· 2017-04-18 · micronesica 35-36:456-513. 2003 the crustaceans and pycnogonids of the mariana...

Micronesica 35-36:456-513. 2003

The crustaceans and pycnogonids of the Mariana Islands

GUSTAV PAULAY1

Marine Laboratory University of Guam

Mangilao, Guam 96923 USA

ROY KROPP

Battelle Marine Sciences Laboratory1529 West Sequim Bay RoadSequim WA 98382-9099 USA

PETER K. L. NG

Raffles Museum of Biodiversity ResearchDepartment of Biological SciencesNational University of SingaporeKent Ridge, Singapore 119260

Republic of Singapore

LUCIUS G. ELDREDGE

Pacific Science AssociationBishop Museum1525 Bernice St.

Honolulu HI 96817 USA

Abstract—The crustacean and pycnogonid fauna of the Mariana Islandsis reviewed, and 839 crustacean and 15 pycnogonid species aredocumented from the archipelago based on literature records and newcollections, including 272 new records. Voucher specimens are listed for615 and photographic records (see http://www.flmnh.ufl.edu/reefs) for358 species. The bulk of the fauna is marine, including 12 terrestrial and12 freshwater decapods with marine larvae. Three copepods and 6cladocerans comprise the known freshwater fauna, and 25 peracarids arecurrently documented on land. Coverage reflects a taxonomically uneveneffort, and is strongly biased toward macrocrustaceans, with decapodsaccounting for 84% and crabs for 50% of the recorded marine crustaceandiversity.

IntroductionThe Mariana Islands form a chain of 15 major volcanic and limestone islands

bordering the western margin of the Pacific plate between 13 and 21oN latitude 1Current address: Florida Museum of Natural History, Gainesville FL 32611-7800 USA, email:

[email protected]

Paulay et al.: Crustaceans and pycnogonids of the Marianas 457

(see Paulay 2003 for a more detailed description of the area). The archipelago isa classic island arc, formed by volcanic eruptions initiated by the subduction ofthe Pacific plate under the Philippine plate, on which the islands lie. The islandgroup is composed of two geologically distinct chains, the volcanically-inactiveand older islands of the Southern Mariana back-arc, extending from Guam toFarallon de Medinilla, and the volcanically-active, younger islands of theNorthern Mariana fore-arc, extending from Anatahan to Uracas (Fig. 1). Thisgeological separation should not be confused with the political division of thearchipelago into the U.S. Territory of Guam, and the Commonwealth of theNorthern Mariana Islands (CNMI) comprising all islands north of Guam. Thus,Rota, Tinian, Aguijan, Saipan and Farallon de Medinilla are part of the SouthernMariana back-arc but are under the jurisdiction of the CNMI.

The Southern Marianas were initiated ~43 Ma when the Pacific plateabruptly shifted spreading from NW to WNW. The archipelago has had acomplex geological history since. The southern islands are composites ofsubmarine and subaerial volcanics and marine sedimentary formations raised byuplift. Volcanic shorelines are uncommon while karstic shores of mostlyMiocene to Pleistocene limestones dominate. Limestone cliffs line large tracks ofshore, and reefs are generally narrow. Large sections of especially windwardshores lack reef protection, with steep fore reefs abuting eroded cliffs, oftenfronted by supratidal limestone benches. Other areas have narrow, largelyintertidal reef flats or shallow, moderately wide (100-1000 m) fringing reefs.Deep moat and lagoonal habitats are uncommon, the only significant ones beingApra Harbor and Cocos lagoon on Guam and Tanapag-Garapan lagoon onSaipan. Only Apra Harbor has substantial lagoonal habitats >10 m deep. Mangalsare consequently poorly developed, and have undergone considerable reductionsince the arrival of humans (Amesbury 1999). The main remaining mangals arein inner Apra Harbor and along the SW shore of Guam.

Subtidal habitats are dominated by oligotrophic reefs with interspersed sandpockets, with finer mobile sediments largely confined to small estuaries, ApraHarbor, and deep water. The fauna of soft bottoms remains poorly studied. Thekarstic limestones bordering and underlying reefs are riddled by caves andcaverns both above and below water. Although marine caverns and crevices arefairly common, they are generally of limited horizontal extent. Large blocksfallen from cliff faces provide additional wall and crevice habitats. Rubble fieldsare also common, and have been the focus of considerable sampling that revealeda rich crustacean community, including many unusual, troglobitic forms livingdeep in rubble (e.g. Ng 2002b, Ng & Ng 2003, Ng & Takeda 2003).

The Northern Marianas are presently uninhabited, active, young volcanoes.They essentially lack reefs, having rocky shore and subtidal habitats with coralcommunities and incipient reefs growing on igneous outcrops and rubble.Consequently there are no lagoonal habitats except for the protected crater ofMaug. Although poorly known, the biota of the Northern Marianas appears to belargely a less diverse version of that of the Southern Marianas, although some

458 Micronesica 35-36, 2003

species are clearly limited to the northern islands, likely specialized to uniquelynorthern habitats.

Figure 1. Map of the Mariana Islands.


CARCINOLOGICAL HISTORYThe first recorded collection of crustaceans from the Marianas was by Quoy

& Gaimard (1824), who described the land crab Thelphusa rotunda (nowDiscoplax rotundum) and listed 5 additional species from Guam. In February1900 the U.S.F.S. “Albatross” Expedition called on Guam, but only a single crabspecies (Ocypode ceratophthalma) was recorded from their visit (Rathbun 1907).William Edwin Safford, Assistant Botanist for the Department of Agriculture andLieutenant U.S. Navy, was stationed on Guam between August 1899 to August1900. He collected a number of crustaceans and wrote short notes on the crabs(Safford 1905). Alvin Seale of the U.S. War Department, spent several monthson Guam in 1900, reported on the birds and fishes (Seale 1901), and depositedmore than 30 lots of crustaceans in the Bishop Museum. Hans GeorgeHornbostel, Lieutenant U.S. Naval Hospital, archaeologist, and collector for theBishop Museum was on Guam between April 1922 and June 1928. He wroteprimarily on archaeology, but also collected nearly 100 lots of crustaceans for theBishop Museum. Naturalist Edwin H. Bryan Jr. collected extensively on Guam in1936, his collections were also deposited at the Bishop Museum. In the wake ofWorld War II Micronesia received considerable attention from the U.S. During1951-1954, the U.S. Army Corps of Engineers and U.S. Geological Surveyconducted field work on Guam for the “Military Geology of Guam”, headed byKenneth O. Emery. Substantial collections were made by several visitors at thistime and deposited at the USNM; these served as the basis of subsequent studieson the marine biota (e.g. Stephenson & Rees 1967). A similar effort headed byPreston Cloud Jr. on Saipan in 1949 included an intensive survey of the marinebiota. Cloud (1959) recorded 125 crustaceans, most identified to species by theauthorities of the time, several of whom published papers based largely on thiscollection (Banner 1956, Holthuis 1953, 1981). Cloud’s specimens, deposited atthe USNM, have been the source of numerous Saipan records in subsequenttaxonomic revisions as well (e.g. Chace & Bruce 1993, Clark & Galil 1993,Castro 2003).

Carcinological studies in the Marianas gained considerable momentum withthe establishment of the University of Guam Marine Laboratory (UOGML) in1970. Lu Eldredge was involved in a number of surveys focused both on shallowand deep water crustaceans, trained several M.S. students in carcinology,distributed material to specialists, and hosted visiting carcinologists. Deep-watermarine resource surveys included an assessment of deep-water shrimp resourcesaround Guam, funded by Sea Grant between 1973-1976 (Wilder 1977), and theResource Assessment Investigation of the Mariana Archipelago (RAIOMA) ofthe National Marine Fisheries Service using the NOAA ship Townsend-Cromwell between May 1982 and June 1984 (Polovina et al. 1985). These effortshave led to a number of publications on the systematics and fisheries potential ofdeep-water crustaceans (Eldredge 1976, 1980, Wilder 1977, Moffitt 1983; King1987, Williams & Moffitt 1991, Williams & Eldredge 1994). A visit by thesubmersible Alvin to the hydrothermal vents of the Mariana back arc resulted in


several papers on the crustacean biota of this isolated vent system (Hessler &Martin 1989, Humes 1990, Hessler & Lonsdale 1991, Child 1989), while asurvey of the much shallower, hydrothermally-active volcano at Esmerelda Bankled to the record of two unusual decapods (Türkay & Sakai 1995).

Several graduate students have pursued carcinological theses at theUniversity of Guam (Tseng 1972, Wortman 1976, Dickinson 1977, Wooster1979, Kropp 1981, Donaldson 1981, Grimm 1983, Camacho 1997, Leberer1997). Wooster’s study culminated in a paper on the Calcinus fauna of Guam(Wooster 1982), while Leberer went on to review the taxonomy of the island’satyids (Leberer & Cai 2003). Roy Kropp and Jane Dominguez undertook anintensive collecting survey of the shallow water crustaceans of Guam; their largecollection now at the USNM has been the basis of numerous publications (e.g.papers by Kropp and co-authors, Child 1991, Ahyong & Erdmann 2003, Tan &Ng 2003b).

During the 1990s the Gustav Paulay’s laboratory has been active in marinebiodiversity surveys on Guam. Material was collected during several organizedsurveys (see Paulay 2003), as well as incidentally by students, faculty, staff andfriends of the University of Guam Marine Laboratory. Many specimensgenerated by these surveys have been sent to specialists for identification andstudy (e.g. McLay 2001, McLaughlin 2002a-b, Ng, 2002a-c). Harry Conley hascontributed an especially large number of noteworthy specimens, collected fromdeep within rubble, a microhabitat whose exploration he pioneered. Hiscollections have led to the discovery of numerous interesting new taxa (e.g.Kensley 2003, Ng et al. papers in this volume).

The Chiba Museum undertook an expedition to the Northern Marianas in1992 and published a comprehensive checklist of their findings in a specialvolume, with several papers dealing with crustaceans (e.g. Asakura et al. 1994,Hayashi et al. 1994, Takeda et al. 1994, Asakura & Tachikawa 2000). Othervisiting carcinologists also published on the fauna (e.g. Belk 1973, Odinetz 1984,Weissleader et al. 1989, Foster 1990, Asami & Yamaguchi 2001, Castro 2003).A visit by the third author and Chia-Hsiang Wang in 2000 led to a review of thecrabs in the University of Guam’s Invertebrate Collection, an effort that hasalready led to several papers that appear in this volume as well as many of theidentifications listed in our checklist. A subsequent collecting visit by their teamin 2001 together with ongoing studies of previously collected material (especiallyof the Grapsidae, Xanthidae and Parthenopidae) will undoubtedly lead tonumerous studies on Guam’s crustaceans in the future.

MethodsThis paper is based on a compilation of records from the literature and

identifications made on new collections. Both efforts are ongoing and certainlyincomplete. Although we strove to check all papers dealing with the Micronesianfauna as well as major taxonomic revisions, additional records of the biota


undoubtedly appear in papers we did not consult. We did not include recordslisted in the voluminous gray literature of environmental surveys published onthe area, as such records are usually difficult or impossible to verify. We haveupdated the nomenclature of literature records as much as possible; changes aregiven as notes in the Appendix. Numerous species listed here have not beenpreviously recorded and comprise new additions to the Marianas’ fauna. Theseare marked by the name of the identifier in the Appendix and identified with aspecimen voucher and/or photograph.

New collections focused largely on stomatopods, decapods, and barnacles.Even among these groups efforts have been uneven, with larger species, crabs,hermit crabs, axiids, and lobsters receiving the greatest attention. Much of thismaterial is housed at the Florida Museum of Natural History, University ofFlorida (UF), U.S. National Museum of Natural History, Smithsonian Institution(USNM), Zoological Reference Collection, Raffles Museum, National Universityof Singapore (ZRC), and the University of Guam (UGI). Most newly collectedspecies were photo documented. Photo vouchers cited in the Appendix areavailable online at http://www.flmnh.ufl.edu/reefs and in the Marine Biodiversityof Guam CD-ROM co-publication.

In addition to the species listed here, numerous remain to be identified andthe identification of many listed species improved. Work is ongoing on these andadditions and modifications will be posted on the website as they are published.Major ongoing efforts include studies on barnacles (W. Newman, A. Ross),paguroids (P. McLaughlin), galatheoids (R. Kropp), and brachyurans (B. Galil,C. McLay, P. Ng et al.).

For the purposes of this checklist we define marine crustaceans as speciesthat have at least one phase of their life cycle in the sea. These include allterrestrial and freshwater decapods in the Marianas, as all are known or presumedto have marine larvae. Truly terrestrial and freshwater crustaceans are listedbelow the marine taxa, and include isopods, talitrid amphipods, copepods, andcladocerans.

Results and DiscussionWe documented 839 crustacean and 15 pycnogonid species from the Mariana

Islands based on literature records and new collections supported by voucherspecimens and/or photographic records (Tables 1, 2, Appendix). 483 species arebased at least in part on literature records, 272 are recorded for the first time inthis paper, while an additional 99 are new records from our recent biodiversitysurveys that have been further documented in other papers in this volume, and inpapers published recently or in press elsewhere.

358 species are photodocumented based on living or fresh specimens. 663species are recorded from Guam, while the remainder only from the CNMI. Mostof the species currently known only from the CNMI likely also occur on Guam,but remain undocumented there because of limited collecting or different


taxonomic emphasis. For example the alpheids of Saipan were reviewed in asubstantial paper by Banner (1956), while those of Guam remain virtuallyunstudied.

Table 1. Diversity of marine arthropod taxa

Taxon Number of spp.

Copepoda 4Ostracoda 45Amphipoda 17Isopoda 3Stomatopoda 36Stenopodidea 1Penaeoidea 4Caridea 139Palinura 11Astacidea 2Thalassinidea 16Anomura 94Brachyura 405Ascothoracida 1Cirripedia 27Pycnogonida 15

TOTAL 820

The bulk (805 species) of the crustaceans are marine, including 12 species ofland and hermit crabs with marine larvae and 12 species of anadromous decapods(Table 2). Three copepods and 6 cladocerans comprise the known freshwaterfauna, and 25 species of peracarids are currently documented on land.

Taxonomic coverage is highly uneven, with macrocrustaceans comprisingthe vast majority of the records (Table 1). Thus decapods comprise 84% (674species) and crabs 50% (405 species) of the marine crustaceans recorded. Othercrustacean taxa remain poorly documented with the exception of stomatopods(36 species) and barnacles (27 species). For example only 20 peracarids and 4copepods are currently recorded in the marine fauna. Even among decapods,coverage varies substantially among families, reflecting taxonomically focusedcollecting and study. Thus brachyurans have received much greater attentionoverall than carideans. Cryptochirids and porcellanids were intensively studiedby Kropp, and are especially well known (see papers by Kropp et al.). The mostspecies rich families are the Xanthidae (114 species), Alpheidae (61 species),Portunidae (55 species), Diogenidae (40 species), Palaemonidae (35 species), andCryptochiridae (28 species).

Even groups that have been the focus of study harbor much undocumenteddiversity. Thus the known paguroid diversity of Guam has more than doubled (31vs. 69 species) in the two decades following Wooster’s (1979, 1982) thesis on thegroup, even though hermit crabs did not receive much specialized attentionduring this period.


It is difficult to estimate the actual diversity of marine arthropods in Guam orthe Marianas, as even macrocrustaceans are but modestly known. Crabs areamong the best studied overall, but the rapid rate of new encounters and backlogof unidentified material implies that perhaps not more than half of the crabdiversity is currently documented. Decapod diversity as a whole will certainlymuch more than double with additional study and the diversity ofmicrocrustaceans cannot be reliably estimated at present.

Table 2. Arthropod diversity by habitat

Habitat Number of spp.

Shallow (<60m) marine 747Deep (>60m) marine 49Anadromous 12Terrestrial with marine larvae 12Freshwater 9Terrestrial 25

AcknowledgementsWe thank the students, faculty and staff at the UOG Marine Lab and

naturalists from throughout the Marianas, especially Tim Allen, Scott Bauman,Ian Elliott, Bruce Henke, Lisa Kirkendale, Chris Meyer, Rob Myers, RaphaelRitson-Williams, Teina Rongo, John Starmer, and Linda Ward for bringingcrustacean specimens, records, and photos to our attention. Special thanks toHarry Conley for his indefatigable collecting over the years. We thank ShaneAhyong, Kiyo Asami, Warren Blow, Yixiong Cai, Peter Castro, Tin-Yam Chan,Bill Dall, Mark Erdmann, Charles Fransen, Bella Galil, Yoshi Hisatsune, BrianKensley, Hung-Chang Liu, Patsy McLaughlin, Colin McLay, Junji Okuno, JohnStarmer, Masatsune Takeda, and Chia-Hsiang Wang for help with identifications.We thank Rob Myers, Tim Allen, Hung-Chang Liu, and Ian Elliott forpermission to use their photographs of species not otherwise recorded fromGuam, and Alain Crosnier and Joseph Poupin for comments on the manuscript.Funding by the U.S. Dept. of Defense at COMNAVMARIANAS and NOAA SeaGrant (SG-NIS-35) to Gustav Paulay for biodiversity surveys is gratefullyacknowledged. Contribution 485 of the University of Guam Marine Laboratory.

