32950.Donacia rugifrons Newman (1838:391). Types of this name

have not been found, but should be in BMNH. Three speci-mens (of P. rufa, however, not P. pusilla) were found in theMCZ that are labelled Trenton Falls, New York, collected byDoubleday. Similar labels are borne by specimens of twoother Newman species D. cincticornis and D. cataractaeand I consider the possibility that these might be specimensof Newman’s D. rugifrons.

1991] Askevold 171

Donacia pyritosa LeConte 1857(1860):66. Lectotype or, MCZtype #4246. This name was formerly used as a variety of P.pusilla, but synonymized by Askevold (1991).

P. robusta (Schaeffer, 1919:318). This name was used as a varietyof P. pusilla until elevated to species status by Askevold(1991).

Donacia pusilla var. robusta Schaeffer (1919:318). Holotype c,USNM #42398, and 3 paratypes USNM #42398.

P. rufa (Say, 1827:283). Neotype c, MCZ #32947. This specieswas formerly called P. sulcicollis Lac. The name D. rufa hasbeen used incorrectly by authors (Marx 1957:250, Schaeffer1925:117), and was transferred from Donacia to Plateumarisby Askevold (1991) [cf. also D. (Donaciomima) tuberculataLacord., below].

Donacia affinis Kirby (1837:224) (not Kunze, 1818). Lectotype9, BMNH. Transferred to synonymy with P. rufa from syn-onymy with P. sulcicollis by Askevold (1991).

Donacia sulcicollis Lacordaire (1845:186). Lectotype cr,BMNH. Synonymized with P. rufa by Askevold (1991).

Donacia chalcea Lacordaire (1845:188). Holotype 9, MLU.Transferred to synonymy with P. rufa from P. flavipes, byAskevold (1991).

Donacia kirbyi Lacordaire (1845:202). Replacement name forD. affinis Kirby; transferred to synonymy with P. rufa fromP. sulcicollis by Askevold (1991).

Donacia jucunda LeConte (1851:315). Holotype c, MCZ#4248. Transferred to synonymy of P. rufa from P. sulcicol-lis by Askevold (1991).

P. schaefferi Askevold (1991). Holotype c, USNM, and 7paratypes, USNM, CUIC and UMMA.

Donacia nitida, sensu Schaeffer (1925:130) (see P. nitida,above).

P. shoemakeri (Schaeffer, 1925:129). This species was formerlyincorrectly known as P. flavipes (see P. flavipes, above), val-idated by Askevold (1991).

Donacia flavipes var. shoemakeri Schaeffer (1925"129). Holo-type cr, USNM #4239-1 and 5 paratypes USNM #42391.

Donaciaflavipes var. lodingi Schaeffer (1925"129). HolotypeUSNM #4239-2 and 1 paratype, USNM #42392. Southern

172 Psyche [Vol. 98

specimens, from Alabama, were given varietal status bySchaeffer (1925), but Askevold (1991) did not give this geo-graphic variation taxonomic status.

Poecilocera Schaeffer (1919:316). Elevated to genus status byAskevold (1990)

P. harrisii (LeConte, 1851:316). Lectotype 9 MCZ #4244, desig-nated by Askevold (1990:635); the abdomen is now missing butwas present when I first examined it. LeConte stated he had atleast two specimens, and 9, although only a single specimenremains in the LeConte collection. P. harrisii was placed inSominella by Goecke (1931), but Marx (1957) reverted to place-ment in Donacia (Poecilocera), evidently unaware of Goecke’streatment.

DONACIINI KIRBY, 1837

Donaciella Reitter (1920:38). Elevated to genus by Askevold(1990), this name has been treated by European authors as asubgenus of Donacia, which has not been recognized in NorthAmerica despite direct comparisons by Schaeffer (1925) withspecies occurring in the Palaearctic Region.

D. pubicollis (Suffrian, 1872:21). Transferred to Donaciella byAskevold (1990). Suffrian did not indicate the number ofspecimens before him, but given a range of size measure-ments, had at least two. A series of 12 specimens are pre-served in MLU, associated with a Suffrian label. Asdetermined previously (see P. chalcea and P. nitida, above)these specimens are also likely original Suffrian material,and are here designated. LECTOTYPE , following a Suf-frian label ["pubicollis m. Illinois", on large, green, black-bordered paper]: "25519 [red handwriting on small whitelabel] / LECTOTYPE r Donacia pubicollis Suffrian 1872design. I.S. Askevold 1991". The remaining 11 specimensare here designated paralectotypes. These bear the same red,handwritten number and my paralectotype label: 25516 (r),25517 (c), 25520 (cy), 25521 (c), 25522 (cr), 25523 (9),25524 (9), 25525 (9), 25526 (9), 33256 (9) and 25518 (sexundetermined, abdomen missing).

1991] Askevold 173

Donacia pubicollis Crotch (1873:21) (not Suffrian, 1872:21).Synonymy with Donacia pubicollis Suffrian by Marx (1957).No specimens of Crotch’s species were found at MCZ indi-cated as type, probably because it was not recognized thatCrotch had also described the species, until Marx (1957). Ipresume the type must be lost, but it is hardly necessary todesignate a neotype for such a well-known species.

Donacia (Donaciomima) Medvedev (1973:876). The subgenusDonaciomima was originally based on an eastern Palaearcticspecies, but was disregarded in subsequent treatments. AllNearctic species of Donacia were formerly placed in Donacia s.str., but Askevold (1990) revalidated Donacia (Donaciomima),and defined it to include the majority of species hitherto placedin the nominate subgenus. Most Nearctic species (23) of Dona-cia are included, of which two remain undescribed.