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Received 3 March 2002

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e fr

om G

uam

unl

ess

pare

nthe

tical

ly in

dica

ted

othe

rwis

e; s

ee u

nder

“Is

land

” be

low

for

key

to i

slan

d na

me

abbr

evia

tions

.H

ab: H

abita

t: 1:

mar

ine,

0-6

0m, 2

: mar

ine,

60-

200m

, 3: m

arin

e, >

200m

, FW

: fre

shw

ater

, T: t

erre

stria

lIs

land

: A

g: A

grih

an, A

l: A

lam

agan

, An:

Ana

taha

n, A

r: A

raka

ne R

eef,

As:

Asu

ncio

n, C

NM

I: C

omm

onw

ealth

of

the

Nor

ther

n M

aria

na I

slan

ds,

isla

nd u

nspe

cifie

d, E

: Esm

eral

da B

ank,

G: G

uam

, Gu:

Gug

uan,

M: M

aug,

MS:

Mar

iana

Isl

ands

, isl

and

unsp

ecifi

ed, M

TV: M

aria

na T

roug

hve

nts,

P: P

agan

, R: R

ota,

S: S

aipa

n, S

a: S

arig

an, T

: Tin

ian,

U: U

raca

s.R

efs:

Ref

eren

ces l

iste

d at

end

of A

ppen

dix

Not

es: l

iste

d at

end

of A

ppen

dix

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndM

AR

INE

TAX

AC

OPE

POD

A: D

IRIV

ULT

IDA

EC

hasm

atop

ontiu

s the

scal

us H

umes

, 199

01,

23

MTV

Styg

iopo

ntiu

s pec

tinat

us H

umes

, 198

72

3M

TVSt

ygio

pont

ius s

tabi

litus

Hum

es, 1

990

1, 2

3M

TVC

OPE

POD

A: E

RG

ASI

LID

AE

Paeo

node

s sub

viri

dis E

l-Ras

hidy

& B

oxsh

all,

2001

BM

NH

199

9.15

29 (h

olot

ype)

140

1G

OST

RA

CO

DA

: PO

DO

CO

PID

ABa

irdo

ppila

ta c

rate

rico

la M

addo

cks,

1969

31

GBi

shop

ina

timor

ensi

s (Fy

an, 1

916)

31

GBo

sase

lla c

f. m

acro

loba

(Hu,

198

1)3

1G

Byth

ocyp

ris a

ff. p

rola

ta M

addo

cks,

1969

31

GC

allis

tocy

ther

e ar

cuat

a B

onad

uce

et a

l., 1

980

31

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndC

allis

tocy

ther

e cf

. kei

jii (H

artm

ann,

197

8)3

1G

Cau

dite

s exm

outh

ensi

s Har

tman

n, 1

978

31

GC

leto

cyth

erei

s ras

trom

argi

nata

(Bra

dy, 1

880)

31

GC

ythe

rello

ida

sp. B

3

1G

Cyt

here

lloid

a sp

. E

31

GH

emic

ythe

re sp

. B

31

GH

emic

ythe

rura

sp. 1

3

11

GH

erm

anite

s del

toid

ea (B

rady

, 189

0)3

1G

Her

man

ites t

rans

ocea

nica

Tee

ter,

1975

31

GK

eijia

fove

ata

(Har

tman

n, 1

978)

31

GLo

xoco

ncha

her

onis

land

ensi

s Har

tman

n, 1

981

31

GLo

xoco

ncha

hua

hine

ensi

s Har

tman

n, 1

984

31

GLo

xoco

ncha

sp. 1

3

21

GLo

xoco

ncha

turn

erae

Wat

son,

198

83

1G

Loxo

conc

hella

ano

mal

a (B

rady

, 188

0)3

1G

Loxo

conc

hella

cf.

mal

aysi

ana

Kei

j, 19

783

1G

Mor

khov

enia

inco

nspi

cua

(Bra

dy, 1

880)

31

GM

ydin

obai

rdia

schy

roco

ncha

(Mad

dock

s, 19

69)

31

GN

eone

side

a cf

. are

nige

ra M

addo

cks,

1969

31

GN

eone

side

a le

ntifi

la H

artm

ann,

198

43

1G

Neo

nesi

dea

schu

lzi (

Har

tman

n, 1

964)

31

GO

rnat

oleb

eris

sp. 1

3

31

GPa

racy

ther

idea

cf.

diss

ecta

Hu,

198

33

1G

Para

cyth

erid

ea ts

chop

pi B

old,

194

63

1G

Para

nesi

dea

cf. a

lgic

ola

Mad

dock

s, 19

693

1G

Para

nesi

dea

sp. A

3

1G

Para

nesi

dea

spon

gico

la M

addo

cks,

1969

31

GPo

ntic

ocyt

here

s ich

thyo

derm

a (B

rady

, 189

0)3

1G

Prop

onto

cypr

is sp

. A

31

GPr

opon

tocy

pris

sp. B

3

1G

Pter

obai

rdia

mad

dock

sae

McK

enzi

e &

Kei

j, 19

773

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndRa

dim

ella

sp. A

3

1G

Rena

udcy

pris

? na

tans

Har

tman

n, 1

984

31

GSa

ipan

etta

clo

udi (

McK

enzi

e, 1

967)

USN

M 1

1343

7 (h

olot

ype,

S)

132

681

SSe

mic

ythe

rura

cf.

cryp

tifer

a (B

rady

, 188

0)3

1G

Tane

lla c

f. gr

acili

s Kin

gma,

194

83

1G

Trie

belin

a br

adyi

Trie

bel,

1948

31

GTr

iebe

lina

sert

ata

Trie

bel,

1948

31

GTr

iebe

lina

sp. 1

3

41

GXe

stol

eber

is sp

p.

31

GA

MPH

IPO

DA

: AM

PHIL

OC

HID

AE

Gita

nops

is sp

. 1

41

MA

MPH

IPO

DA

: AM

PITO

IDA

EAm

pith

oe ra

mon

di A

udou

in, 1

826

51

SAm

pith

oe sp

. 1

41

MAm

pith

oe sp

. 2

41

Gu

Ampi

thoe

sp. 3

4

1M

Gru

bia

filos

a (S

avig

ny, 1

826)

51

SPa

radu

sa sp

. 1

41

MA

MPH

IPO

DA

: CO

LOPH

IID

AE

Gam

mar

opsi

s sp.

1

41

MA

MPH

IPO

DA

: GA

MM

AR

IDA

EC

erad

ocus

rubr

omac

ulat

us (S

timps

on, 1

855)

51

SEr

asm

ops s

p. 1

4

1M

Mae

ra in

sign

is (C

hevr

eux,

190

1)5

1S

Mae

ra sp

. 1

41

MA

MPH

IPO

DA

: HY

ALI

DA

EH

yale

sp. 1

4

1M

Hya

le sp

. 2

41

MH

yale

sp. 3

4

1M

AM

PHIP

OD

A: I

SCH

YR

OC

ERID

AE

Ambi

chol

este

s squ

amife

rus (

Just

, 198

4)U

GI (

para

type

)6

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndA

MPH

IPO

DA

: LEU

CO

THO

IDA

ELe

ucot

hoe

sp. 1

4

1M

ISO

POD

A: C

YM

OTH

OID

AE

Cer

atot

hoa

angu

lata

(Ric

hard

son,

191

0)U

SNPC

088

995

71

GC

ymot

hoa

bych

owsk

yi A

vdee

v, 1

979

USN

PC 0

8899

6 7

1G

Elth

usa

rayn

audi

i (M

ilne-

Edw

ards

, 184

0)U

SNPC

088

997

71

GST

OM

ATO

POD

A: B

ATH

YSQ

UIL

LID

AE

Bath

ysqu

illa

mic

rops

(Man

ning

, 196

1)U

SNM

228

146

8,9

3G

STO

MA

TOPO

DA

: CO

RO

NID

IDA

EM

orte

nsen

enus

pau

lay

Ahy

ong

& E

rdm

ann,

200

3A

M P

(hol

otyp

e, M

) 9

1M

Neo

coro

nida

mar

tens

i Man

ning

, 197

8A

M P

9

1M

STO

MA

TOPO

DA

: EU

RY

SQU

ILLI

DA

EM

anni

ngia

zehn

tner

i Man

ning

, 197

4U

SNM

307

137

91

GST

OM

ATO

POD

A: G

ON

OD

AC

TYLI

DA

EG

onod

acty

lace

us fa

lcat

us (F

orss

kål,

1775

)U

F 26

5, U

SNM

304

400

5, 9

, 10

GP6

94-1

45

1G

, SG

onod

acty

lace

us te

rnat

ensi

s (de

Man

, 190

2)U

SNM

304

416

91

GG

onod

acty

lellu

s erd

man

ni A

hyon

g, 2

001

USN

M 3

0440

3,

USN

M 3

0465

3 (R

) 9

1G

, R

Gon

odac

tyle

llus e

spin

osus

(Bor

rada

ile 1

898)

USN

M 3

0444

59

1G

Gon

odac

tyle

llus m

icro

nesi

cus (

Man

ning

, 197

1)U

SNM

304

408

91

GG

onod

acty

lus c

hild

i Man

ning

, 197

1U

SNM

304

444

9R

FM1

GG

onod

acty

lus c

hira

gra

(Fab

riciu

s, 17

81)

UF

239,

USN

M 3

0448

95,

9, 1

0G

P359

-21

1G

, SG

onod

acty

lus p

laty

som

a W

ood-

Mas

on, 1

895

UF

233,

USN

M 3

0445

59,

11

GP4

64-1

06

1G

Gon

odac

tylu

s sm

ithii

Poco

ck, 1

893

UF

235,

USN

M 3

0445

3 9

GP4

04-3

01

GH

oplo

squi

lla sa

id E

rdm

ann

& M

anni

ng, 1

998

USN

M 3

0608

0 9

1G

STO

MA

TOPO

DA

: IN

DO

SQU

ILLI

DA

EIn

dosq

uilla

man

ihin

ei In

gle

& M

erre

tt, 1

971

USN

M 2

3347

7,

USN

M 2

2841

7 (P

) 8,

93

G, P

STO

MA

TOPO

DA

: LY

SIO

SQU

ILLI

DA

ELy

sios

quill

ina

mac

ulat

a (F

abric

ius,

1793

)U

F 23

2, U

SNM

304

425

91

GLy

sios

quill

ina

sulc

ata

(Man

ning

, 197

8)

12R

FM 2

261

G, S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndST

OM

ATO

POD

A: N

AN

NO

SQU

ILLI

DA

EPu

llosq

uilla

lito

ralis

(Mic

hel &

Man

ning

, 197

1)U

SNM

304

502

91

GPu

llosq

uilla

mal

ayen

sis (

Man

ning

, 196

8)U

SNM

304

529

91

GPu

llosq

uilla

thom

assi

ni M

anni

ng, 1

978

USN

M 3

0450

0 9

1G

STO

MA

TOPO

DA

: OD

ON

TOD

AC

TYLI

DA

EO

dont

odac

tylu

s scy

llaru

s (Li

nnae

us, 1

758)

USN

M 3

0443

5 9

RFM

98a

1G

STO

MA

TOPO

DA

: PR

OTO

SQU

ILLI

DA

EC

hori

squi

lla g

yros

a (O

dhne

r, 19

23)

USN

M 3

0444

8 9

RK

1

GC

hori

squi

lla k

ropp

i Ahy

ong

& E

rdm

ann,

200

3U

SNM

304

499

(hol

otyp

e)9

1G

Echi

nosq

uilla

gue

rini

(Whi

te, 1

861)

USN

M 3

0442

49

RK

1

GH

apto

squi

lla g

lypt

ocer

cus (

Woo

d-M

ason

, 187

5)U

F 23

7, U

SNM

120

271

9G

P467

-33

1G

Hap

tosq

uilla

tane

nsis

(Fuk

uda,

191

1)N

eoty

pe: U

SNM

304

475

91

GST

OM

ATO

POD

A: P

SEU

DO

SQU

ILLI

DA

EPs

eudo

squi

lla c

iliat

a (F

abric

ius,

1787

)U

SNM

304

428

9R

FM 2

06b

1G

Pseu

dosq

uilla

orn

ata

Mie

rs, 1

880

5, 1

01

SPs

eudo

squi

llana

rich

eri (

Moo

sa, 1

991)

USN

M 3

0443

2 9

1G

Raou

lser

enea

hie

rogl

yphi

ca (M

anni

ng, 1

972)

USN

M 3

0442

79

1G

Raou

lser

enea

kom

aii (

Moo

sa, 1

991)

UF

236,

USN

M 3

0443

0 9

GP6

50-3

21

G, A

s, M

,A

gRa

ouls

eren

ea o

xyrh

ynch

a (B

orra

daile

, 189

8)U

SNM

304

426

91

GST

OM

ATO

POD

A: S

QU

ILLI

DA

ELe

ptos

quill

a sc

hmel

tzii

(A. M

ilne-

Edw

ards

, 187

3)U

SNM

304

446

91

GO

rato

squi

lla fa

bric

ii (H

olth

uis,

1941

)U

SNM

304

447

9G

P396

-30

1G

Parv

isqu

illa

dom

ingu

ez A

hyon

g &

Erd

man

n, 2

003

USN

M 3

0452

5 (h

olot

ype)

91

GST

OM

ATO

POD

A: T

AK

UID

AE

Mes

actu

rus d

icru

rus K

ropp

& D

omin

guez

, 199

0U

SNM

244

236

(hol

otyp

e),

USN

M 2

4423

8 (p

arat

ype,

S)

9, 1

3, 1

41

G, S

, M

STEN

OPO

DID

EA: S

TEN

OPO

DID

AE

Sten

opus

his

pidu

s (O

livie

r, 18

11)

USN

M 1

7159

8, B

PBM

287

35,

10,

15

GP5

16-2

31

G, S

, Gu,

P,

M

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndPE

NA

EOID

EA: P

ENA

EID

AE

Met

apen

aeop

sis c

omm

ensa

lis B

orra

daile

, 189

8U

F 89

1D

GP2

64-1

41

GPe

naeu

s mon

odon

Fab

riciu

s, 17

9816

71

GLi

tope

naeu

s sty

liros

tris

(Stim

pson

, 187

4)16

7, 6

91

GLi

tope

naeu

s van

nam

ei (B

oone

, 193

1)17

71

GC

AR

IDEA

: ALP

HEI

DA

EAl

pheo

psis

dia

bolu

s Ban

ner,

1956

USN

M (h

olot

ype,

S)

188

1S

Alph

eops

is e

qual

is C

outiè

re, 1

896

NN

M D

-480

87 (A

s)

5, 1

81

As,

SAl

pheo

psis

tetr

arth

ri B

anne

r, 19

56U

SNM

(hol

otyp

e, S

)18

1S

Alph

eus b

iden

s? (O

livie

r, 18

11)

151

Gu

Alph

eus b

rady

pus C

outiè

re, 1

905

181

SAl

pheu

s bre

vipe

s Stim

pson

, 186

0U

SNM

5,

18

1S

Alph

eus b

ucep

halu

s Cou

tière

, 190

5U

SNM

5, 1

81

SAl

pheu

s cf.

crin

itus D

ana,

185

215

1A

gAl

pheu

s cha

mor

ro B

anne

r, 19

56U

SNM

(hol

otyp

e, S

) 18

1S

Alph

eus c

oetiv

ensi

s Cou

tière

, 190

818

91

SAl

pheu

s col

lum

ianu

s Stim

pson

, 186

0N

NM

D-4

8070

(As)

, U

GI 6

629

(CN

MI)

5, 1

810

1A

s, S

Alph

eus c

rock

eri (

Arm

stro

ng, 1

941)

NN

M D

-480

68 (A

s),

UG

I 662

8 (C

NM

I)18

111

As,

S

Alph

eus d

eute

ropu

s Hilg

endo

rf, 1

878

UF

829,

USN

M 2

3340

1 5,

18

GP7

58-1

11

G, S

Alph

eus d

iade

ma

Dan

a, 1

852

NN

M-D

4807

6 (A

g)

5, 1

81

Ag,

SAl

pheu

s dol

erus

Ban

ner,

1956

USN

M (h

olot

ype,

S)

181

SAl

pheu

s eul

imen

e de

Man

, 191

1U

SNM

5, 1

81

SAl

pheu

s fro

ntal

is H

. Miln

e Ed

war

ds, 1

837

UF

825,

NN

M D

-480

74 (A

l) 15

, 18

GP4

54-1

71

G, A

l, S,

P,

MAl

pheu

s gra

cilip

es S

timps

on, 1

860

USN

M5,

18

1S

Alph

eus g

raci

lis H

elle

r, 18

6218

121

SAl

pheu

s haa

ni O

rtman

n, 1

890

51

SAl

pheu

s idi

oche

les C

outiè

re, 1

905

USN

M 2

5598

0 5,

18,

19

131

G, S

Alph

eus l

adro

nis B

anne

r, 19

56U

SNM

(hol

otyp

e, S

)18

1S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndAl

pheu

s lep

toch

irus

Cou

tière

, 190

518

1S

Alph

eus l

ottin

i Gué

rin, 1

829

UF

939

(Al),

UF

985

(As)

, U

F 82

8, U

GI 6

627

(CN

MI)

,N

NM

-D48

064

(Ag)

, BPB

M

5, 1

5, 1

8G

P784

-26

141

G, A

g, A

l,A

n, A

s, S

Alph

eus l

utin

i Cou

tière

, 190

5U

SNM

5, 1

81

SAl

pheu

s mal

leod

igitu

s (B

ate,

188

8)U

F 82

6, U

SNM

223

423

5, 1

8, 1

915

1G

, SAl

pheu

s mic

rost

ylus

(Bat

e, 1

888)

USN

M5,

18

1S

Alph

eus o

ahue

nsis

(Ban

ner,

1953

)18

1S

Alph

eus o

beso

man

us D

ana,

185

2U

SNM

255

979

15, 1

91

G, M

Alph

eus o

valic

eps C

outiè

re, 1

905

181

SAl

pheu

s pac

hych

irus

Stim

pson

, 186

0U

SNM

5, 1

81

SAl

pheu

s pac

ificu

s Dan

a, 1

852

15, 1

81

S, G

u, A

l,M

Alph

eus p

arac

rini

tus M

iers

, 188

1U

SNM

5,

18

161

SAl

pheu

s par

viro

stri

s Dan

a, 1

852

USN

M5,

18

1S

Alph

eus p

ercy

i Cou

tière

, 190

815

171

As

Alph

eus p

erpl

exus

Ban

ner,

1956

USN

M (h

olot

ype,

S)

18, 2

01

G, S

Alph

eus r

apax

Fab

riciu

s, 17

98B

PBM

S68

5018

1G

Alph

eus s

tanl

eyi C

outiè

re, 1

908

1819

1S

Alph

eus s

taph

ylia

nus C

outiè

re, 1

908

UG

I 662

2 (C

NM

I)F

1C

NM

IAl

pheu

s sul

catu

s Kin

gsle

y, 1

878

NN

M D

-480

75 (A

l)F

1A

lAl

pheu

s sup

erci

liari

s Cou

tière

, 190

5U

F 96

2 (A

g), U

F 96

1 (A

s), U

F96

0 (M

), N

NM

D-4

8078

(As)

F

1A

s, M

, Ag

Alph

eus s

p. 1

(n. s

p.)

181

SAl

pheu

s sp.

2 (n

. sp.

) 18

1S

Alph

eus s

p. 3

(n. s

p.)

181

SAl

pheu

s sp.

4 (n

. sp.

) 18

181

SAr

ete

iphi

anas

sa d

e M

an, 1

910

USN

M5,

18

1S

Atha

nas a

rete

form

is C

outiè

re, 1

903

15, 1

820

1M

, SAt

hana

s djib

outie

nsis

Cou

tière

, 189

7U

SNM

5, 1

81

S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndAt

hana

s dor

salis

(Stim

pson

, 186

1)U

F 96

8 (M

), B

PBM

, N

NM

D-4

8088

(As)

F

1G

, As,

M,

Ag

Atha

nas d

ubiu

s Ban

ner,

1956

USN

M (h

olot

ype,

S),

BPB

M18

1G

, SAu

tom

ate

dolic

ogna

tha

de M

an, 1

888

5, 1

5, 1

821

1A

g, S

Auto

mat

e jo

hnso

ni C

hace

, 195

5U

SNM

18

1S

Met

alph

eus p

arag

raci

lis (C

outiè

re, 1

897)

NN

M D

-480

86 (M

)5,

18

221

M, S

Met

alph

eus r

ostr

atip

es (P

ococ

k, 1

890)

5, 1

5, 1

823

1M

, URa

ciliu

s com

pres

sus P

auls

on, 1

875

151

Ag

Syna

lphe

us tu

mid

oman

us (P

auls

on, 1

875)

151

Gu,

MSy

nalp

heus

anc

eps B

anne

r, 19

56U

SNM

(hol

otyp

e, S

), B

PBM

18

1G

, SSy

nalp

heus

cha

ron

(Hel

ler,

1861

)B

PBM

, NN

M D

-480

84 (A

g)

15, 1

824

1G

, Ag,

M, S

Syna

lphe

us e

xilip

es C

outiè

re, 1

905

USN

M

1825

1S

Syna

lphe

us p

aran

eom

eris

Cou

tière

, 190

5N

NM

D-4

8082

(As)

5,

15,

18

1A

s, S,

MSy

nalp

heus

stim

pson

ii (d

e M

an, 1

888)

NN

M D

-480

85 (A

s)F

GP8

30-3

1G

, As

CA

RID

EA: A

TYID

AE

Ante

cari

dina

laue

nsis

Edm

onds

on, 1

935

UF/

ZRC

21FW

GAt

yoid

a pi

lipes

(New

port,

184

7)U

F 81

221

FWG

Car

idin

a br

achy

dact

yla

(de

Man

, 190

8)U

F/ZR

C21

FWG

Car

idin

a m

erto

ni R

oux,

191

1U

F/ZR

C21

FWG

Car

idin

a ty

pus H

. Miln

e Ed

war

ds, 1

837

UF/

ZRC

21FW

GH

aloc

arid

inid

es sp

. 1 (n

. sp.

) U

F/ZR

C21

FWG

CA

RID

EA: B

RES

ILII

DA

EC

horo

cari

s van

dove

rae

Mar

tin &

Hes

sler

, 199

02,

22

3M

TVD

isci

as m

usic

us H

olth

uis,

1981

USN

M 9

4774

(hol

otyp

e, S

)5,

10,

24

261

SC

AR

IDEA

: GN

ATH

OPH

YLL

IDA

EG

nath

ophy

llum

am

eric

anum

Gué

rin-M

énev

ille,

185

5U

F 45

6, B

PBM

5, 1

0G

P363

-27

1G

, SG

nath

ophy

llum

pre

cipu

um T

itgen

, 198

9U

F 80

8 P

1G

Pycn

ocar

is c

hago

ae B

ruce

, 197

2N

NM

D-4

8092

FG

P415

-18

1G

CA

RID

EA: H

IPPO

LYTI

DA

ETh

inor

a m

aldi

vens

is (B

orra

daile

, 191

5)5,

10

1S

Hip

poly

te e

dmon

dson

i Hay

ashi

, 198

215

1G

u

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndH

ippo

lyte

sp. 1

U

F 81

9F

1G

Lysm

ata

ambo

inen

sis (

de M

an, 1

888)

PR

FM1

GLy

smat

a sp

. 1

UF

884

FG

P778

-36

1G

Parh

ippo

lyte

mis

ticia

(Cla

rk, 1

989)

U

F 80

7P

GP8

26-1

1G

Saro

n m

arm

orat

us (O

livie

r, 18

11)

?UF

457

5, 1

0G

P516

-28

1G

, SSa

ron

negl

ectu

s de

Man

, 190

25,

10

1S

Saro

n sp

. 1

USN

M?

PR

K1

GTh

or a

mbo

inen

sis (

de M

an, 1

888)

UF

821

PR

FM-1

86a

1G

Thor

pas

chal

is (H

elle

r, 18

62)

5, 1

0, 2

51

SC

AR

IDEA

: HY

MEN

OC

ERID

AE

Hym

enoc

era

pict

a D

ana,

185

2 U

F 82

2P

GP6

53-3

11

GC

AR

IDEA

: PA

LAEM

ON

IDA

EBr

achy

carp

us b

iung

uicu

latu

s (Lu

cas,

1846

)U

F 93

5, N

NM

D-4

8096

(As)

F

1G

, As

Con

chod

ytes

mel

eagr

inae

Pet

ers,

1852

NN

M D

-480

95F

1G

Cor

allio

cari

s gra

min

ea (D

ana,

185

2)

UF

842

FG

P821

-37

1G

Cor

allio

cari

s sup

erba

(Dan

a, 1

852)

5, 1

01

SD

acty

loni

a as

cidi

cola

(Bor

rada

ile, 1

898)

NN

M D

-480

6613

41

GEx

opon

toni

a m

alle

atri

x B

ruce

, 198

8N

NM

D-4

8065

F

1G

Har

pilio

psis

bea

upre

sii (

Aud

ouin

, 182

6)15

1M

Har

pilio

psis

dep

ress

a (S

timps

on, 1

860)

151

M, G

uH

arpi

liops

is sp

inig

era

(Ortm

ann,

189

0)N

NM

D-4

8079

(Ag)

F1

Ag

Isch

nopo

nton

ia lo

phos

(Bar

nard

, 196

2)15

1A

gJo

cast

e ja

poni

ca (O

rtman

n, 1

890)

151

Ag

Joca

ste

luci

na (N

obili

, 190

1)5,

10

1S

Lean

der p

lum

osus

Bru

ce, 1

993

PTA

199

91

GLe

ande

r ten

uico

rnis

(Say

, 181

8)U

F 81

8, N

NM

D-4

8093

(S)

FG

P410

-11

1G

, SM

acro

brac

hium

equ

iden

s (D

ana,

185

2)

ZRC

N1

GM

acro

brac

hium

lar (

Fabr

iciu

s, 17

98)

BPB

M-S

117

27, 1

36FW

GM

acro

brac

hium

rose

nber

gii (

de M

an, 1

879)

167

FWG

Mac

robr

achi

um sp

. 1

2627

FWG

Ony

coca

ris q

uadr

atop

htha

lma

(Bal

ss, 1

921)

151

M

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndPa

laem

on d

ebili

s Dan

a, 1

852

281

GPa

laem

onel

la ro

tum

ana

(Bor

rada

ile, 1

898)

5, 1

028

1S

Pala

emon

ella

spin

ulat

a Y

okoy

a, 1

936

151

Ag

Peri

clim

enae

us q

uadr

iden

tatu

s (R

athb

un, 1

906)

5, 1

01

SPe

ricl

imen

aeus

trid

enta

tus (

Mie

rs, 1

884)

10, 2

91

SPe

ricl

imen

ella

spin

ifera

(de

Man

, 190

2)N

NM

D-4

8067

, UG

I 606

45,

10

GP3

63-3

729

1G

, SPe

ricl

imen

es b

revi

carp

alis

(Sch

enke

l, 19

02)

UF

905?

P

GP7

85-1

01

GPe

ricl

imen

es e

lega

ns/g

rand

is(P

auls

on,1

875)

/(Stim

pson

,186

0)

NN

M D

-480

73F

GP5

13-2

31

G

Peri

clim

enes

impe

rato

r (Sc

henk

el, 1

880)

UF

841,

NN

M D

-480

94F

RFM

5-9

81

GPe

ricl

imen

es so

ror N

obili

, 190

4N

NM

D-4

8071

, UF

3031

30G

P739

-18

1G

Peri

clim

enes

sp. 1

(n. s

p. a

ff. h

olth

uisi

) U

F 83

8 F

GP8

40-1

71

GPl

iopo

nton

ia fu

rtiv

a B

ruce

, 197

3U

F 84

3 F

GP7

41-1

91

GPo

nton

iops

is c

oman

thi B

orra

daile

, 191

55,

10,

29

1S

Steg

opon

toni

a co

mm

ensa

lis N

obili

, 190

6P

GP5

18-2

01

GU

roca

ride

lla a

nton

bruu

nii/c

yrto

rhyn

cha

(Bru

ce, 1

967)

/Fu

jino

& M

iyak

e, 1

969

UF

813

OG

P413

-25

1G

Vir o

rien

talis

(Dan

a, 1

852)

5, 1

0, 2

91

SC

AR

IDEA

: PA

ND

ALI

DA

EH

eter

ocar

pus d

orsa

lis B

ate,

188

831

3M

SH

eter

ocar

pus e

nsife

r A. M

ilne

Edw

ards

, 188

1U

SNM

233

349

31, 3

23

GH

eter

ocar

pus g

ibbo

sus B

ate,

188

831

3M

SH

eter

ocar

pus l

aevi

gatu

s Bat

e, 1

888

31, 3

23

GH

eter

ocar

pus l

epid

us d

e M

an, 1

917

313

MS

Het

eroc

arpu

s lon

giro

stri

s Mac

Gilc

hris

t, 19

0531

, 33

3M

SH

eter

ocar

pus s

ibog

ae d

e M

an, 1

917

313

MS

Het

eroc

arpu

s tri

cari

natu

s Alc

ock

& A

nder

son,

189

431

3M

SPl

esio

nika

edw

ards

ii (B

rand

t, 18

51)

313

MS

Ples

ioni

ka e

nsis

(A. M

ilne

Edw

ards

, 188

1)31

3M

SPl

esio

nika

mar

tia (A

. Miln

e Ed

war

ds, 1

883)

313

MS

Ples

ioni

ka o

cellu

s (B

ate,

188

8)31

3M

S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndC

AR

IDEA

: PR

OC

ESSI

DA

EN

ikoi

des s

ibog

ae d

e M

an, 1

918

5, 2

51

SN

ikoi

des s

tein

ii (E

dmon

dson

, 193

5)5,

10

301

SC

AR

IDEA

: RH

YN

CH

OC

INET

IDA

EC

inet

orhy

nchu

s con

colo

r (O

kuno

, 199

4)O

GP8

15-2

11

GC

inet

orhy

nchu

s haw

aiie

nsis

Oku

no &

Hoo

ver,

1998

OG

P648

-81

GC

inet

orhy

nchu

s hen

ders

oni (

Kem

p, 1

925)

?UF

824

OG

P648

-19

1G

Cin

etor

hync

hus h

iatti

(Hol

thui

s & H

ayas

hi, 1

967)

U

F 90

334

GP8

15-2

1G

Cin

etor

hync

hus r

etic

ulat

us O

kuno

, 199

7O

GP5

16-2

51

GC

inet

orhy

nchu

s str

iatu

s (N

omur

a &

Hay

ashi

, 199

2)O

GP8

16-1

51

GPA

LIN

UR

A: P

ALI

NU

RID

AE

Just

itia

long

iman

us (H

. Miln

e Ed

war

ds, 1

837)

UF

563

P2

GPa

libyt

hus m

agni

ficus

Dav

ie, 1

990

UF

1003

PG

P422

-26

2G

Palin

urel

lus w

iene

ckii

(de

Man

, 188

1)U

F 12

34P

GP8

26-1

01

GPa

nulir

us fe

mor

istr

iga

(Von

Mar

tens

, 187

2)U

F 89

4, U

GI 2

960

P1

GPa

nulir

us lo

ngip

es b

ispi

nosa

Bor

rada

ile, 1

899

UF

705

PG

P826

-81

GPa

nulir

us o

rnat

us (F

abric

ius,

1798

) P

IE1

GPa

nulir

us p

enic

illat

us (O

livie

r, 17

91)

UG

I 295

915

GP8

12-2

91

G, M

Panu

lirus

ver

sico

lor (

Latre

ille,

180

4)

UG

I 296

1P

1G

PALI

NU

RA

: SC

YLL

AR

IDA

EBi

arct

us v

itien

sis (

Dan

a, 1

852)

UF

2282

CG

P818

-91

GG

alea

rctu

s aur

ora

Hol

thui

s, 19

81U

F 22

81C

GP4

22-7

2G

Parr

ibac

us a

ntar

ctic

us (L

und,

179

3)U

F 89

5, B

PBM

PG

P816

-41

GA

STA

CID

EA: E

NO

PLO

MET

OPI

DA

EEn

oplo

met

opus

hol

thui

si G

ordo

n, 1

968

PG

P826

-51

GEn

oplo

met

opus

occ

iden

talis

(Ran

dall,

184

0)P

GP8

15-1

71

GTH

ALA

SSIN

IDEA

: AX

IID

AE

Alla

xius

cly

peat

us (d

e M

an, 1

887)

USN

M 2

4349

435

1G

Alla

xius

pic

teti

(Zeh

ntne

r, 18

94)

UF

468,

USN

M 2

4350

235

GP7

64-1

71

GAx

iops

is p

ica

Ken

sley

, 200

3U

SNM

296

400

(hol

otyp

e)35

GP5

74-2

51

GAx

iops

is sp

. 1 (a

ff. p

ica

Ken

sley

, 200

3)

UF

59

PG

P864

-13

311

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndAx

iops

is se

rrat

ifron

s (A

. Miln

e Ed

war

ds, 1

873)

USN

M 2

4336

235

GP2

49-3

81

GBo

uvie

raxi

us sp

ring

eri K

ensl

ey, 1

996

UF

58, U

SNM

296

398

35G

P527

-51

GM

anax

ius p

itatu

cens

is (d

e M

an, 1

925)

USN

M 2

4337

8 35

1G

Mar

iana

xius

kro

ppi K

ensl

ey, 1

996

USN

M 2

4353

0 (h

olot

ype)

35

, 36

1G

Para

scyt

olep

tus t

ride

ns (R

athb

un, 1

906)

USN

M 2

3935

235

1G

Para

xiop

sis b

isqu

amos

a (d

e M

an, 1

905)