D. assimilis Lacordaire (1845:153). Holotype c, BMNH. Lacor-daire had before him a single specimen of undetermined sex:"TYPE [red] / TYPE [white] / E. Coll. Lafert6 / Donaciaassimilis type Lacord. / Donacia assimilis Dej. h. in Ameri.bor. J. Leconte".

Donacia glabrata Schaeffer (1919:313) (not Solsky, 1872).Based on single c, USNM holotype: "Type / Clementon,VIII. 6. N.J. / Brooklyn Museum Colln. 1929 / TYPE USNM42387 / D. glabrata type Schffr. / Donacia assimilis Lac."Synonymy with D. assimilis by Schaeffer (1925).

D. biimpressa Melsheimer (1847:159). Lectotype cr, MCZ #33365.Melsheimer had before him at least two specimens; the singleremaining specimen is here designated as lectotype" "Melsh.biimpres / Type [red] / biimpressa [large Melsheimer label,handwritten] / LECTOTYPE Donacia biimpressaMelsheimer design. I.S. Askevold 1990 [red] / MCZ. Lecto-type 33365 [red]".

Donacia aurichalcea Melsheimer (1847:159). Evidently basedon a single specimen. Holotype or, MCZ #33364: "Melsh.aurichalcea / Type [red] / dissected and remounted ISAskevold 1983 / aurichalcea [large handwritten Melsheimerlabel] / Donacia biimpressa Melsheimer 1847 det. I.S.

174 Psyche [Vol. 98

Askevold 1983 /M.C.Z. Lectotype 33364 [red]". Synonymywith D. biimpressa by Schaeffer (1925).

Donacia torosa LeConte (1851:313). LeConte did not specifynumber of specimens, but three specimens found in theLeConte collection are here designated as types. Lectotype c,MCZ: "pink disc / LECTOTYPE [blue trimmed disc] / TYPE4243 [red] / D. torosa Lec. / J. LECONTE COLLECTION /

LECTOTYPE cy Donacia torosa LeConte 1851 designated ISAskevold 1983 [red]". Paralectotype cy: "pink disc / Type4243 [red] / torosa 2 / J. LECONTE COLLECTION / PAR-ALECTOTYPE cy Donacia torosa LeConte [red]"; paralecto-type [sex undetermined, abdomen missing] "pink disc /torosa 3 / J. LECONTE COLLECTION / PARALECTO-TYPE Donacia torosa LeConte [red]". Synonymy with D.biimpressa by Schaeffer (1925).

D. caerulea Olivier (1795:10). Neotype cy, CNC #19492, desig-nated by Askevold (1987b:347). Discussed by Askevold(1987b), this species was formerly known as D. aequalis; D.caerulea was used incorrectly by Schaeffer (1925) and Marx(1957), which they applied to certain specimens of D. prox-ima.

Donacia aequalis Say (1824:248). Neotype c, MCZ #32948,designated by Askevold (1987b:347); placed in synonymywith D. caerulea by Askevold (1987b).

Donacia confusa Lacordaire (1845; 109) (not LeConte, 1851).Lectotype cy, BMNH, designated by Askevold (1987b:347);placed in synonymy with D. caerulea by Askevold (1987b).

Donacia confusa LeConte (1851:313) (not Lacordaire, 1845).Lectotype cy, MCZ #32949, designated by Askevold(1987b:347); placed in synonymy with D. caerulea byAskevold (1987b).

D. cazieri Marx (1957:213). Holotype cy, CNC, allotype , CNC,four paratopotypes, CNC, and one paratopotype, AMNH,Goose Bay, Labrador (various dates and collectors, see Marx1957).

D. confluenta Say (1827:293). Neotype cy, MCZ #32956, desig-nated by Askevold (1987a:634); removed from synonymywith D. subtilis Kunze by Askevold (1987a).

1991] Askevold 175

D. cuprea Kirby (1837:225) (not Melsheimer, 1847). LectotypeBMNH, designated, and removed from synonymy with D.quadricollis Say, by Askevold (1987a:633).

Donacia curticollis Knab (1905:122) (not Haupt, 1956).Removed from synonymy of D. quadricollis Say byAskevold (1987a). Lectotype 9, and three paratypes, INHS;designation and details given by Frison (1927:160).

D. dissimilis Schaeffer (1925:115). Holotype 9, USNM: "Floridaex Liebeck / Charles Schaeffer Collection / HS BarberBequest 1950 / dissimilis Holotype C. Schaeffer coll. 1935Barber / Donacia dissimilis Schffr.". Allotype c, PUR:"Dunedin [Florida] 4.16.24 W.S.B. coll./2724 / c Allotype /Donacia dissimilis cr allotype Schaeffer".

D. distincta LeConte (1851:313). LeConte did not state the numberor sex specimens before him; two specimens were found inthe LeConte collection. Lectotype cr and paralectotypeMCZ #4242, here designated: "cr [pale blue disc] / LECTO-TYPE [blue trimmed disc] / Type 4242 [red] / D. distinctaLec. aequalis Kirby / LECTOTYPE cy Donacia distinctaLeConte 1851 designated 1983 IS Askevold [red]" [specimendissected, with genitalia placed in microvial pinned beneathspecimen]; paralectotype 9, "9 [pale blue disc] / Type 4242[red] / distincta 2 / J. LECONTE COLLECTION / PAR-ALECTOTYPE 9 Donacia distincta LeConte [red]".