5, 1

0, 3

51

SPa

raxi

opsi

s pau

layi

Ken

sley

, 200

3U

SNM

296

395

(hol

otyp

e)35

GP5

43:1

21

GPa

raxi

opsi

s plu

mos

iman

us K

ensl

ey, 2

003

USN

M 2

9639

4 (h

olot

ype)

351

GTH

ALA

SSIN

IDEA

: CA

LLIA

NA

SSID

AE

Cor

allic

hiru

s bay

eri K

ensl

ey, 2

001

USN

M 3

1067

7 (h

olot

ype)

, U

F 12

2737

1G

Para

glyp

turu

s cal

deru

s Tür

kay

& S

akai

, 199

538

1E

THA

LASS

INID

EA: C

ALL

IAN

IDEI

DA

EC

allia

nide

a ty

pa H

. Miln

e Ed

war

ds, 1

837

UF

470,

USN

M 2

4360

5, B

PBM

39, 4

0G

P275

-19

1G

, PTH

ALA

SSIN

IDEA

: STR

ATH

LAX

IID

AE

Nea

xius

aca

nthu

s (A

. Miln

e Ed

war

ds, 1

879)

UF

142,

USN

M 2

4360

45,

10,

35

GP5

75-4

321

G, S

AN

OM

UR

A: A

LBU

NEI

DA

EAl

bune

a sp

ecio

sa D

ana,

185

2U

F 31

17P

1-2

GA

NO

MU

RA

: CO

ENO

BIT

IDA

EBi

rgus

latr

o (L

inna

eus,

1767

)ZR

C5,

40,

41,

42, 9

3G

P575

-973

TG

, S, M

, As,

P, G

u, T

Coe

nobi

ta b

revi

man

us D

ana,

185

2U

F 28

3, U

F 58

6 (S

)40

, 42

GP8

11-8

TG

, S, A

n,G

uC

oeno

bita

cav

ipes

Stim

pson

, 185

8U

F 23

040

, 42

GP8

11-3

TG

, An,

Ag

Coe

nobi

ta p

erla

tus H

. Miln

e Ed

war

ds, 1

837

UF

583,

UF

600

(S)

40, 4

2G

P809

-18

TG

, S, A

n, S

,G

u, A

l, P,

Ag,

As,

MC

oeno

bita

pur

pure

us S

timps

on 1

858

401

MC

oeno

bita

rugo

sus H

. Miln

e Ed

war

ds, 1

837

UF

581,

UF

603

(S)

40, 4

2T

G, S

, An,

Gu,

Al,

P,A

g, M

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndA

NO

MU

RA

: DIO

GEN

IDA

EAn

icul

us a

nicu

lus (

Fabr

iciu

s, 17

87)

UF

40G

P584

-31

331

G, A

n,G

, Al,

Ag,

M, A

sAn

icul

us m

axim

us E

dmon

dson

, 195

2U

F42

GP4

69-1

51

GAn

icul

us re

tipes

Lew

inso

hn, 1

982

UF

430

42G

P453

-13

1G

Anic

ulus

urs

us (O

livie

r, 18

11)

USN

M 1

5247

5, U

F42

GP6

15-7

341

GAn

icul

us sp

. 1

UF

953

P1

GC

alci

nus a

reol

atus

Rah

ayu

& F

ores

t, 19

99U

F 44

9M

GP7

70-2

91

GC

alci

nus a

rgus

Woo

ster

, 198

4B

PBM

-S 1

0589

(hol

otyp

e)42

, 43

1G

Cal

cinu

s ele

gans

(H. M

ilne

Edw

ards

, 183

6)U

F 32

5 40

, 42,

43

GP8

55-1

1G

, An,

S,

Gu,

Al,

P,A

g, A

s, M

Cal

cinu

s gua

men

sis W

oost

er, 1

984

BPB

M-S

105

88 (h

olot

ype)

, UF

40, 4

2, 4

3G

P864

-23

1G

, Gu,

Ag,

MC

alci

nus h

aiga

e W

oost

er, 1

984

AH

F42

, 43

1G

Cal

cinu

s isa

bella

e Po

upin

, 199

7U

F 73

242

, 43

GP7

37-5

351

GC

alci

nus k

uroz

umii

Asa

kura

& T

achi

kaw

a, 2

000

CB

M-Z

C 4

057

(hol

otyp

e, P

)44

1P

Cal

cinu

s lae

vim

anus

(Ran

dall,

184

0)U

F 60

140

, 42,

43, 1

35G

P850

-20

1G

, S, A

n,A

l, G

u, P

,A

g, M

Cal

cinu

s lat

ens (

Ran

dall,

184

0)U

SNM

255

985,

UF

460

40, 4

2,43

, 45

GP7

70-3

61

G, A

n, A

l,G

u, M

Cal

cinu

s lin

eapr

opod

us M

orga

n &

For

est,

1991

UF

1322

42, 4

3G

P751

-24

371

GC

alci

nus m

inut

us B

uite

ndijk

, 193

7U

F 13

2142

, 43

GP5

90-3

31

GC

alci

nus m

orga

ni R

ahay

u &

For

est,

1999

UF

5, 1

0, 4

0,42

, 43

GP5

88-1

138

1G

, S, A

n,A

l, M

Cal

cinu

s pul

cher

For

est,

1958

UF

MG

P519

-26

1G

Cal

cinu

s rev

i Pou

pin

& M

cLau

ghlin

, 199

844

1M

, A,

Cal

cinu

s seu

rati

Fore

st, 1

951

UF

562

40G

P854

-28

1G

, An,

S,

Gu,

P, A

g,M

, U

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndC

iliop

agur

us m

ajor

For

est,

1995

462

GC

iliop

agur

us p

acifi

cus F

ores

t, 19

9546

2G

Cili

opag

urus

stri

gatu

s (H

erbs

t, 18

04)

AH

F 59

1-01

, UF

42, 4

6G

P751

-27

1G

Clib

anar

ius c

oral

linus

(H. M

ilne

Edw

ards

, 184

8)U

SNM

270

096,

UF

42G

P753

b-10

1G

Clib

anar

ius e

urys

tern

us H

ilgen

dorf

, 187

8U

F42

GP8

50-2

51

GC

liban

ariu

s hum

ilis (

Dan

a, 1

851)

UF

40, 4

2G

P850

-18

1G

, Gu,

Al

Clib

anar

ius s

trio

latu

s Dan

a, 1

852

UF

42G

P759

-11

1G

Clib

anar

ius v

ires

cens

(Kra

uss,

1843

)U

F42

GP8

50-9

1G

Dar

danu

s cra

ssim

anus

(H. M

ilne

Edw

ards

, 183

6)42

1G

Dar

danu

s def

orm

is (H

. Miln

e Ed

war

ds, 1

836)

UF

42G

P413

-28

1G

Dar

danu

s gem

mat

us (H

. Miln

e Ed

war

ds, 1

848)

UF

42G

P833

-13

1G

Dar

danu

s gut

tatu

s (O

liver

, 181

1)U

F42

GP5

17-2

61

GD

arda

nus l

agop

odes

(For

sskå

l, 17

75)

UF

326

42G

P764

-31

1G

Dar

danu

s meg

isto

s (H

erbs

t, 18

04)

UF

950

42G

P681

-21

1G

Dar

danu

s scu

tella

tus (

H. M

ilne

Edw

ards

, 184

8)U

F42

GP4

13-3

81

GD

ioge

nes p

alle

scen

s Whi

tele

gge,

189

7U

F 14

1 M

GP7

70-2

51

GPa

guri

stes

jalu

r Mor

gan,

199

2U

FM

GP5

44-3

01

GPa

guri

stes

mon

opor

us M

orga

n, 1

987

UF

MG

P848

-27

1G

Pseu

dopa

guri

stes

jane

tkae

McL

augh

lin, 2

002

47G

P754

-92

GPs

eudo

pagu

rist

es sp

. 2 (n

. sp.

) M

GP7

71-1

42

GA

NO

MU

RA

: GA

LATH

EID

AE

Allo

gala

thea

ele

gans

(Whi

te, 1

847)

UF

277

PG

P762

-27

1G

Gal

athe

a am

amie

nsis

Miy

ake

& B

aba,

196

640

1M

Mun

idop

sis m

aria

nica

Will

iam

s & B

aba,

198

92,

55

3M

TVA

NO

MU

RA

: HIP

PID

AE

Hip

pa p

acifi

ca D

ana,

185

2U

F 25

240

GP2

54-1

51

G, A

gA

NO

MU

RA

: LIT

HO

DID

AE

Lith

odes

sp. 1

(n. s

p.)

UF

2283

CG

P527

-27

3G

Para

lom

is h

aiga

e El

dred

ge, 1

976

BPB

M-S

852

4 (h

olot

ype)

483

GPa

ralo

mis

seag

rant

i Eld

redg

e, 1

976

BPB

M-S

852

6 (h

olot

ype)

483

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndA

NO

MU

RA

: PA

GU

RID

AE

Cat

apag

uroi

des f

ores

ti M

cLau

ghlin

, 200

2U

F 5

491

GPa

guri

tta k

ropp

i McL

augh

lin &

Lem

aitre

, 199

3U

SNM

259

396

(hol

otyp

e)50

GP4

69-2

539

1G

Pagu

rixu

s anc

eps (

Fore

st, 1

954)

401

Gu,

Ag,

M,

Al

Pagu

rixu

s bon

inen

sis (

Mel

in, 1

939)

UF

645

40G

P859

-33

1G

, PPa

guri

xus c

f. la

evim

anus

(Ortm

ann,

189

2)A

HF

5, 1

0G

P846

-18

361

GPa

guri

xus c

f. tw

eedi

ei (F

ores

t, 19

56)

401

MPa

guri

xus m

aoru

s (N

obili

, 190

6)U

FM

GP7

51-3

21

GPa

guri

xus s

p. 2

(n. s

p. fa

st p

urpl

e)

UF

432

MG

P813

-31

GPa

guri

xus s

p. 3

(sla

te)

UF

MG

P820

-10

1G

Pagu

rixu

s sp.

8 (p

ale)

U

F51

GP8

60-8

401

GPa

guri

xus s

p. 9

(n. s

p., p

urpl

e cl

aw)

UF

MG

P860

-21

1G

Pagu

rus h

irtim

anus

Mie

rs, 1

880

UF

307

MG

P846

-13

1G

Pagu

rus?

sp. 1

M

GP8

67-2

1G

Porc

ella

nopa

guru

s sp.

1 (n

. sp.

) Kro

pp &

Eld

redg

e, M

SR

KK

3G

Pylo

pagu

rops

is fi

mbr

iata

McL

augh

lin &

Hai

g, 1

989

USN

M 2

3141

3 (h

olot

ype)

, U

F 59

752

GP8

47-1

01

G

Pylo

pagu

rops

is k

eijii

McL

augh

lin &

Hai

g, 1

989

USN

M 2

3141

5 (p

arat

ype)

52G

P845

-25

1G

Tric

hopa

guru

s tri

chop

htha

lmus

(For

est,

1954

)U

F 64

4M

GP8

67-5

1G

AN

OM

UR

A: P

AR

APA

GU

RID

AE

Sym

pagu

rus d

ofle

ini (

Bal

ss, 1

912)

5341

3G

Tyla

spis

ano

mal

a H

ende

rson

, 188

554

3G

AN

OM

UR

A: P

OR

CEL

LAN

IDA

EN

eope

trol

isth

es m

acul

atus

(H. M

ilne

Edw

ards

, 183

7)U

SNM

255

986

KG

P776

-24

1G

Pach

yche

les g

arci

aens

is (W

ard,

194

2)U

SNM

255

976

40R

K G

UM

-21

8I1

G, M

Pach

yche

les p

isoi

des (

Hel

ler,

1865

)U

SNM

255

959

K1

GPa

chyc

hele

s spi

nipe

s (A

. Miln

e Ed

war

ds, 1

873)

USN

M 2

5595

5 K

1G

Petr

olis

thes

asi

atic

us (L

each

, 182

0)U

SNM

233

491

231

G, S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndPe

trol

isth

es b

ispi

nosu

s Bor

rada

ile, 1

900

USN

M 2

5720

7 56

RK

GU

M-

221C

1G

, S

Petr

olis

thes

bor

rada

ilei K

ropp

, 198

4Pa

raty

pe: U

SNM

210

633

23, 4

0R

K G

UM

-21

4A1

G, S

, An,

Gu,

P, M

Petr

olis

thes

car

inip

es (H

elle

r, 18

61)

USN

M 2

5723

8 K

1G

Petr

olis

thes

coc

cine

us (O

wen

, 183

9)U

F 47

7, U

SNM

257

230

40G

P750

-28

1G

, MPe

trol

isth

es d

ecac

anth

us O

rtman

n, 1

897

USN

M 2

1053

4 56

RK

GU

M-

208P

1G

, S

Petr

olis

thes

don

anen

sis O

saw

a, 1

997

KC

KR

K G

UM

-20

8M1

G

Petr

olis

thes

eld

redg

ei H

aig

& K

ropp

, 198

8Pa

raty

pe: U

SNM

222

328

561

GPe

trol

isth

es e

lega

ns H

aig,

198

1U

SNM

257

218

KR

K G

UM

-22

1G1

G

Petr

olis

thes

ext

rem

us K

ropp

& H

aig,

199

4 A

HF

571

GPe

trol

isth

es h

eter

ochr

ous K

ropp

, 198

6U

SNM

222

529

(hol

otyp

e)58

1G

, Gu,

SPe

trol

isth

es la

mar

ckii

(Lea

ch, 1

820)

USN

M 1

8936

1 23

, 40

GP7

51-5

1G

, S, A

nPe

trol

isth

es m

asak

ii M

iyak

e, 1

943

USN

M 2

1053

9 K

1G

Petr

olis

thes

mes

odac

tylo

n K

ropp

, 198

4U

SNM

210

624

(hol

otyp

e, P

) 23

, 40

1P,

Gu,

An,

MPe

trol

isth

es m

iyak

ei K

ropp

, 198

4U

SNM

210

628

(hol

otyp

e)23

, 40

1G

, MPe

trol

isth

es p

ubes

cens

Stim

pson

, 185

8U

SNM

257

240

40, 5

81

G, M

, An

Petr

olis

thes

tom

ento

sus (

Dan

a, 1

852)

UF

334,

USN

M 2

5760

6 58

GP7

51-6

1G

, P, S

BR

AC

HY

UR

A: A

ETH

RID

AE

Aeth

ra e

dent

ata

Edm

onds

on, 1

951

UF

481,

ZR

C

72G

P780

-10

1G

Aeth

ra sc

rupo

sa (L

inna

eus,

1764

) U

SNM

282

895

PG

P366

-71

GB

RA

CH

YU

RA

: BY

THO

GR

AEI

DA

EAu

stin

ogra

ea w

illia

msi

Hes

sler

& M

artin

, 198

92,

59

3M

TVB

RA

CH

YU

RA

: CA

LAPP

IDA

EC

alap

pa c

alap

pa (L

inna

eus,

1758

)U

F 75

, BPB

MN

GP8

56-7

1G

Cal

appa

gal

lus (

Her

bst,

1803

)U

F 82

, USN

M 2

7762

8,

UG

I 258

0G

GP8

14-2

61

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndC

alap

pa h

epat

ica

(Lin

naeu

s, 17

58)

UF

81, B

PBM

, UG

I 257

568

, 137

GP4

10-7

1G

, SC

alap

pa c

f. m

atsu

zaw

a G

alil,

199

7U

F/ZR

CN

GP5

79-3

5 1

GC

alap

pa sp

. 1 (a

ff. s

ebas

tiani

Gal

il, 1

997)

U

SNM

NR

K3

GC

alap

pa sp

. 2

UF

85

NG

P819

-33

1G

Cal

appa

sp. 3

(n. s

p.) N

g et

al.,

in p

rep.

UF

3094

, ZR

CN

GP7

39-2

71

GC

yclo

es g

ranu

losa

de

Haa

n, 1

837

USN

MK

1G

BR

AC

HY

UR

A: C

AR

PILI

IDA

EC

arpi

lius c

onve

xus (

Fors

skål

, 177

5)

UF

869,

USN

MK

GP8

31-3

61

GC

arpi

lius m

acul

atus

(Lin

naeu

s, 17

58)

UF

64, B

PBM

81, 1

37G

P820

-13

1G

BR

AC

HY

UR

A: C

RY

PTO

CH

IRID

AE

Cry

ptoc

hiru

s cor

allio

dyte

s Hel

ler,

1861

USN

M 2

5755

1 97

1G

Dac

ryom

aia

edm

onso

ni (F

ize

& S

erèn

e, 1

956)

USN

M

981

GD

acry

omai

a ja

poni

ca (T

aked

a &

Tam

ura,

198

1)K

CK

1G

Dac

ryom

aia

sp. 1

(n. s

p.) K

ropp

, MS

KC

K1

GD

acry

omai

a sp

. 2 (n

. sp.