Donacia distincta occidentalis Mead (1938:115)..HolotypeCAS #4691, allotype 9, CAS #4692, and paratype, MCZ [exFall Coll.], Luthers Pass, Cal. vi-15-1930 AT McClay. Syn-onymy with D. distincta by Marx (1957), to which I agree.

D. fulgens LeConte (1851:312). Lectotype cr, MCZ #4240, desig-nated by Askevold (1987a:634). At the time of this designa-tion, another syntype specimen was not sent to me; I here adda 9 paralectotype from the LeConte collection: "pale bluedisc, two cuts / Type #4240 [red] / D. fulgens Lec. / J.LECONTE COLLECTION / PARALECTOTYPE 9 Donaciafulgens LeConte [red]". Removed from synonymy of D. sub-tilis Kunze by Brivio and Balsbaugh (1984).

D. hirticollis Kirby (1837:226). Holotype c, BMNH. Kirby statedhe had before him a single specimen, which bears the follow-ing labels: "TYPE [red trimmed disc] / N. Amer. 5954a

176 Psyche [Vol. 98

[white disc] / Donacia hirticollis Kirby N. Ameri. 5954 Rev.Wm. Kirby [type, on underside]". The specimen lacks theright fore leg and left hind leg, and right elytron smashed bypin.

Donacia rudicollis Lacordaire, 1845:108. LECTOTYPE 9,BMNH, by present designation. Lacordaire stated he hadbefore him two, supposedly female, specimens. The singlespecimen found is here designated: "LECTOTYPE [bluetrimmed disc] /TYPE [red trimmed disc] /TYPE [white] /E.Coll. Lafert6 / [green, square paper, 4 mm] / 67.56 / Donaciarudicollis Type Lac. / Donacia rudicollis. Reiche Amer. bor./LECTOTYPE 9 Donacia rudicollis Lacordaire 1845 desig-nated 10.vii.1984 I.S. Askevold [red] / Donacia hirticollisKirby det. I.S. Askevold 1984". I have not determined theoriginal source of synonymy.

D. liebecki Schaeffer (1919:314). Holotype c, USNM #42390:"Type cr / Wyandanch L Isld. / Brooklyn Museum Colin.1929 / liebecki type USNM 42390 [red] / Donacia liebeckiSchffr. [red bordered]". Allotype 9: "Type 9 Allotype /

Wyandanch L Isld./liebecki Schffr. Allotype USNM 42390[red] / Brooklyn Museum Colin. 1929". Five paratypes MCZ#27228 (two ex Fall Colin.), Malaga 1.8 N.J. One specimenlabelled as a paratype (USNM) may be a paratype, though itdoes not bear labels indicating it is part of the original syn-type series of 11 specimens: "Lakehurst NJ / Ed. A. BischoffCollection / Brooklyn Museum Colin. 1929 / liebecki USNMparatype ? 42390 [red]". Another specimen labelled as aparatype (USNM) cannot be a paratype because it is fromWading River, Long Island, a locality not mentioned bySchaeffer.

Donacia pallipes Lacordaire (1845:149) (not Kunze, 1818).Holotype 9, BMNH. Lacordaire had before him a singlespecimen which he supposed to be male. It is female, how-ever. "TYPE [red trimmed disc] / TYPE [white] / 67.56 / D.pallipes Lac. cr type / Donacia type 32. pallipes Lac. 119 39Philadelp Am. Sept. Boston D. Green [green, folded] / Dona-cia liebecki Schaeffer det. IS Askevold 1985." D. pallipeswas listed in early catalogues as a synonym of D. aequalisSay; Schaeffer (1925) realized that D. pallipes Lac. was a

1991] Askevold 177

senior synonym of D. liebecki, but preoccupied, thus D.liebecki remains valid. Synonymy with D. liebecki bySchaeffer (1925).

D. limonia Schaeffer (1925). This name was described as a varietyof D. biimpressa Melsh.; Marx (1957) reduced it to syn-onymy with D. biimpressa. I here remove it from synonymy,and recognize D. limonia as a valid species on the basis ofdistinctive structure of male genitalia; further treatment ofthis species as a member of the D. distincta-Group isplanned. NEW STATUS

D. biimpressa var. limonia Schaeffer (1925:100). Holotype c,USNM: "Indian Pass N.Y. VII. 11. / Essex Co. Quirsfeld /

Brooklyn Museum Coll. 1929 / limonia Schffr. TYPE USNM42388 / Donacia biimpressa limonia Schffr.". Allotype 9,USNM, Van Cortlandt Park, N.Y. (additional details notrecorded), and five paratypes, USNM (label details notrecorded).

D. rnagnifica LeConte (1851:310). LeConte did not state the num-ber of specimens before him, but stated he had both male andfemale specimens. Four specimens in the Leconte collectionbear appropriate label data, and are here designated, lecto-type c, MCZ #4237, one cr and two 99 paralectotypes, MCZ#4237: "LECTOTYPE [blue trimmed disc] / cr [pale bluedisc] / Type 4237 [red] / Donacia magnifica Lec./LECTO-TYPE c Donacia magnifica LeConte 1851 designated I.S.Askevold 1984 [red]"; the three paralectotypes are labelled"magnifica 2", "magnifica 3" and "magnifica 4", each withthe labels "Type 4237" [red] and "PARALECTOTYPEDonacia magnifica LeConte [red]". In earlier catalogues, D.magnifica was listed under D. cincticornis.

D. megacornis Blatchley (1910:1103). Lectotype cr, PUR. Blatch-ley did not state the number of specimens before him, butdesignated a lectotype c later (Blatchley 1930): "Lake Co.,Ind. W.S.B. 5-28-1903 / Donacia megacornis, sp. nov. [hand-written]".

Donacia megatocera [sic] Weise (1913:118). Unnecessaryreplacement name. Clavareau (1913:23) altered the spellingof megatocera to megalocera, probably supposing that it wasa printer’s error. Weise proposed the name D. megalocera to

178 Psyche [Vol. 98

replace Blatchley’s D. megacornis, on the basis that mega-cornis is a name formed from two languages; the namemegacornis is valid, and Schaeffer (1925) restored use of thename D. megacornis, and synonymized D. megatocera.

D. porosicollis Lacordaire (1845:150). Holotype 9, BMNH. Lacor-daire described this species from a single, apparently femalespecimen (which it is): "TYPE [red trimmed disc] / TYPE[white] /Donacia Chvt longipennis parraceicollis Lac. typeAmerica bor. Boston D. Greene 627 [green] / 67.56 / E. Coll.Chevt. / 21. [folded paper] / D. porosicollis Lac. type". Thespecimen is a little crushed and is missing the right middleleg.

D. pubescens LeConte (1868:55). Described from a unique speci-men, which LeConte stated was from the Smoky Hill River.Holotype 9, MCZ #4234: "Type [red] / D. pubescens Lec.".

D. rugosa LeConte (1878:415). Described from a unique speci-men, from Enterprise, Florida. Holotype 9, MCZ #4241:"Enterprise May 20 Fla. / 402 / Type 4241 [red] / D. rugosaLec./J. LECONTE COLLECTION".

D. subtilis Kunze (1818:12) (replacement name for D. aeneaAhrens).

Donacia aenea Ahrens (1810:21) (not Hoppe 1795:44; not Gis-tel 1857:524). Neotype cy, CMP; designation and discussionby Askevold (1987a:633).

Donacia quadricollis Say (1827:282). Neotype cy, MCZ #32951,designated by Askevold (1987a:633). The name D. quadri-collis was formerly used by authors (e.g. Schaeffer 1925,Wilcox 1954, Marx 1957), applied to the species now knownas D. cuprea Kirby (see above) but was synonymized with D.subtilis by Askevold 1987a).

Donacia aerea Lacordaire (1845:148) (not Schrank, 1798). Lec-totype cr, BMNH, designated by Askevold (1987a). I havenot determined the original source of synonymy.

Donacia subtilis magistrigata Mead (1938:113). HolotypeCAS #4689, allotype 9, CAS #4690, two paratypes, CAS,and one paratype, MCZ #27230 (ex Fall Colin.), Clarksburg,Cal, June-4-1931, A.T. McClay. Synonymized with D. ful-gens by Marx (1957) but transferred to synonymy with D.subtilis by Askevold (1987a).

1991 Askevold 179

Donacia tuberculata Lacordaire (1845:155). Lectotype c andparalectotype 9, BMNH. Two specimens were found thatappeared to be authentic Lacordaire specimens, which arehere designated: "LECTOTYPE [blue trimmed disc] / D.tuberculata Lac / 67.56 / LECTOTYPE cr Donacia tubercu-lata Lacordaire 1845 designated 10.vii.1984 I.S. Askevold[red]". A female is labelled: "PARALECTOTYPE 9 [paleblue-trimmed disc] / TYPE [red trimmed disc] / TYPE[white] / E. Coll. Lafert6 / Donacia tuberculata mihi h. inAmeri. bor. D. Leconte [green, folded] / 67.56 / PARALEC-TOTYPE 9 Donacia tuberculata Lacordaire 1845 designated10.vii.1984 I.S. Askevold [red]." Donacia tuberculata wassynonymized with D. rufa Say by Schaeffer (1925), anarrangement followed by Marx (1957). Leng (1891) had usedD. rufa in the sense used above, as a species of Plateumaristo which it was transferred by Askevold (1991). With trans-ferral of D. rufa to Plateumaris, D. tuberculata becomesthe senior, available name for the taxon previously recog-nized as D. rufa, sensu Schaeffer (1925) and Marx (1957).NEW STATUS

Donacia rutila Melsheimer (1847:160). Lectotype 9, MCZ#32952, by present designation: "LECTOTYPE [bluetrimmed disc] / Melsh. rutila / M.C.Z. Lectotype 32952 [red]/ tuberculata [large handwritten Melsh. label] / LECTOTYPE9 Donacia rutila Melsheimer, design. I.S. Askevold 1990[red]". Earlier authors (e.g. Leng 1891) placedD, rutilaunder D. tuberculata. However, this name was transferred tosynonymy of D. rufa. Because D. tuberculata is revalidated,D. rutila becomes its synonym. NEW SYNONYMY

D. tuberculifrons Schaeffer (1919:315). Holotype c, USNM:"Toronto VII.7 Can. / Brooklyn Museum Coll. 1929 / TYPEUSNM 42389 / D. tuberculifrons type Schffr.". Allotype 9and one paratype, USNM (label details not recorded). Fourparatypes, MCZ #27229 (1 ex Fall Colln.): "Toronto, Can.VII.I".This species was treated as a member of the D. subtilis-Group by Askevold (1987a).