) Kro

pp, M

SK

CK

1G

Fize

sere

neia

hei

mi (

Fize

& S

erèn

e, 1

956)

USN

M

981

GFi

zese

rene

ia la

tisel

la K

ropp

, 199

4K

C99

1G

Fize

sere

neia

stim

pson

i (Fi

ze &

Ser

ène,

195

6)U

SNM

257

250

K1

GFu

ngic

ola

utin

omii

(Fiz

e &

Ser

ène,

195

6)U

SNM

K

1G

Hap

aloc

arci

nus m

arsu

pial

is S

timps

on, 1

859

USN

M 1

0049

355,

98

GP6

44-2

31

G, S

Hir

oia

krem

pfi (

Fize

& S

erèn

e, 1

956)

USN

M 2

5724

5 K

1G

Lith

osca

ptus

hel

leri

(Fiz

e &

Ser

ène,

195

7)U

SNM

257

345

K1

GLi

thos

capt

us p

arad

oxus

A. M

ilne

Edw

ards

, 186

2U

SNM

257

368

971

GLi

thos

capt

us p

rion

otus

Kro

pp, 1

994

KC

991

GLi

thos

capt

us tr

i (Fi

ze &

Ser

ène,

195

6)U

SNM

257

336

K1

GLu

ciad

es a

gana

Kro

pp &

Man

ning

, 199

6U

SNM

268

890

(hol

otyp

e)10

01

GN

eotr

oglo

carc

inus

hon

gkon

gens

is (S

hen,

193

6)U

SNM

257

311

K1

GO

peca

rcin

us a

uran

tius K

ropp

, 198

9U

SNM

234

266

(hol

otyp

e)

101

1G

Ope

carc

inus

cre

scen

tus (

Edm

onds

on, 1

925)

USN

M 2

5747

8 10

11

GO

peca

rcin

us g

ranu

latu

s (Sh

en, 1

936)

USN

M 2

3426

4 10

11

GO

peca

rcin

us lo

bifr

ons K

ropp

, 198

9U

SNM

234

269

(hol

otyp

e)10

11

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndO

peca

rcin

us p

hole

ter K

ropp

, 198

9U

SNM

234

275

(hol

otyp

e)

101

1G

Ope

carc

inus

sier

ra K

ropp

, 198

9U

SNM

234

278

(hol

otyp

e)10

11

GPe

lyco

mai

a m

inut

a (E

dmon

dson

, 193

3)U

SNM

K

1G

Pseu

docr

ypto

chir

us v

irid

is H

iro, 1

938

USN

M 2

5731

4 10

21

GPs

eudo

hapa

loca

rcin

us ra

nson

i Fiz

e &

Ser

ène,

195

6U

SNM

257

285

981

GU

tinom

iella

dim

orph

a (H

ende

rson

, 190

6)U

SNM

257

512

981

GXy

nom

aia

shen

i (Fi

ze &

Ser

ène,

195

6)U

SNM

257

277

981

GB

RA

CH

YU

RA

: DA

IRID

AE

Dai

ra p

erla

ta (H

erbs

t, 17

90)

UF

1953

5, 1

0, 6

2G

P845

-41

G, S

, Gu,

P,

Al

BR

AC

HY

UR

A: D

RO

MII

DA

EC

rypt

odro

mia

falla

x (L

amar

ck, 1

818)

ZRC

200

0.07

5160

, 61,

62

421

G, M

Cry

ptod

rom

ia h

ilgen

dorf

i de

Man

, 188

8 U

SNM

128

571,

ZR

C 2

000.

2119

611

GC

rypt

odro

mia

pen

tago

nalis

Hilg

endo

rf, 1

879

ZRC

200

0.21

1561

1G

Cry

ptod

rom

ia p

itien

sis M

cLay

, 200

1 M

NH

N-B

2730

3 (h

olot

ype)

611

GC

rypt

odro

mia

tum

ida

Stim

pson

, 185

8 ZR

C 2

000.

2113

611

GC

rypt

odro

mio

psis

plu

mos

a (L

ewin

sohn

, 198

4)

ZRC

200

0.21

1261

1G

Cry

ptod

rom

iops

is u

nide

ntat

a (R

üppe

ll, 1

830)

U

GI 6

020

60, 6

1G

P380

-32

1G

Dro

mia

dor

mia

(Lin

naeu

s, 17

63)

601

SD

rom

idio

psis

leth

rinu

sae

(Tak

eda

& K

urat

a, 1

976)

ZR

C 2

000.

2121

611

GTa

kedr

omia

cri

stat

ipes

(Sak

ai, 1

969)

ZR

C 2

000.

0755

611

GB

RA

CH

YU

RA

: DY

NO

MEN

IDA

ED

ynom

ene

guam

ensi

s McL

ay, 2

001

MN

HN

-B26

474

(hol

otyp

e)61

RK

GU

M22

1F1

G

Dyn

omen

e hi

spid

a G

uérin

-Mén

evill

e, 1

832

621

MD

ynom

ene

krop

pi M

cLay

, 200

1M

NH

N-B

2647

6 (h

olot

ype)

, U

F 40

161

GP8

32-1

81

G

Dyn

omen

e pr

aeda

tor M

ilne

Edw

ards

, 187

9 ZR

C 2

000.

0743

61, 6

31

GH

irsu

tody

nom

ene

spin

osa

(Rat

hbun

, 191

1)ZR

C 2

000.

0745

, UF

313

61, 6

3G

P532

-31

1G

, As

Para

dyno

men

e tu

berc

ulat

a Sa

kai,

1963

ZR

C 2

000.

2122

, UF

2161

, 63

GP8

19-2

51

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndB

RA

CH

YU

RA

: ER

IPH

IID

AE

Dac

ryop

ilum

nus e

rem

ita N

obili

, 190

662

1M

Dac

ryop

ilum

nus r

athb

unae

Bal

ss, 1

932

USN

M5,

10

1G

, SEp

ixan

thoi

des a

nom

alus

Bal

ss, 1

935

UF

70, U

SNM

, ZR

CK

RK

-GU

M20

8G1

G

Epix

anth

us c

orro

sus A

. Miln

e Ed

war

ds, 1

873

621

MEr

iphi

a sc

abri

cula

Dan

a, 1

852

UF

113,

USN

M 1

5414

8, B

PBM

62G

P403

-21

G, A

lEr

iphi

a se

bana

(Sha

w &

Nod

der,

1803

)U

SNM

, ZR

CK

GP3

57-3

21

GG

lobo

pilu

mnu

s sp.

1

UF/

ZRC

NG

P829

-20

1G

Hyp

otha

lass

ia a

rmat

a (d

e H

aan,

183

5)U

SNM

, UG

I, U

F 28

4212

8R

K3

GLy

dia

annu

lipes

(H. M

ilne

Edw

ards

, 183

4)U

F 28

2, B

PBM

, ZR

C62

GP4

14-4

1G

, An,

Ag

Ozi

us ru

gulo

sus S

timps

on, 1

858

UF

293,

USN

M 1

5484

2 K

GP4

63-3

71

GO

zius

tube

rcul

osus

H. M

ilne

Edw

ards

, 183

4U

F 86

8, U

SNM

, UG

I62

GP7

86-1

71

G, A

n, G

u,A

g, A

sB

RA

CH

YU

RA

: GEC

AR

CIN

IDA

EC

ardi

som

a ca

rnife

x (H

erbs

t, 17

96)

UF

114

92G

P791

-29

TG

Dis

copl

ax h

irtip

es (D

ana,

185

1)U

F 14

5, B

PBM

, ZR

C92

GP3

31-3

2T

GD

isco

plax

long

ipes

A. M

ilne

Edw

ards

, 186

7U

F 14

7, U

SNM

181

977,

ZR

C92

, 133

GP8

09-1

TG

Dis

copl

ax ro

tund

a (Q

uoy

& G

aim

ard,

182

4)

UF

144,

USN

M 1

5504

9, Z

RC

62

, 92,

93

GP2

75-2

964

TG

, MEp

igra

psus

not

atus

(Hel

ler,

1865

)U

F 89

, ZR

C92

GP3

33-3

4T

GEp

igra

psus

sp. 1

(n. s

p.) N

g, 2

003

ZRC

94PK

LN1

GB

RA

CH

YU

RA

: GR

APS

IDA

EG

eogr

apsu

s cri

nipe

s (D

ana,

185

1)U

F 11

162

GP8

10-7

1G

, Gu,

MG

eogr

apsu

s gra

yi (H

. Miln

e Ed

war

ds, 1

853)

62

1A

n, G

u, U

Geo

grap

sus s

torm

i de

Man

, 189

5U

F 92

62G

P811

-20

1G

, MG

raps

us a

lbol

inea

tus L

amar

ck, 1

818

ZRC

L1

GG

raps

us lo

ngita

rsis

Dan

a, 1

851

UF

9362

GP3

38-2

51

G, A

n, A

gG

raps

us te

nuic

rust

atus

(Her

bst,

1783

)5,

62

GP8

09-3

01

G, S

, An,

Gu,

Sa,

P,

Ag,

M, A

lG

raps

us sp

. 1 (a

ff. l

ongi

tars

is D

ana,

185

1)

UF

134

PG

P414

-11

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndM

etop

ogra

psus

latif

rons

(Whi

te, 1

874)

UF

101

PG

P409

-34

1G

Met

opog

raps

us o

cean

icus

(Jac

quin

ot, 1

852)

UF

103

PG

P526

-28

1G

Met

opog

raps

us th

ukuh

ar (O

wen

, 183

9)U

F 95

, UF

591

(S)

PG

P409

-71

G, S

Pach

ygra

psus

min

utus

A. M

ilne

Edw

ards

, 187

3U

F 12

3, U

F 59

0 (S

)62

GP8

71-2

01

G, S

, M, A

lPa

chyg

raps

us p

lani

fron

s de

Man

, 188

8U

F 12

912

71

GPa

chyg

raps

us p

licat

us (H

. Miln

e Ed

war

ds, 1

837)

UF

130,

UF

584

(S),

BPB

M62

GP4

10-2

1G

, S, P

BR

AC

HY

UR

A: H

OM

OLI

DA

EH

omol

a di

ckin

soni

Eld

redg

e, 1

980

BPB

M-S

859

5 (h

olot

ype)

, ZR

C,

UG

I 676

764

, 65

3G

Hom

ola

orie

ntal

is H

ende

rson

, 188

8B

PBM

S10

479,

UG

I 676

464

3G

Lam

oha

pers

onat

a (G

uino

t & R

iche

r de

Forg

es, 1

981)

ZRC

663

GLa

moh

a w

illia

msi

(Tak

eda,

198

0)B

PBM

(par

atyp

e)14

271

3S

BR

AC

HY

UR

A: G

ON

EPLA

CID

AE

Con

leyu

s def

odio

Ng

& N

g, 2

003

UF

2098

(hol

otyp

e),

ZRC

200

2.17

82G

P694

-21

1G

Inte

sius

sp. 1

(n. s

p.) C

rosn

ier &

Ng,

200

3B

PBM

143

3G

Para

pilu

mnu

s ory

ctus

Ng,

200

3ZR

C 2

002.

174

(hol

otyp

e)83

GP4

69-3

31

GB

RA

CH

YU

RA

: HY

MEN

OSO

MA

TID

AE

Elam

ena

sp. 1

U

F 62

8T

GP3

91-9

1G

BR

AC

HY

UR

A: L

EUC

OSI

IDA

EAl

ox ru

gosu

m (S

timps

on, 1

858)

UF

729

P1

GD

rach

iella

ang

ulat

a (I

hle,

191

8)U

F 10

19B

GP4

08-2

41

GEb

alia

(?) s

p. 1

U

F 72

8N

GP5

37-1

1G

Ebal

iops

is e

rosa

(A. M

ilne

Edw

ards

, 187

4)

UF

621

BG

P408

-22

1G

Het

eron

ucia

ven

usta

Nob

ili, 1

906

UF

610

BG

P533

-12

1G

Leuc

osia

sp. 1

U

F 67

BG

P596

-23

1G

Myr

a fu

gax

(Fab

riciu

s, 17

98)

UF

2937

P1

GN

ew g

enus

? sp

. 1

UF

1021

B1

GN

ucia

spec

iosa

Dan

a, 1

852

UF

670

5, 1

0G

P597

-26

1G

, SN

ucia

? sp

. 1

UF

285

NG

P527

-21

1G

Nuc

ia?

sp. 2

U

F/ZR

C

NG

P818

-20

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndO

reot

los l

agar

odes

Tan

& N

g, 1

995

USN

M 2

6680

1 (p

arat

ype)

691

GO

reot

los p

ala

Tan

& N

g, 1

995

USN

M 2

5598

4 (h

olot

ype)

, U

F 53

069

1G

"Ran

dalli

a" se

rene

i Ric

her d

e Fo

rges

, 198

3U

F 85

6N

GP7

33-2

43

GB

RA

CH

YU

RA

: MA

JID

AE

Acha

eus l

acer

tosu

s? S

timps

on, 1

907

UF

629

NG

P823

-28

1G

Acha

eus l

aevi

ocul

is (d

e M

an, 1

905)

UF

690

NG

P583

-61

GBo

thro

mai

a gr

iffin

i Will

iam

s & M

offit

t, 19

91U

SNM

250

884

(hol

otyp

e, S

) 67

3S,

EC

ampo

scia

retu

sa L

atre

ille,

182

9U

SNM

282

898,

UF

2918

PG

P367

-13

1G

Cri

ocar

cinu

s sup

erci

liosu

s (Li

nnae

us, 1

767)

UF

855,

ZR

C, U

GI 2

945

NG

P692

-22

1G

Cyc

lax

subo

rbic

ular

is (S

timps

on, 1

858)

UF

15, U

GI

NG

P463

-11

G, A

sC

yrto

mai

a in

term

edia

Sak

ai, 1

938

UF

857

703

GD

umea

sp. 1

(aff

. tai

wan

icus

(Loh

& W

u, 1

998)

) ZR

CN

GP3

57-1

21

GH

ueni

a pr

oteu

s? d

e H

aan,

183

9U

F 39

4, U

GI 2

652

NG

P590

-81

GH

yast

enus

con

vexu

s Mie

rs, 1

884

UF

655

N1

GH

yast

enus

hilg

endo

rfi d

e M

an, 1

887

UF

850

N1

GH

yast

enus

pla

nasi

us (A

dam

s & W

hite

, 184

8)U

F 28

9, U

GI

NG

P576

-10

1G

Men

aeth

ius m

onoc

eros

(Lat

reill

e, 1

825)

UF

79, U

SNM

134

298,

U

GI 2

646

5, 1

0, 6

2G

P814

-13

1G

, S, G

u

Mic

ippa

par

ca A

lcoc

k, 1

895

5, 1

01

SM

icip

pa p

latip

es R

üppe

ll, 1

830

UF

666,

UG

I5,

10

GP4

14-2

31

G, S

Peri

nia

tum

ida

Dan

a, 1

852

UF

665

5, 1

0, 6

21

S, M

Pseu

dom

icip

pe e

ldre

dgei

Grif

fin &

Tra

nter

, 198

6A

HF

(hol

otyp

e), U

F 42

5, Z

RC

130

GP5

37-1

71

GRo

chin

ia d

ecip

iata

Will

iam

s & E

ldre

dge,

199

4U

SNM

251

434

(hol

otyp

e)71

3G

Schi

zoph

roid

a hi

lens

is (R

athb

un, 1

906)

UF

667,

ZR

C, U

GI

NG

P540

-61

GSc

hizo

phry

s asp

era

(H. M

ilne-

Edw

ards

, 183

4)U

F 28

7, U

GI

NG

P826

-31

1G

Sim

ocar

cinu

s obt

usir

ostr

is (M

iers

, 187

9)U

F 68

8, U

GI

5, 1

0G

P405

-343

1G

, STi

arin

ia c

orni

gera

(Lat

reill

e, 1

825)

UF

312,

UG

IN

GP8

37-2

21

GTi

arin

ia sp

inig

era

Stim

pson

, 190

762

1A

gTy

loca

rcin

us st

yx (H

erbs

t, 18

03)

5, 1

044

1S

Xeno

carc

inus

dep

ress

us M

iers

, 187

4U

F 65

9N

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndB

RA

CH

YU

RA

: OC

YPO

DID

AE

Mac

roph

thal

mus

con

vexu

s Stim

pson

, 185

8 U

F 11

8, B

PBM

, UG

I 673

7K

GP4

12-1

31

GM

acro

phth

alm

us la

trei

llei (

Des

mar

est,

1822

)U

F 28

46N

721

GM

acro

phth

alm

us v

erre

auxi

H. M

ilne

Edw

ards

, 184

8U

F 98

, BPB

M, Z

RC

KG

P412

-25

1G

Mac

roph

thal

mus

sp. 1

U

F 97

NG

P409

-17

1G

Ocy

pode

cer

atop

htha

lma

(Pal

las,

1772

) U

F 98

7, B

PBM

5, 6

2G

P336

-31

G, S

, Ag

Ocy

pode

cor

dim

ana

Des

mar

est,

1825

UF

115,

BPB

M,

USN

M 1

6846

2, Z

RC

62

GP3

36-1

71

G, P

Uca

cra

ssip

es (A

dam

s & W

hite

, 184

8)

UF

96T

GP4

08-3

11

GU

ca v

ocan

s (Li

nnae

us, 1

758)

UF

119

TG

P412

-34

1G

Ocy

podi

dae

sp. 2

U

F/ZR

CP

1G

BR

AC

HY

UR

A: P

ALI

CID

AE

Cro

ssot

onot

us sp

inip

es (d

e M

an, 1

888)

UF

517

TG

P540

-29

1G

Exop

alic

us m

acul

atus

(Edm

onds

on, 1

930)

UF

330

RG

P597

-22

1G

BR

AC

HY

UR

A: P

AR

THEN

OPI

DA

EAu

laco

lam

brus

hop

lono

tus (

Ada

ms &

Whi

te, 1

848)

UF

2099

, ZR

C 2

002.