D. vicina Lacordaire (1845:154) (not Haupt, 1956). Holotype 9,BMNH. Lacordaire described the species from a single

180 Psyche [Vol. 98

specimen received from Reiche, incorrectly stating it to bemale: "HOLOTYPE [red trimmed disc] / TYPE [white] / E.Coll. Lafert6 / Donacia vicina Dej. h. in Amer. bor. D.Leconte [green]".

Donacia (Donacia) Fabricius, 1775. All Nearctic species ofDonacia were formerly placed in Donacia s. str. (e.g. Leng1891, Jacoby and Clavareau 1904, Clavareau 1913, Jolivet1970, Borowiec 1984), but Askevold (1990) restricted thenominate subgenus to include 10 Nearctic species.

D. cincticornis Newman (1838"39i). Lectotype 9, BMNH, by pres-ent designation. Newman did not state the number of speci-mens before him. The single specimen remaining is thereforehere designated: "LECTOTYPE [blue trimmed disc] / TYPEH.T. [red trimmed disc] / Ent. Club. 44-12 / R. Foster Tren-ton Falls, New York / Donacia cincticornis type Newm. /

LECTOTYPE 9 Donacia cincticornis Newman 1838 desig-nated 11.vii. 1984 I.S. Askevold 1984".

Donacia lucida Lacordaire (1845:106). Lectotype, BMNH, bypresent designation. The species was described from at leasttwo specimens. The single remaining specimen under thisname is selected as lectotype, which bears the followinglabels: "LECTOTYPE [blue trim disc] / TYPE [red trim disc]/ Donacia Lucida Lac Type / TYPE / E. Coll. Chevt. / D.lucida cr Lac. type / Donacia type 626 lucida (Chvt) Lac. mon6 Am. bor. Boston D. Greene [green label] / 67.56 / LECTO-TYPE [abdomen missing, probably male] Donacia lucidaLacordaire 1845 designated 10.VII.1984 I.S. Askevold [red] /

Donacia cincticornis Newman det. I.S. Askevold 1984". Syn-onymy with D. cincticornis was apparently first by Leng(1891).

Donacia rufipennis Lacordaire (1845:110). Lectotype cy,BMNH, by present designation. Lacordaire described thisspecies from four specimens. A male is here selected as lec-totype, bearing the following labels: LECTOTYPE [bluetrimmed disc, added by me] ! TYPE [red] ! Donacia rufipen-nis Type Lac. [white] ! TYPE [white] ! Donacia type picipen-nis Chevt rufipennis Lac. mon. Ameri. bor. Boston D. Greene[green] / 12 / 67.56 / LECTOTYPE Donacia rufipennis

19911 Askevold 181

Lacordaire 1845 designated 10.VII. 1984 I.S. Askevold [red] /

Donacia cincticornis Newman det. I.S. Askevold 1984".Synonymy with D. lucida by Crotch (1873), therefore placedas synonym of D. cincticornis by later authors.

Donacia pulchella LeConte (1851:312). LeConte apparentlydescribed a single specimen, stating at least that he had nomales. The single specimen found in the LeConte collectionis therefore assumed to be the holotype, which LeConteincorrectly sexed. Holotype 9, MCZ #4235: "pink disc / Type4235 [red] / D. pulchella LeC.". Synonymy with D. lucida byCrotch (1873), therefore placed as synonym of D. cincticor-nis by later authors.

Donacia antillarum Suffrian (1864:282). Suffrian described thisfrom at least two specimens, because he gave a range of size.A single specimen here designated as lectotype was found inMLU, associated with the Suffrian label: "Antillarum m.Cuba [large, green, black-bordered label]". LECTOTYPE(sex undetermined, abdomen missing): "25004. [red hand-writing on small label] / LECTOTYPE Donacia antillarumSuffrian 1864 design. I.S. Askevold 1991". Treated as a sub-species of D. cincticornis by Schaeffer (1925); synonymywith D. cincticornis by Marx (1957).

Donacia cincticornis var. tryphera Schaeffer (1925:81). Holo-type cr, USNM: "Centerton VII N.J. / Brooklyn MuseumColl. 1929 / tryphera TYPE USNM 42385". Allotype 9, and11 paratypes, USNM (label data not recorded). Marx (1957)reduced Schaeffer’s subspecies to synonymy; I agree withthis synonymy.

Donacia cincticornis var. tenuis Schaeffer (1925:82). Holotypec, USNM" "Watershield/Swansea SO 11 Aug. 11 Fredk.Knab. / Type c / tenuis TYPE USNM 42386 / D. cincticornistenuis Schffr.". Allotype 9, and 16 paratypes, USNM (labeldata not recorded). Two paratypes, MCZ #27232 (ex FallColin.): "Spencer L., N.Y. viii.26.17, H. Dietrich". Marx(1957) reduced Schaeffer’s subspecies to synonymy; I agreewith this synonymy.

D. edentata Schaeffer (1919:313). Holotype or, USNM: "Crum Lk.N.Y. / U.S.N.M. Type No. 42383 / D. edentata type Schffr.".Allotype 9 and five paratypes, USNM (label data not

182 Psyche [Vol. 98

recorded). One paratype CAS, and eight paratypes, MCZ,"Centerton N.J., 5.31".

D. hypoleuca Lacordaire, 1845:101. Lectotype c, BMNH, by pres-ent designation. Lacordaire lescribed this species from two,supposedly 9, specimens; however, there is only a single crspecimen under hypoleuca which bears an original Lacor-daire label: "Lectotype cr [blue trim disc added by me] / E.Coll. Lafert6 / 67.56 / LECTOTYPE c Donacia hypoleucaLacordaire 1845 designated 11.VII.1984 I.S. Askevold [red,added by me]". Also in the BMNH is another specimen (c)which bears a type label; it bears the labels "TYPE [white] /This specimen cannot possibly be the type of hypoleuca Lac.The labels have evidently been misplaced. C.G. / Donaciahypoleuca. Van Wintheim h. in Amer. bor. Neu Orleans D. v.Wintheim [green Lacordaire label]". This latter label is evi-dently genuine, according to information given by Lacor-daire, mentioning van Wintheim specifically (1845:101);however, the specimen does not agree with the description atall, and is in fact a specimen of D. palmata Olivier. Whetherthe specimen is a syntype or not is irrelevant, for the otherspecimen is here designated lectotype in the interest of con-serving the traditional use of D. hypoleuca.

D. militaris Lacordaire (1845"115). Lectotype c, BMNH, by pres-ent designation. Lacordaire did not state the number of speci-mens before him, only that he had no female specimens; acrspecimen was found: "LECTOTYPE [blue trimmed disc] /

TYPE [red trimmed disc] / Donacia militaris type Lac. /

TYPE [white] / E. Coll. Chevt. / 67.56 / D. militaris Lac. [??]type / Donacia type 630 bispinosa (Chvt) 9 militaris Lacmon. America bor. Boston D. Greene [green, folded] LEC-TOTYPE Donacia militaris Lacordaire 1845 designated10.vii.1984 I.S. Askevold [red]". Formerly misidentified as asynonym of D. palmata, but removed from synonymy bySchaeffer (1925).

Donacia floridae Leng (1891:166). No type specimen specifi-cally identified was found in USNM (R.E. White, pers.comm.); however, a specimen, USNM type #558 was foundamong specimens in "regular collection", which is assumed

1991] Askevold 183

to be a paratype. Synonymized by Schaeffer (1925), which isdoubtless correct.

D. palmata Olivier (1795:8). Type(s) almost certainly destroyed;no neotype is designated because Olivier provided a figure ofthe protarsus of male specimens; the species is unique amongDonaciinae in this structure.

Donacia claudicans Germar (1821:173). Germar evidentlydescribed this from a single male specimen, specifically dis-cussing the unusual and unmistakable protarsal structure.According to Suffrian (1872), he had the Germar collectionin his possession (see extended discussion of P. nitida byAskevold, 1991). Thus, Germar’s type, if existing, should bein the Suffrian collection. Three specimens (cc) are associ-ated with the large, green, black-bordered Suffrian label inMLU: "palmata Oliv. Lec. claudicans Grm. Illinois". Neitherof these fits especially the specific colours that Germardescribed, and so it is quite likely that none of these is theGermar specimen. However, it is not necessary to designate aneotype because the species is so distinctive and Germardescribed the distinguishing character. I have not traced thesource of original synonymization.

Donacia palmata angustipes Marx (1957"237). Holotype c andallotype 9 #8452, and 19 paratypes, CAS, "Devils Riv. nearDel Rio, Texas, vi-16-32, J.O. Martin"; 2 paratypes, CAS,"Devils Riv. near Del Rio, Texas, v-25-32, J.O. Martin".These Texas specimens differ little from other, typical D.palmata from other regions, differing essentially only in thatmales have a less broadly palmate basal protarsomere. NEWSYNONYMY

D. parvidens Schaeffer (1919:311). Holotype c, USNM: "LahawayOcean Co. V. N.J. / Brooklyn Museum Coll. 1929 / U.S.N.M.Holotype 42384 / D. parvidens type Schffr.". Four paratypes,USNM (label data not recorded) and one paratype, MCZ,"Lahaway, Ocean Co., N.Y."

D. piscatrix Lacordaire (1845:113). Lectotype c, BMNH, by pres-ent designation. Lacordaire did not state the number of speci-mens before him, only that he had no male specimens. One 9specimen was found: "LECTOTYPE [blue trimmed disc] /

TYPE [red] / Donacia piscatrix Lacord. / E. Coll. Lafert6 / D.

184 Psyche [Vol. 98

piscatrix Lac./Donacia Am. bor. [green] / 67.56 / LECTO-TYPE 9 Donacia piscatrix Lacordaire 1845 designated10.vii. 1984 I.S. Askevold [red]".

Donacia carolina Lacordaire (1845:114). Lectotype or, BMNH.Lacordaire stated he had both a male and female before him.The single, apparently authentic, Lacordaire specimen foundamong specimens of D. piscatrix, is here designated" "LEC-TOTYPE [blue trimmed disc] / TYPE [red] / Donacia car-olina Lacord. / TYPE [white] / 67.56 / Donacia carolina mihi.h. in Amer. bor. D. Latreille [green] / Donacia piscatrix det.I.S. Askevold ’84". I did not trace the original source of syn-onymization.

Donacia cuprea Melsheimer (1847:158) (not Kirby, 1837). Lec-totype or, MCZ, by present designation. Described from atleast two specimens, for Melsheimer gave a size range andindicated a "var. a". The single remaining specimen is heredesignated: "cy [discoloured square] / Melsh. cuprea / Type[red] / cupria [sic] [large handwritten Melsheimer label] /

LECTOTYPE c Donacia cuprea Melsheimer design. I.S.Askevold 1990 [red] / M.C.Z. Lectotype 33366 [red]". Syn-onymy with D. piscatrix apparently first by Schaeffer (1925).