201

73G

P414

-12

1G

Aula

cola

mbr

us h

ystr

icos

us T

an &

Ng,

200

3U

F 21

01 (h

olot

ype)

, ZR

C 2

002.

204

731

G

Cer

tola

mbr

us p

ugila

tor (

A. M

ilne

Edw

ards

, 187

3)U

F 9,

ZR

C 2

002.

206

731

GD

aldo

rfia

hor

rida

(Lin

naeu

s, 17

58)

UF

6, U

SNM

5, 1

0, 7

3,74

GP8

36-3

1G

, S

Dal

dorf

ia le

pros

a (N

obili

, 190

5)U

F/ZR

C73

, 74

1G

Furt

ipod

ia g

emm

a Ta

n &

Ng,

200

3ZR

C 2

002.

211

(hol

otyp

e),

UF

2103

731

G

Furt

ipod

ia p

etro

sa (K

lunz

inge

r, 19

06)

UF

8, Z

RC

200

2.21

373

GP5

40-8

1G

Lam

brac

haeu

s ram

ifer A

lcoc

k, 1

895

UF/

ZRC

144

1G

Nei

kola

mbr

us p

olem

iste

s Tan

& N

g, 2

003

ZRC

200

2.21

4 (h

olot

ype)

731

GPa

rthe

nopo

ides

car

iei B

ouvi

er, 1

914

UF

3773

, 74

RK

GU

M-

221D

1G

Rhin

olam

brus

rudi

s (R

athb

un, 1

916)

UF

2102

, ZR

C 2

002.

215

73G

P250

-35

1G

Thyr

olam

brus

effl

ores

cens

(Alc

ock,

189

5)U

F 41

9 73

, 74

GP8

36-8

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndB

RA

CH

YU

RA

: PIL

UM

NID

AE

Actu

mnu

s sp.

1 (n

. sp.

) Ng,

in p

rep.

UF

358

NG

P534

-21

GC

aeco

pilu

mnu

s hir

sutu

s (B

orra

daile

, 190

3)U

F/ZR

CP

GP5

80-3

1

GEc

hino

ecus

pen

tago

nus (

A. M

ilne

Edw

ards

, 187

9)U

SNM

154

036

N66

1G

Gla

brop

ilum

nus d

ispa

r (D

ana,

185

2)

UF

715

NG

P592

-21

1G

Gla

brop

ilum

nus l

aevi

man

us (D

ana,

185

2)U

F 71

6N

?GP5

32-1

91

GLo

phop

lax?

sp. 1

U

F 78

5N

GP8

58-1

51

GN

anop

ilum

nus c

oral

lioph

ilus (

Take

da &

Miy

ake,

196

9)

621

MPe

rman

otus

pur

pure

a (G

ordo

n, 1

934)

UF

482

75G

P783

-22

661

GPi

lum

nus c

urso

r A. M

ilne

Edw

ards

, 187

35,

10

1S

Pilu

mnu

s cf.

oahu

ensi

s Edm

onds

on, 1

931

UF

723

N1

GPi

lum

nus c

f. sc

abri

uscu

lus A

dam

s & W

hite

, 184

9U

F/ZR

CN

GP3

93-3

11

GPi

lum

nus l

ongi

corn

is H

ilgen

dorf

, 189

8U

F 72

4N

1G

Pilu

mnu

s ser

enei

Ng,

198

8U

F 78

7N

1G

Pilu

mnu

s tae

niol

a R

athb

un, 1

906

UF

725

NG

P466

-26

1G

Pilu

mnu

s ves

pert

ilio

(Fab

riciu

s, 17

98)

USN

M, U

F/ZR

CK

1G

Pilu

mnu

s sp.

1 (n

. sp.

) Ng,

in p

rep.

UF/

ZRC

NG

P466

-22

1G

Pseu

dolit

ochi

ra in

tege

r (M

iers

, 188

4)

5, 1

050

1S

Tana

oche

les s

teno

chilu

s Kro

pp, 1

984

USN

M 2

1063

6 (h

olot

ype)

89, 1

311

GTi

aram

edon

spin

osum

(Mie

rs, 1

879)

USN

M, U

F/ZR

CR

GP4

66-7

661

GVi

ader

iana

ara

nea

(Tes

ch, 1

918)

5, 1

045

1S

Viad

eria

na q

uadr

ispi

nosa

(Zeh

ntne

r, 18

94)

UF

80, U

GI

NG

P814

-17

1G

Viad

eria

na sp

. 4

UF

310

NG

P837

-29

1G

Viad

eria

na ty

pica

? W

ard,

194

2U

F/ZR

CN

GP5

28-2

1G

Viad

eria

na?

sp. 5

U

F/ZR

CN

GP8

27-2

01

GB

RA

CH

YU

RA

: PIN

NO

THER

IDA

EAp

hano

dact

ylus

sp. 1

(ter

ebel

lid c

omm

ensa

l) U

F/ZR

CN

GP7

86-1

31

GB

RA

CH

YU

RA

: PLA

GU

SIID

AE

Perc

non

abbr

evia

tum

(Dan

a, 1

851)

UF

106

PG

P808

-21

1G

Perc

non

affin

e (H

. Miln

e Ed

war

ds, 1

853)

UF

88P

GP5

12-2

41

GPe

rcno

n gu

inot

ae C

rosn

ier,

1965

U

F 10

4, Z

RC

PG

P512

-10

1G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndPe

rcno

n pl

anis

sim

um (H

erbs

t, 18

04)

BPB

M 1

671

62G

P424

-71

G, A

n, P

,A

g, M

Plag

usia

imm

acul

ata

Lam

arck

, 181

8ZR

C62

?, 1

29G

P512

-22

651

G, ?

Ag

Plag

usia

spec

iosa

Dan

a, 1

851

UF

87, B

PBM

125

1G

Plag

usia

squa

mos

a (H

erbs

t, 17

90)

ZRC

62?,

129

GP5

83-3

665

1G

, ?A

gB

RA

CH

YU

RA

: PO

RTU

NID

AE

Atop

ortu

nus g

usta

vi N

g &

Tak

eda,

200

3U

F 20

91 (h

olot

ype)

, ZR

C 2

002.

173

76G

P832

-12

1G

Cap

hyra

rotu

ndifr

ons (

A. M

ilne

Edw

ards

, 186

9)U

F 16

7, U

F 58

8 (S

)5,

10

GP7

90-2

31

G, S

Cap

hyra

sp. 1

(on

Aste

rosp

icul

aria

rand

alli)

U

F 25

88N

1G

Cap

hyra

uni

dent

ata?

Len

z, 1

910

UF

411

SG

P834

-34

1G

Car

upa

ohas

hii T

aked

a, 1

993

UF

152

PG

P818

-32

1G

Car

upa

sp. 2

(n. s

p.) N

g &

Tak

eda,

in p

rep.

UF/

ZRC

TG

P767

-41

GC

arup

a te

nuip

es D

ana,

185

1U

F 54

PG

P694

-24

1G

Cat

optr

us n

itidu

s A. M

ilne

Edw

ards

, 187

0U

F 18

2, U

SNM

P

GP5

71-4

1G

Cha

rybd

is e

ryth

roda

ctyl

a (L

amar

ck, 1

818)

UF

1915

621

G, A

gC

hary

bdis

haw

aien

sis E

dmon

dson

, 195

4 U

F 15

8N

GP8

28-2

41

GC

hary

bdis

hel

leri

i (A

. Miln

e Ed

war

ds, 1

867)

U

F 15

1N

GP7

49-1

21

GC

hary

bdis

obt

usifr

ons L

eene

, 193

6 U

F 19

2P

GP5

97-1

81

GC

hary

bdis

ori

enta

lis D

ana,

185

2U

F 17

9 N

GP5

13-1

91

GC

hary

bdis

pau

cide

ntat

a (A

. Miln

e Ed

war

ds, 1

861)

UF

3149

P1

GC

hary

bdis

sp. 1

(n. s

p. a

ff. o

man

ensi

s Lee

ne, 1

934)

U

F/ZR

CN

GP4

22-1

2G

Coe

loca

rcin

us fo

liatu

s Edm

onds

on, 1

930

UF

472

77G

P617

-15

1G

Lale

onec

tes n

ippo

nens

is (S

akai

, 193

8)

UF

2071

, UF

972

(As)

PG

P592

-12

1G

, As

Liby

stes

niti

dus?

A. M

ilne

Edw

ards

, 186

7 U

FP

GP4

11-2

31

GLi

byst

es v

illos

us R

athb

un, 1

924

UF

163

PG

P410

-14

1G

Liss

ocar

cinu

s lae

vis M

iers

, 188

6U

F 20

4P

GP5

33-7

1G

Liss

ocar

cinu

s orb

icul

aris

Dan

a, 1

852

UF

169,

BPB

M5,

10

GP6

93-1

01

G, S

Lupo

cycl

us q

uinq

uede

ntat

us R

athb

un, 1

906

UF/

ZRC

PG

P598

-16

1G

Podo

phth

alm

us v

igil

(Web

er, 1

795)

UG

I 672

9P

GP4

59-6

1G

Port

unus

alc

ocki

(Nob

ili, 1

905)

U

F 24

1N

GP3

97-2

51

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndPo

rtun

us c

onve

xus d

e H

aan,

183

3U

F 20

9P

GP8

10-2

81

GPo

rtun

us g

ranu

latu

s (H

. Miln

e Ed

war

ds, 1

834)

U

F 16

6, B

PBM

PG

P847

-19

1G

Port

unus

long

ispi

nosu

s Rat

hbun

, 190

6 U

F 17

1N

GP5

16-3

01

GPo

rtun

us m

acro

phth

alm

us R

athb

un, 1

906

UF

183

NG

P387

-19

1G

Port

unus

pet

reus

(Alc

ock,

189

9)U

F 18

8 N

GP5

80-1

61

GPo

rtun

us sa

ngui

nole

ntus

(Her

bst,

1783

)U

SNM

781

GPo

rtun

us tu

berc

ulos

us (A

. Miln

e Ed

war

ds, 1

861)

U

F 16

5N

GP3

91-1

71

GSc

ylla

serr

ata

(For

sskå

l, 17

75)

791

GTh

alam

ita a

dmet

e (H

erbs

t, 18

03)

UF

164,

USN

M 1

4961

6, B

PBM

5, 1

0, 6

2,78

, 141

GP4

14-3

61

G, S

, Gu,

M,

UTh

alam

ita a

uaue

nsis

Rat

hbun

, 190

6U

SNM

781

GTh

alam

ita c

f. ce

rasm

a W

ee &

Ng,

199

5U

F 21

8P

GP4

67-2

41

GTh

alam

ita c

oeru

leip

es Ja

cqui

not,

1846

UF

149,

USN

M78

GP8

50-3

01

GTh

alam

ita c

orru

gata

Ste

phen

son

& R

ees 1

961

UF

180

PG

P356

-25

1G

Thal

amita

cre

nata

H. M

ilne

Edw

ards

, 183

4 U

F 20

5, B

PBM

78

GP4

09-2

81

GTh

alam

ita d

akin

i Mon

tgom

ery,

193

1U

SNM

781

GTh

alam

ita e

xeta

stic

a A

lcoc

k, 1

899

UF

206

NG

P412

-30

1G

Thal

amita

gat

avak

ensi

s Nob

ili, 1

906

801

STh

alam

ita in

tegr

a D

ana,

185

2U

F 20

0, U

F 58

7 (S

), U

SNM

78G

P749

-24

1G

Thal

amita

med

ipac

ifica

Edm

onds

on, 1

954

UF

197

N1

GTh

alam

ita m

itsie

nsis

Cro

snie

r, 19

62U

F 19

0N

1G

Thal

amita

pic

ta S

timps

on, 1

958

UF

177,

UG

I5,

10,

62

GP5

40-1

21

G, S

, Al

Thal

amita

pilu

mno

ides

Bor

rada

ile, 1

903

5,10

1S

Thal

amita

pry

mna

(Her

bst,

1803

) U

F 22

1, B

PBM

78G

P576

-19

1G

Thal

amita

pse

udop

elsa

rti C

rosn

ier,

2002

UF

156,

UG

I, ZR

C12

6G

P382

-31

GTh

alam

ita q

uadr

iloba

ta M

iers

, 188

4 U

F 20

1N

GP5

44-1

31

GTh

alam

ita si

ma

H. M

ilne

Edw

ards

, 183

414

11

GTh

alam

ita sp

inife

ra B

orra

daile

, 190

2 U

F 20

8, U

SNM

255

982

NG

P813

-19

1G

Thal

amita

spin

imer

a St

ephe

nson

& R

ees,

1967

U

SNM

112

418

(hol

otyp

e),

UF

173

78G

P576

-36

1G

Thal

amita

woo

dmas

oni A

lcoc

k, 1

899

5, 1

01

S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndTh

alam

itoid

es q

uadr

iden

s A. M

ilne

Edw

ards

, 186

9U

F 17

0, U

SNM

255

987

5, 1

0G

P361

-14

1G

, SB

RA

CH

YU

RA

: PO

UPI

NII

DA

EPo

upin

ia h

irsu

ta G

uino

t, 19

91U

SNM

250

886

673

Ar

BR

AC

HY

UR

A: P

SEU

DO

ZIID

AE

Eury

oziu

s? sp

. 1

UF

302

NG

P850

-26

671

GPs

eudo

zius

cay

stru

s (A

dam

s & W

hite

, 184

8)

UF

294,

BPB

M, Z

RC

62G

P836

-23

671

G, S

a, G

u,M

, UB

RA

CH

YU

RA

: RA

NIN

IDA

ER

anin

idae

sp. 1

U

F 10

13

P1

PB

RA

CH

YU

RA

: SES

AR

MID

AE

Chi

rom

ante

s obt

usifr

ons (

Dan

a, 1

852)

UF

90, Z

RC

NG

P423

-81

GC

listo

coel

oma

sp. 1

U

F/ZR

CN

GP7

58-2

81

GLa

buan

ium

rotu

ndat

um (H

ess,

1865

)U

F 91

, ZR

CN

GP6

56-1

1T

GM

etas

esar

ma

rous

seau

xi H

. Miln

e Ed

war

ds, 1

853

UF

108,

ZR

C5,

10

GP7

38-9

TG

, SN

eosa

rmat

ium

four

man

oiri

Ser

ène,

197

3U

F 99

, ZR

CN

GP8

10-1

21

GN

eosa

rmat

ium

mei

nert

i (de

Man

, 188

7)U

F 10

7, Z

RC

N1

GN

eosa

rmat

ium

tris

pino

sum

Dav

ie, 1

994

ZRC

NG

P409

-22

1G

Para

sesa

rma

pict

um (D

e H

aan,

183

5)U

F 12

8N

GP7

58-1

71

GSe

sarm

a s.l

. sp.