Donacia alutacea LeConte (1851:311). LeConte had at leasttwo specimens before him. Two specimens found in theLeConte collection are here designated: lectotype cr, MCZ#4239, "LECTOTYPE [blue trimmed disc] / Type [partlyred] / c [pink disc] / Type 4239 [red] / D. alutacea Lec.cuprea//Mels. / LECTOTYPE c Donacia alutacea LeConte1851 designated I.S. Askevold 1984 [red]"; and paralectotypec, "9 [pink disc] / Type 4239 [red] alutacea 2 PARALECTO-TYPE cr Donacia alutacea LeConte [red]". Synonymy with D.piscatrix by Crotch (19873).

Donacia congener LeConte (1851:310). LeConte did not state num-ber of specimens before him, only that he had no males. Twospecimens in the LeConte collection bear orange discs, so itappears he had at least two specimens, and I here designatethem. Lectotype 9, MCZ #4238: "LECTOTYPE [blue trimmeddisc] / 9 [orange disc] / Type 4238 [red] / D. congener Lec. /

LECTOTYPE 9 Donacia congener LeConte 1851 designated I.S.Askevold 1984"; and one 9 paralectotype, MCZ #4238 "9

1991 Askevold 185

[orange disc] / Type 4238 [red] / congener 2 / PARALECTO-TYPE 9 Donacia congener LeConte [red]". Synonymy with D.piscatrix by Crotch (1873), treated as subspecies by Schaeffer(1925), resynonymized by Marx (1957).

D. proxima Kirby (1837:225). Lectotype 9, BMNH, designated byAskevold (1987b:347).

Donacia episcopalis Lacordaire (1845:105). Lectotype c, BMNH,designated by Askevold (1987b:348). Formerly placed in syn-onymy with D. proxima, transferred to synonymy with D.caerulea by Schaeffer (1925), who used this name used incor-rectly; D. episcopalis was transferred back to junior synonymywith D. proxima by Askevold (1987b).

Donacia californica LeConte (1861:357). Holotype 9, MCZ #4236(cf. Askevold 1987b:348). Given subspecies status by Marx(1957), but this classification rejected by Askevold (1987b).

Donacia caerulea, sensu Schaeffer (1925) and Marx (1957). SeeAskevold (1987b) for discussion.

D. rufescens Lacordaire (1845:112). Lectotype c, BMNH, by presentdesignation. Lacordaire had before him two specimens, suppos-edly female. However, both are male, of which one is here des-ignated: "LECTOTYPE [blue trimmed disc] TYPE [redtrimmed disc] TYPE [white] Donacia type 629 calcarata Chvt8 rufescens Lac. mon. America bor. Boston D. Greene [green,folded] / 15. / 67.56 / LECTOTYPE cr Donacia rufescens Lacor-daire 1845 designated 10.vii.1984 I.S. Askevold [red]".

D. texana Crotch (1873:22). Lectotype 9, MCZ #5032, by present des-ignation. Crotch did not state the number of specimens beforehim, but three specimens already indicated as types were foundin the MCZ. Lectotype 9, MCZ #5032, "Tex. Type 5032 [red] /

D. texana Cr. / LECTOTYPE 9 Donacia texana Crotch design.IS Askevold 1990 [red]"; paralectotype 9, MCZ #5032, "Tex./

Type 5032 [red] / texana 2 / PARALECTOTYPE Donacia tex-ana Crotch [red]" and paralectotype or, MCZ #5032, "Tex.Type 5032 [red] texana 3 PARALECTOTYPE Donacia tex-ana Crotch [red]".

Donacia texana var. minor Schaeffer (1925:72). Holotype c,USNM: "Centerton V. 28. N.J. / Donacia texana minor Schffr. c/ U.S.N.M. Type No. 42382". Allotype 9, and 6 paratypes,USNM (label data not recorded). Marx (1957) did not recognize

186 Psyche [Vol. 98

D. minor as a subspecies and therefore synonymized D. minor,to which I agree.

HAEMONIINI CHEN, 1941

Neohaemonia Sz6kessy (1941:148). This genus was formerly consid-ered congeneric with the Palaearctic genus, MacropleaSamouelle. Revision of genus and establishment of genus statusby Askevold (1988).

N. flagellata Askevold (1988:390). Holotype or, CNC, and 72paratypes of various locality data deposited in CARR, CAS,CNC, CUIC, EGRC, ISAC, JBWM, ROM, UAE, UMMA,USNM.

N. flohri (Jacoby, 1884:126). Holotype 9, MCZ #8468. Details givenby Askevold (1988).

N. melsheimeri (Lacordaire, 1845:211). Holotype cr, BMNH. Detailsgiven by Askevold (1988).

N. minnesotensis Askevold (1988:388). Holotype cr, CNC #19495, and38 paratypes with various locality data deposited in AMNH,CAS, CMP, CNC, DEFW, EGRC, FMNH, FSCA, ISAC,JBWM, LSUC, MCZ, NMDC, SCSU, UMMA, USNM.

N. nigricornis (Kirby, 1837:222). Holotype 9, BMNH. Details givenby Askevold (1988).

Haemonia americana Gu6rin-M6neville (1844:259). Holotype 9,BMNH. Details given by Askevold (1988).

ACKNOWLEDGEMENTS

I owe the many curators of museums cited above profuse thanks fortheir inestimable help with my studies about Donaciinae. Without theirassistance, this and other publications on this group would suffer sig-nificant incompleteness. Mostly I examined all types myself, butRichard E. White is owed particular thanks for providing details ofSchaeffer and Leng types that I was neglectful in recording data aboutpersonally, or did not personally examine.