2

UF/

ZRC

?P

1G

Sesa

rmoi

des g

uam

ensi

s Ng,

200

2eU

F, Z

RC

124

PKLN

1G

Stel

gist

ra st

orm

i de

Man

, 189

5U

F 12

5 N

HC

L1

GB

RA

CH

YU

RA

: TR

APE

ZIID

AE

Qua

drel

la sp

. 85

1G

Tetr

alia

cin

ctip

es P

auls

on, 1

875

SMF

2692

785

1G

Tetr

alia

fulv

a Se

rène

, 198

4U

SNM

(S)

851

STe

tral

ia ru

brid

acty

la G

arth

, 197

1U

SNM

, SM

F 26

282

851

G, S

Tetr

alia

van

nini

i Gal

il &

Cla

rk, 1

988

UF

911,

SM

F 26

929

85G

P856

-27

1G

Tetr

aloi

des h

eter

odac

tyla

(Hel

ler,

1861

)U

SNM

(S)

851

STe

tral

oide

s nig

rifr

ons (

Dan

a, 1

852)

USN

M 2

8598

0, S

MF

2628

4 85

1G

, STr

apez

ia c

ymod

oce

(Her

bst,

1799

)B

PBM

162

9, S

MF

2517

185

, 90

1G

Trap

ezia

dig

italis

Lat

reill

e, 1

828

SMF

2628

585

1G

, M

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndTr

apez

ia fe

rrug

inea

Lat

reill

e, 1

828

BPB

M-S

118

80, S

MF

2628

985

GP6

56-2

0 1

G, S

, As,

MTr

apez

ia fl

avop

unct

ata

Eydo

ux &

Sou

leye

t, 18

42ZR

C85

GP6

56-2

1

G, M

Trap

ezia

form

osa

Smith

, 186

9SM

F 26

286

851

G, M

Trap

ezia

gar

thi G

alil,

198

4M

NH

N-B

261

3285

1G

, STr

apez

ia g

utta

ta R

üppe

ll, 1

830

USN

M, S

MF

2520

1, U

GI 2

607

85, 9

11

G, S

Trap

ezia

lute

a C

astro

, 199

7B

PBM

S16

29, U

SNM

85G

P290

-15

1G

, STr

apez

ia n

egle

cta

Cas

tro, 2

003

SMF

2629

0 (h

olot

ype)

851

GTr

apez

ia p

lana

War

d, 1

941

USN

M 2

6083

5 (S

)85

1S,

Ag

Trap

ezia

rufo

punc

tata

(Her

bst,

1799

)B

PBM

-S 1

1879

, USN

M85

GP6

44-3

11

G, S

Trap

ezia

sept

ata

Dan

a, 1

852

USN

M 2

8648

5, M

NH

N-B

971

485

, 91

1G

Trap

ezia

sere

nei O

dine

tz, 1

984

MN

HN

B-9

681

(hol

otyp

e),

USN

M 2

8593

3 85

, 91

GP6

56-2

6 1

G

Trap

ezia

spec

iosa

Dan

a, 1

852

USN

M

851

G, S

BR

AC

HY

UR

A: V

AR

UN

IDA

EC

yclo

grap

sus i

nteg

er H

. Miln

e Ed

war

ds, 1

837

621

MC

yclo

grap

sus s

p. 1

U

F 13

3P

GP7

38-2

1G

Cyc

logr

apsu

s sp.

2

UF/

ZRC

PG

P738

-30

1G

Hel

ice

subq

uadr

ata

(Dan

a, 1

851)

UF/

ZRC

N1

GO

rcov

ita m

ollit

ia N

g, G

uino

t, Ili

ffe,

199

6 U

F 11

2, Z

RC

95G

P733

-22

FWG

Ptyc

hogn

athu

s sp.

1

UG

I 262

1P

FWG

Ptyc

hogn

athu

s sp.

2 (c

f. ba

rbat

us (A

. Miln

e Ed

war

ds, 1

873)

)U

F 12

6N

GP7

38-1

81

GSc

utum

ara

sp. 1

U

F/ZR

CN

PKLN

1G

Varu

na li

ttera

ta (F

abric

ius,

1798

)U

GI 2

626

PFW

GXe

nogr

apsu

s nov

aein

sula

ris T

aked

a &

Kur

ata,

197

7 SM

F 22

945

961

EB

RA

CH

YU

RA

: XA

NTH

IDA

EAc

taea

pol

yaca

ntha

(Hel

ler,

1861

)U

F 38

65,

10

GP8

58-2

31

G, S

Acta

ea sp

inos

issi

ma

Bor

rada

ile, 1

902

UF

758

N1

GAc

taeo

des h

irsu

tissi

mus

(Rüp

pell,

183

0)U

SNM

154

036,

UF

2991

5, 1

046

1G

, SAc

taeo

des t

omen

tosu

s (H

. Miln

e Ed

war

ds, 1

834)

UF

585,

BPB

M, U

GI 2

593

KG

P782

-12

1G

Ater

gatis

dila

tatu

s de

Haa

n, 1

835

5, 1

047

1S

Ater

gatis

flor

idus

(Lin

naeu

s, 17

67)

UF

1977

, BPB

M 1

616

5, 1

0, 1

37G

P366

-27

1G

, S

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndAt

erga

tis g

ranu

latu

s de

Man

, 188

9U

F 87

4P

GP5

98-3

11

GAt

erga

tis la

tissi

mus

(H. M

ilne

Edw

ards

, 183

4)U

F 70

6P

GP3

67-1

81

GAt

erga

tis n

itidu

s A. M

ilne

Edw

ards

, 186

5U

F 71

4N

481

GAt

erga

tis su

bden

tatu

s de

Haa

n, 1

835

UF

272

PG

P598

-61

GAt

erga

tops

is g

ranu

latu

s A. M

ilne

Edw

ards

, 186

1U

F 31

47P

1G

Ater

gato

psis

luca

si M

ontro

uzie

r, 18

65

UF

875,

UG

I 675

3N

1G

Ater

gato

psis

sign

atus

(Ada

ms &

Whi

te, 1

848)

UF

880

PG

P819

-16

1G

Bana

reia

cf.

arm

ata

A. M

ilne

Edw

ards

, 186

9 U

F 79

7N

1G

Chl

orod

iella

bar

bata

(Bor

rada

ile, 1

900)

5, 1

01

SC

hlor

odie

lla c

ythe

rea

(Dan

a, 1

852)

U

F 39

15,

10,

62

GP8

59-2

11

G, S

, P, M

Chl

orod

iella

laev

issi

ma

(Dan

a, 1

852)

5,

10

1S

Cym

o an

dreo

ssyi

(Aud

ouin

, 182

6)U

F 78

85,

10

1G

, SC

ymo

quad

rilo

batu

s Mie

rs, 1

884

UF

863

PG

P656

-61

GD

omec

ia g

labr

a A

lcoc

k, 1

899

UF

711

5, 1

0, 6

21

G, S

, MD

omec

ia h

ispi

da E

ydou

x &

Sou

leye

t, 18

4262

1A

sEt

isus

mol

okai

ensi

s (R

athb

un, 1

906)

5, 1

01

SEt

isus

ana

glyp

tus H

. Miln

e Ed

war

ds, 1

834

UF

76, U

SNM

NG

P827

-25

1G

Etis

us b

ifron

talis

(Edm

onds

on, 1

935)

UF

804

N1

GEt

isus

dem

ani O

dhne

r, 19

25U

F 34

1, U

GI

NG

P808

-91

GEt

isus

den

tatu

s (H

erbs

t, 17

85)

UF

740,

USN

M 1

5415

3, B

PBM

PR

FM-

145N

1G

Etis

us e

lect

ra (H

erbs

t, 18

01)

USN

M 1

5415

4 5,

10

1G

, SEt

isus

fron

talis

(Dan

a, 1

852)

UF

757

N1

GEt

isus

laev

iman

us R

anda

ll, 1

840

UF

589

(S),

UG

IP

1G

Etis

us sp

lend

idus

Rat

hbun

, 190

6U

F 87

1 P

GP5

97-3

31

GEt

isus

util

is Ja

cqui

not,

1854

UF

3560

, USN

MK

1G

Euxa

nthu

s exs

culp

tus (

Her

bst,

1790

)U

F 86

2, B

PBM

KG

P319

-32

1G

Fore

stia

dep

ress

a (W

hite

, 184

7)62

1P,

MG

ailla

rdie

llus o

rien

talis

(Odh

ner,

1925

)U

F 86

1N

GP4

14-1

71

GG

ailla

rdie

llus s

uper

cilia

ris (

Odh

ner,

1925

)5,

10

491

SH

ypoc

olpu

s abb

otti

(Rat

hbun

, 189

4)U

F 72

6 P

GP8

28-1

81

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndJo

nesi

us tr

iung

uicu

latu

s (B

orra

daile

, 190

2)U

SNM

84

, 85

1G

Juxt

axan

thia

s liv

idus

(Lam

arck

, 181

8)U

SNM

K1

GJu

xtax

anth

ias t

etra

odon

(Hel

ler,

1865

)86

1G

Kra

ussi

a ru

gulo

sa (K

raus

s, 18

43)

UF

886

5, 1

0, 6

2G

P814

-28

1G

, S, A

lLa

chno

podu

s bid

enta

tus (

A. M

ilne

Edw

ards

, 186

7)U

SNM

134

551

K1

GLa

chno

podu

s rod

gers

i Stim

pson

, 185

862

1M

Lach

nopo

dus s

ubac

utus

(Stim

pson

, 185

8)U

F 86

0 N

GP7

51-2

11

GLe

ptod

ius e

xara

tus (

H. M

ilne

Edw

ards

, 183

4)U

SNM

, UG

IK

1G

Lept

odiu

s nud

ipes

(Dan

a, 1

852)

UF

423,

USN

MK

1G

Lept

odiu

s san

guin

eus (

H. M

ilne

Edw

ards

, 183

4)U

F 29

2, U

SNM

134

558,

UG

I62

GP8

36-3

11

G, A

an, G

u,P,

Ag,

M,

Al

Lioc

arpi

lode

s arm

iger

pac

ificu

s Bal

ss, 1

938

USN

M 1

5471

6 5,

10

511

G, S

Lioc

arpi

lode

s har

msi

Bal

ss, 1

934

6252

1G

u, M

Lioc

arpi

lode

s int

eger

rim

us (D

ana,

185

2)U

SNM

154

732

5, 1

01

G, S

Liom

era

bella

(Dan

a, 1

852)

USN

M 1

5413

9 5,

10

551

G, S

Liom

era

cinc

timan

a (W

hite

, 184

7)U

F 56

8, U

SNM

KG

P832

-24

1G

Liom

era

mon

ticul

osa

(A. M

ilne

Edw

ards

, 187

3)5,

10

531

SLi

omer

a ru

gata

(H. M

ilne

Edw

ards

, 183

4)62

RK

-GU

M-

208F

1G

, P, M

Liom

era

sem

igra

nosa

de

Man

, 188

85,

10

541

SLi

omer

a st

imps

oni (

A. M

ilne

Edw

ards

, 186

5)U

F 71

8 N

GP4

56-2

61

GLi

omer

a st

riol

ata

(Odh

ner,

1925

)U

SNM

155

973

K1

GLi

omer

a tr

istis

(Dan

a, 1

852)

UF

777,

USN

M 9

3940

5, 1

0G

P357

-356

1G

, SLi

omer

a? sp

. 1

UF/

ZRC

NG

P865

-51

GLi

omer

a sp

. 2 (n

. sp.

? af

f. tr

istis

(Dan

a, 1

852)

) U

F 77

9N

GP6

15-9

1G

Loph

ozoz

ymus

ber

tonc

inia

e G

uino

t & R

iche

r de

Forg

es, 1

981

UF

896

NG

P422

-29

2G

Loph

ozoz

ymus

edw

ards

i (O

dhne

r, 19

25)

UF

805

NG

P519

-18

1G

Loph

ozoz

ymus

inci

sus (

H. M

ilne

Edw

ards

, 183

4)U

F 80

3, U

SNM

286

496

NG

P544

-10

1G

Loph

ozoz

ymus

pic

tor (

Fabr

iciu

s, 17

98)

BPB

M S

1615

K1

GLo

phoz

ozym

us p

ulch

ellu

s A. M

ilne

Edw

ards

, 186

75,

10,

62

1S,

Ag

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndLy

bia

tess

elat

a (L

atre

ille,

181

2)U

F 77

0, B

PBM

5, 1

0, 6

2G

P538

-24

1G

, S, M

Lybi

a sp

. 1

UF

383

PG

P858

-51

GM

acro

med

aeus

cra

ssim

anus

(A. M

ilne

Edw

ards

, 186

7)U

F/ZR

CN

GP4

67-1

81

GM

acro

med

aeus

nud

ipes

(A. M

ilne

Edw

ards

, 186

7)U

F 39

862

GP8

51-1

01

G, M

Med

aeus

? or

natu

s Dan

a, 1

852

5, 1

01

SN

eolio

mer

a ri

chte

rsoi

des?

Sak

ai, 1

969

UF

495

PG

P402

-32

1G

Neo

liom

era

sp. 1

(pin

k-re

d)

UF

29

KG

P829

-28

1G

Neo

liom

era

sp. 2

(ora

nge)

U

F 20

10K

GP8

29-1

81

GN

eolio

mer

a sp

. 3 (f

lesh

) U

F 42

0N

GP8

29-3

41

GN

eolio

mer

a st

riat

a B

uite

ndijk

, 194

1U

F 50

0, U

SNM

, ZR

CK

1G

New

gen

us (a

ff. P

aram

edae

us) s

p. 1

(n. s

p.) N

g, in

pre

p.U

F 42

NG

P832

-22

1G

Ole

noth

us u

ogi N

g, 2

003

UF

2096

(hol

otyp

e),

ZRC

200

2.17

187

GP5

76-3

01

G

Pala

pedi

a sp

. 1

UF

396

PG

P851

-16

1G

Pala

pedi

a sp

. 2

UF

461

PG

P786

-23

1G

Para

ctae

a ru

fopu

ncta

ta c

ompl

ex (H

. Miln

e Ed

war

ds, 1

834)

UF

3N

GP8

47-1

31

GPa

ram

edae

us n

oele

nsis

(War

d, 1

934)

UF

756

NG

P457

-51

GPa

ram

edae

us si

mpl

ex (A

. Miln

e Ed

war

ds, 1

873)

UF

380

NG

P836

-15

1G

Para

xant

hias

not

atus

(Dan

a, 1

852)

UF

416,

USN

M 1

5484

6 5,

10,

62

GP7

51-1

61

G, S

, MPa

raxa

nthi

as p

arvu

s (B

orra

daile

, 190

0)U

SNM

5, 1

057

1G

, SPh

ymod

ius n

itidu

s (D

ana,

185

2)5,

10

1S

Phym

odiu

s ung

ulat

us (H

. Miln

e Ed

war

ds, 1

834)

UF

881,

USN

M 1

5495

4 5,

10

1G

, SPi

lodi

us a

reol

atus

(H. M

ilne

Edw

ards

, 183

4)U

F 36

, USN

MK

GP2

73-2

21

GPi

lodi

us g

ranu

latu

s Stim

pson

, 185

9 U

F 71

2N

GP4

63-1

81

GPi

lodi

us m

elan

ospi

nis (

Rat

hbun

, 191

1)U

F 76

4 N

1G

Pilo

dius

pau

mot

ensi

s Rat

hbun

, 190

7U

SNM

134

593

62, 8

81

G, M

Pilo

dius

pilu

mno

ides

(Whi

te, 1

848)

U

F/ZR

C5,

10,

88

581

G, S

Pilo

dius

pub

esce

ns D

ana,

185

2 5,

10,

88

591

SPi

lodi

us p

ugil

Dan

a, 1

852

5, 1

0, 8

860

1S

Pilo

dius

scab

ricu

lus D

ana,

185

2 88

1S

Pilo

dius

sere

nei M

iyak

e &

Tak

eda,

196

8U

F 78

0N

611

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndPi

lodi

us sp

. 1U

F/ZR

CN

GP7

66-1

11

GPi

lodi

us sp

inip

es H

elle

r, 18

61U

F 76

7, U

SNM

KG

P534

-16

1G

Plat

ypod

ia a

nagl

ypta

(Hel

ler,

1861

)U

F 77

4N

?GP7

91-1

11

GPl

atyp

odia

sem

igra

nosa

(Hel

ler,

1861

)U

F 77

65,

10

1G

, SPo

lyde

ctus

cup

ulife

r (La

treill

e, 1

812)

UF

783

P1

GPs

aum

is c

avip

es (D

ana,

185

2)U

F 76

55,

10

?GP8

59-2

462

1G

, SPs

eudo

liom

era

gran

osim

ana

(A. M

ilne

Edw

ards

, 186

5)62

1M

Pseu

dolio

mer

a he

lleri

(A. M

ilne

Edw

ards

, 186

5)62

1M

, UPs

eudo

liom

era

lata

(Bor

rada

ile, 1

902)

5, 1

063

1S

Pseu

dolio

mer

a re

mot

a (R

athb

un, 1

907)

UF

864

62G

P533

-81

G, M

Pseu

dolio

mer

a ru

ppel

lioid

es (O

dhne

r, 19

25)

UF

775

NG

P821

-21

GPs

eudo

liom

era

spec

iosa

(Dan

a, 1

852)

621

M, U

Twee

diei

a la

ysan

i (R

athb

un, 1

906)

UF

763

NG

P466

-21

1G

Xant

hias

aff

. che

rbon

nier

i Gui

not,

1964

UF/

ZRC

N1

GXa

nthi

as c

anal

icul

atus

Rat

hbun

, 190

6U

F 72

1N

1G

Xant

hias

gilb

erte

nsis

(Bal

ss, 1

938)

621

MXa

nthi

as la

mar

cki (

H. M

ilne

Edw

ards

, 183

4)U

F 71

75,

10

GP4

63-2

31

G, S

Xant

hias

latif

rons

(de

Man

, 188

8)U

F 79

2N

GP4

56-1

91

GXa

nthi

as li

vidu

s (La

mar

ck, 1

818)

621

MXa

nthi

as p

unct

atus

(H. M

ilne

Edw

ards

, 183

4)U

F 86

5, U

SNM

5,

10

GP4

01-4

1G

, SZo

sim

us a

eneu

s (Li

nnae

us, 1

758)

USN

M, U

GI 2

608

KG

P596

-24

1G

ASC

OTH

OR

AC

IDA

: SY

NA

GO

GID

AE

Sess

ilogo

ga e

long

ata

Gry

gier

, 199

0U

SNM

228

258

(hol

otyp

e)10

31

GC

IRR

IPED

IA: A

RC

HA

EOB

ALA

NID

AE

Con

opea

folli

culu

s Hiro

, 193

710

6, 1

101?