Research over a number of years, and costs of publication, weresupported by NSERC grant A0428 held by R.E. Roughley, Universityof Manitoba, and in part also by an NSERC PGS Scholarship, an ErnstMayr Grant in aid of research at MCZ, and funds provided by CAS to

1991 Askevold 187

research at that institution. Some costs were supported by a grant fromCSRS, USDA to Florida A. & M. University.

SUMMARY

The Nearctic species of Donaciinae are catalogued according to themost recent classification of its members at tribal, generic andsubgeneric levels. Annotations are provided where significant depar-tures have been made from their preceding classification. Donaciatuberculata Lacordaire is removed from synonymy with D. rufa Say(now placed in Plateumaris). Donacia limonia Schaeffer is removedfrom synonymy with D. biimpressa Melsheimer. Donacia angustipesMarx is reduced to synonymy of D. palmata Olivier; D. rutilaMelsheimer is synonymized with D. tuberculata Lacordaire. Details oflabel data, number, kind and location of preservation of type speci-mens (if existing) of all described taxa are given. Lectotypes (and par-alectotypes, if any) are designated for the following nominal taxa: D.alutacea LeConte, D. biimpressa Melsheimer, D. carolina Lacordaire,D. cincticornis Newman, D. congener LeConte, D. cupreaMelsheimer, D. distincta LeConte, D. hypoleuca Lacordaire, D. lucidaLacordaire, D. magnifica LeConte, D. militaris Lacordaire, D. pisca-trix Lacordaire, D. rudicollis Lacordaire, D. rufescens Lacordaire, D.rufipennis Lacordaire, D rutila Melsheimer, D. texana Crotch, D.torosa LeConte and D. tuberculata Lacordaire. Types of D. palmataOlivier and D. claudicans Germar are believed lost, but are distinctivetaxa whose identity is not in doubt.

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forschenden Gesellschaft zu Halle 1: 9-48.ASKEVOLD, I.S.

1987a. The identity of Donacia cuprea Kirby, 1837 and Donacia quadricollis Say,1827, with a taxonomic revision of members of the Donacia subtilisKunze group (Coleoptera: Chrysomelidae: Donaciinae). The CanadianEntomologist 119: 629-645.

1987b. The identity of Donacia caerulea Olivier, 1795 (Coleoptera: Chrysomeli-dae: Donaciinae). The Coleopterists’ Bulletin 41: 345-349.

1988. The genus Neohaemonia Sz6kessy in North America (Coleoptera:Chrysomelidae: Donaciinae): systematics, reconstructed phylogeny andgeographic history. Transaction of the American Entomological Society113: 360-430.

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1990. Reconstructed phylogeny and reclassification of the genera of Donaciinae(Coleoptera: Chrysomelidae). Quaestiones Entomologicae 26: 601-664.

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192 Psyche [Vol. 98

INDEX TO SPECIES, GENUS AND FAMILY GROUP NAMES

aenea, 178aerea, 178aequalis, 174, 176affinis, 171alutacea, 184americana, 186angustipes, 183antillarum, 181assimilis, 173aurichalcea, 173aurifera, 168balli, 168biimpressa, 173, 177binodosa, 170caerulea, 174, 185californica, 185carolina, 184cataractae, 169, 170cazieri, 174chalcea, 171cincticornis, 170,

177, 180, 181claudicans, 183confluenta, 174confusa, 174congener, 184cuprea, 175, 184curticollis, 175dissimilis, 175distincta, 175diversa, 168dives, 169Donacia (Donacia),

180Donacia (Dona-

ciomima), 173Donaciella, 172Donaciini, 172dubia, 168, 169edentata, 181emarginata, 169, 170episcopalis, 185femoralis, 169

flagellata, 186flavipennis, 169flavipes, 169, 171flohri, 186floridae, 182frosti, 169fulgens, 175, 178fulvipes, 169gentilis, 169germari, 169glabrata, 173harrisii, 172hirticollis, 175hypoleuca, 182idola, 169indica, 169juncina, 170jucunda, 171kirbyi, 171!iebecki, 176limonia, 177lodingi, 171longicollis, 170lucida, 180, 181Macroplea, 186magistrigata, 178magnifica, 177megacornis, 177megalocera, 177megatocera, 177melsheimeri, 186metallica, 169militaris, 182minnesotensis, 186minor, 185, 186nana, 169Neohaemonia, 186neomexicana, 170nigricornis, 186nitida, 170, 171notmani, 170occidentalis, 175pacifica, 170

pallipes, 176palmata, 182, 183parva, 169parvidens, 183piscatrix, 183, 184Plateumarini, 168Plateumaris, 168Poecilocera, 172porosicollis, 178proxima, 174, 185pubescens, 178pubicollis, 172pulchella, 181pusilla, 170, 171pyritosa, 171quadricollis, 175,

178robusta, 171rudicollis, 176rufa, 170, 171,179rufescens, 185rufipennis, 180rugifrons, 170rugosa, 178rutila, 179serricauda, 169schaefferi, 170, 171shoemakeri, 169, 171Sominella, 172subtilis, 174, 175,

178sulcicollis, 171tenuis, 181texana, 185torosa, 174tryphera, 181tuberculata, 171,179tuberculifrons, 179vermiculata, 170vicina, 179wallisi, 169

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