SC

IRR

IPED

IA: B

ALA

NID

AE

Bala

nus a

mph

itrite

Dar

win

, 185

4 U

F 33

610

61

GBa

lanu

s ebu

rneu

s Gou

ld, 1

841

UF

223

106

GP8

42-1

31

GC

IRR

IPED

IA: C

HTH

AM

ALI

DA

EC

htha

mal

us m

alay

ensi

s Pils

bry,

191

6U

SNM

, SIO

105,

106

1G

Cht

ham

alus

pro

teus

Dan

do &

Sou

thw

ard,

198

0U

F 26

4, S

IO10

6, 1

071

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndEu

raph

ia h

embe

li C

onra

d, 1

837

USN

M

106,

107

,10

81

G

Pseu

doeu

raph

ia m

ontg

omer

yi (F

oste

r, 19

90)

USN

M 2

3782

0 (h

olot

ype)

105,

106

1G

Nes

ocht

ham

alus

inte

rtex

tus (

Dar

win

, 185

4)

UF

222

107,

108

1G

CIR

RIP

EDIA

: CR

YPT

OPH

IALI

DA

EC

rypt

ophi

alus

cor

onop

horu

s Sm

yth,

198

6U

SNM

222

986

(hol

otyp

e)10

6, 1

12,

113

1G

Cry

ptop

hial

us c

f. zu

lloi T

omlin

son,

197

310

6, 1

13,

114

1G

CIR

RIP

EDIA

: LIT

HO

GLY

PTID

AE

Lith

ogly

ptes

miti

s Tom

linso

n, 1

969

106,

113

,11

41

G

CIR

RIP

EDIA

: NEO

VER

RU

CID

AE

Neo

verr

uca

brac

hyle

pado

form

is N

ewm

an, 1

989

USN

M2,

104

3M

TVC

IRR

IPED

IA: P

AC

HY

LASM

ATI

DA

ETe

trap

achy

lasm

a or

natu

m Jo

nes,

2000

YPM

930

3 (h

olot

ype,

S),

SIO

, U

F 48

010

6, 1

09G

P768

-21

G, S

CIR

RIP

EDIA

: PY

RG

OM

ATI

DA

EC

ante

llius

eus

pinu

losu

m (B

roch

, 193

1)10

61

GC

ante

llius

pal

lidus

(Bro

ch, 1

931)

106

1S

Can

telli

us se

ptim

us (H

iro, 1

938)

10

61

GN

eotr

evat

hana

elo

ngat

a (H

iro, 1

931)

106

1G

Savi

gniu

m c

rena

tum

(Sow

erby

, 182

3)10

61

GTr

evat

hana

den

tata

(Dar

win

, 185

4)10

6, 1

11G

P263

-21

GTr

evat

hana

ori

enta

le (R

en, 1

986)

10

6G

P529

-20

1G

Trev

atha

na p

aula

yi A

sam

i & Y

amag

uchi

, 200

1N

SMT-

Cr1

3681

(hol

otyp

e)10

6, 1

11G

P263

-61

GW

anel

la m

illep

orae

(Dar

win

, 185

4)10

6G

P263

-91

GC

IRR

IPED

IA: S

CA

LPEL

LID

AE

Lith

otry

a ni

coba

rica

Rei

nhar

dt, 1

850

SIO

106

1G

CIR

RIP

EDIA

: TET

RA

CLI

TID

AE

Tetr

aclit

ella

div

isa

(Nils

son-

Can

tell,

192

1)U

SNM

10

5, 1

061

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndTe

sser

opor

a pa

cific

a Pi

lsbr

y, 1

928

USN

M10

5, 1

06G

P420

-21

1G

Tess

erop

ora

sp. 1

10

61

GC

IRR

IPED

IA: V

ERR

UC

IDA

EVe

rruc

a co

okei

Pils

bry,

192

8U

SNM

10

5, 1

061

GPY

CN

OG

ON

IDA

: AM

MO

THEI

DA

EAc

helia

ass

imili

s (H

asw

ell,

1884

)U

SNM

11

51

GAm

mot

hella

sp. i

ndet

. U

SNM

11

51

GAm

mot

hella

tipp

ula

Chi

ld, 1

983

USN

M 1

9537

4 (h

olot

ype)

11

5, 1

16,

117

1G

Anop

loda

ctyl

us c

ham

orru

s Chi

ld, 1

983

USN

M 1

9537

6 (h

olot

ype)

115,

116

,11

71

G

Anop

loda

ctyl

us g

land

ulife

r Sto

ck, 1

954

USN

M

115

1G

Anop

loda

ctyl

us lo

ngic

eps S

tock

, 195

1U

SNM

11

51

GAn

oplo

dact

ylus

pyc

noso

ma

(Hel

fer,

1938

)U

SNM

11

5, 1

171

GAn

oplo

dact

ylus

tenu

icor

pus C

hild

, 199

1U

SNM

11

51

GEu

rycy

de se

rtul

a C

hild

, 199

1U

SNM

234

586

(hol

otyp

e)11

51

GSe

rico

sura

coc

hlei

fove

a C

hild

, 198

9U

SNM

234

505

(hol

otyp

e)

2, 1

183

MTV

Tany

styl

um re

hder

i Chi

ld, 1

970

USN

M

115,

117

1G

PYC

NO

GO

NID

A: C

ALL

IPA

LLEN

IDA

EC

allip

alle

ne n

ovae

zeal

andi

ae (T

hom

son,

188

4)U

SNM

11

51

GSe

guap

alle

ne m

icro

nesi

ca C

hild

, 198

3U

SNM

11

51

GPY

CN

OG

ON

IDA

: EN

DEI

DID

AE

Ende

is m

ollis

(Car

pent

er, 1

904)

USN

M

115

1G

PYC

NO

GO

NID

A: N

YM

PHO

NID

AE

Nym

phon

dia

bolis

Chi

ld, 1

988

USN

M11

51

GFR

ESH

WA

TER

TA

XA

CO

PEPO

DA

: PH

YLL

OG

NA

THO

POD

IDA

EPh

yllo

gant

hopu

s vig

uier

i (M

aupa

s, 18

92)

120

FWG

, Ag

CO

PEPO

DA

: CY

CLO

PID

AE

Bryo

cycl

ops a

nnin

ae (M

enze

l, 19

26)

139

FWG

Mes

ocyc

lops

leuc

kart

i (C

laus

, 185

7)13

9FW

G

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndC

LAD

OC

ERA

: SID

IDA

ED

iaph

anos

oma

sars

i Ric

hard

, 189

4 s.

l. U

SNM

141

128

138

FWG

Lato

nops

is a

ustr

alis

Sar

s, 18

88U

SNM

141

128

119,

138

FWG

CLA

DO

CER

A: M

AC

RO

THR

ICID

AE

Mac

roth

rix

latic

orni

s (Ju

rine,

182

0)11

9FW

GC

LAD

OC

ERA

: MO

INID

AE

Moi

na a

ustr

alie

nsis

Sar

s, 18

9611

9FW

GM

oina

mac

roco

pa m

acro

copa

(Stra

us, 1

820)

119

FWG

Moi

na w

eism

anni

Ishi

kaw

a, 1

896

119

FWG

TER

RES

TRIA

L TA

XA

AM

PHIP

OD

A: T

ALI

TRID

AE

Flor

esor

ches

tia a

nom

ala

(Che

vreu

x, 1

901)

121

TA

g, A

lFl

ores

orch

estia

sp. 1

12

1T

SaIS

OPO

DA

: AR

MA

DIL

LID

AE

Cub

aris

sp. 1

12

2T

SH

ybod

illo

sp. 1

12

3T

P, A

n, G

uSp

haer

illo

sp. 1

12

3T

PSp

haer

illo

sp. 2

12

3T

M, A

s, A

g,Sa

, An

Spha

erill

o sp

. 3

123

TG

uSp

haer

illo

sp. 4

12

3T

P, A

nIS

OPO

DA

: LIG

IID

AE

Ligi

a sp

. 1 (a

ff. y

aman

ishi

i) N

unom

ura,

199

012

3T

Ag

ISO

POD

A: O

NIS

CID

AE

Oni

scid

ae sp

. 1

123

TP

ISO

POD

A: P

HIL

OSC

IID

AE

Burm

onis

cus b

icol

or N

unom

ura,

200

112

2T

SPh

ilosc

iidae

sp. 1

12

3T

Ag

Seta

phor

a sp

. 1

123

TA

s, A

g, P

,A

l, G

u, S

a

App

endi

x: C

heck

list o

f the

cru

stac

eans

and

pyc

nogo

nids

of t

he M

aria

nas /

(con

tinue

d)

Taxo

n M

useu

mR

efs

Phot

oN

otes

Hab

Isla

ndSe

taph

ora

sp. 2

12

3T

M, A

g, A

l,Sa

Seta

phor

a sp

. 3

123

TA

gIS

OPO

DA

: PO

RC

ELLI

ON

IDA

EPo

rcel

lioni

des p

ruin

osus

(Bra

ndt,

1833

)12

3T

M, A

sIS

OPO

DA

: SC

YPH

AC

IDA

EAl

loni

scus

sp. 1

(aff

. bon

inen

sis)

Nun

omur

a, 1

984

123

TA

sAl

loni

scus

sp. 2

12

3T

M, A

lAl

loni

scus

saip

anen

sis N

unom

ura,

200

112

2T

SAl

loni

scus

alls

pach

i Nun

omur

a, 2

001

122

TS

ISO

POD

A: T

RA

CH

ELIP

IDA

EN

agur

us sp

. 1

123

TM

, Ag,

Gu

Nag

urus

sp. 2

12

3T

M, A

nN

agur

us sp

. 3

123

TM

Nag

urus

sp. 4

12

3T

Al

ISO

POD

A: T

RIC

HO

NIS

CID

AE

Tric

honi

scid

ae sp

. 1

123

TA

g, A

l, A

n,P

Ref

eren

ces:

Lite

ratu

re re

cord

s lis

ted

by n

umer

ic c

ode,

whi

le id

entif

ier o

f spe

cies

not

pre

viou

sly

docu

men

ted

from

the

Mar

iana

s lis

ted

by a

lpha

betic

code

as f

ollo

ws.

B) W

. Blo

w, C

) T.-Y

. Cha

n, D

) W. D

all,

F) C

. Fra

nsen

, G) B

. Gal

il, K

) R. K

ropp

, L) H

.-C. L

iu, M

) P. M

cLau

ghlin

, N) P

. Ng,

O) J

. Oku

no, P

) G. P

aula

y, R

) P. C

astro

, S) J

. Sta

rmer

, T) M

. Tak

eda.

1) H

umes

199

0, 2

) Hes

sler

& L

onsd

ale

1991

, 3) W

eiss

lead

er e

t al.

1989

,4)

Kom

ai 1

994,

5) C

loud

195

9, 6

) Jus

t 199

8, 7

) Will

iam

s et a

l. 20

00, 8

) Man

ning

et a

l. 19

90, 9

) Ahy

ong

& E

rdm

ann

2003

, 10)

Hol

thui

s 19

53,

11) E

dmon

dson

192

1, 1

2) M

anni

ng 1

978,

13)

Kro

pp &

Dom

ingu

ez 1

990,

14)

Ham

ano

1994

, 15)

Hay

ashi

et a

l. 19

94, 1

6) E

ldre

dge

1994

, 17)

Vic

Cam

acho

, Uni

v. o

f Gua

m, p

ers.

com

m. 2

000,

18)

Ban

ner 1

956,

19)

Kro

pp 1

987,

20)

Ban

ner &

Ban

ner 1

985,

21)

Leb

erer

& C

ai 2

003,

22)

Mar

tin &

Hes

sler

199

0, 2

3) K

ropp

198

3[19

84],

24)

Hol

thui

s 19

81,

25)

Cha

ce 1

997,

26)

Cam

acho

199

7, 2

7) G

rimm

198

3, 2

8) B

est

&D

avid

son

1981

, 29)

Cha

ce &

Bru

ce 1

993,

30)

Bru

ce &

Coo

mbe

s 19

95, 3

1) K

ing

1987

, 32)

Wild

er 1

977,

33)

Mof

fitt 1

983,

34)

Oku

no 1

994,

35) K

ensl

ey 2

003,

36)

Ken

sley

199

6, 3

7) K

ensl

ey 2

001,

38)

Tür

kay

& S

akai

199

5, 3

9) K

ensl

ey &

Hea

rd 1

991,

40)

Asa

kura

et a

l. 19

94, 4

1)A

mes

bury

198

0, 4

2) W

oost

er 1

979,

43)

Woo

ster

198

2, 4

4) A

saku

ra &

Tac

hika

wa

2000

, 45)

Asa

kura

198

7, 4

6) F

ores

t 199

5, 4

7) M

cLau

ghlin

2002

a, 4

8) E

ldre

dge

1976

, 49)

McL

augh

lin 2

002b

, 50)

McL

augh

lin &

Lem

aitre

199

3, 5

1) M

cLau

ghlin

& H

aig

1984

, 52)

McL

augh

lin &

Hai

g19

89, 5

3) D

unn

et a

l. 19

80, 5

4) L

emai

tre 1

998,

55)

Will

iam

s & B

aba

1990

, 56)

Hai

g &

Kro

pp 1

987,

57)

Kro

pp &

Hai

g 19

94, 5

8) K

ropp

198

6,59

) Hes

sler

& M

artin

198

9, 6

0) M

cLay

199

3, 6

1) M

cLay

200

1, 6

2) T

aked

a et

al.

1994

, 63)

McL

ay 1

999,

64)

Eld

redg

e 19

80, 6

5) G

uino

t &

Ric

her

de F

orge

s 19

95, 6

6) N

g &

Wan

g 20

01, 6

7) W

illia

ms

& M

offit

t 199

1, 6

8) G

alil

1997

, 69)

Tan

& N

g 19

95, 7

0) G

uino

t & R

iche

r de

Forg

es 1

985,

71)

Will

iam

s &

Eld

redg

e 19

94, 7

2) N

g &

Tak

eda

1998

, 73)

Tan

& N

g 20

03a,

74)

Tan

& N

g 20

03b,

75)

Ste

vcic

et a

l. 19

88, 7

6)N

g &

Tak

eda

2003

, 77)

Ng

2002

a, 7

8) S

teph

enso

n &

Ree

s 19

67, 7

9) D

icki

nson

197

7, 8

0) W

ee &

Ng

1995

, 81)

Ver

mei

j 197

8, 8

2) N

g &

Ng

2003

, 83)

Ng

2002

b, 8

4) G

alil

& T

aked

a 19

86, 8

5) C

astro

200

3, 8

6) T

aked

a 19

76, 8

7) N

g 20

02c,

88)

Cla

rk &

Gal

il 19

93, 8

9) K

ropp

198

4, 9

0)G

alil

& C

lark

199

0, 9

1) O

dine

tz 1

984,

92)

Ng

& G

uino

t 200

1, 9

3) Q

uoy

& G

aim

ard

1824

, 94)

Ng

2003

, 95)

Ng

et a

l. 19

96, 9

6) T

ürka

y &

Saka

i 199

5, 9

7) K

ropp

198

8a, 9

8) K

ropp

199

0, 9

9) K

ropp

199

4, 1

00)

Kro

pp &

Man

ning

199

6, 1

01)

Kro

pp 1

989,

102

) K

ropp

198

8b, 1

03)

Gry

gier

199

0, 1

04)

New

man

& H

essl

er 1

989,

105

) Fo

ster

199

0, 1

06)

Paul

ay &

Ros

s 20

03,

107)

Sou

thw

ard

et a

l. 19

98,

108)

Fos

ter

&N

ewm

an 1

987,

109

) Jo

nes

2000

, 110

) H

iro 1

937,

111

) A

sam

i & Y

amag

uchi

200

1, 1

12)

Smyt

h 19

86, 1

13)

Smyt

h 19

89, 1

14)

Smyt

h 19

90,

115)

Chi

ld 1

991,

116

) Chi

ld 1

983,

117

) Sto

ck 1

994,

118

) Chi

ld 1

989,

119

) Bel

k 19

73, 1

20) K

ikuc

hi 1

994,

121

) Mor

ino

1994

, 122

) Nun

omur

a20

01, 1

23)

Nun

omur

a 19

94, 1

24)

Ng

2002

d, 1

25)

Ng

et a

l 200

0, 1

26)

Cro

snie

r 20

02, 1

27)

Edm

onds

on 1

959,

128

) K

oh &

Ng

2000

, 129

)Sc

huba

rt &

Ng

2000

, 130

) G

riffin

& T

rant

er 1

986,

131

) N

g &

Cla

rk 2

000,

132

) M

cKen

zie

1967

, 133

) N

g 19

98a,

134

) Fr

anse

n 20

02, 1

35)

Mor

eira

& N

elso

n 19

90, 1

36) N

elso

n &

Kro

pp 1

985,

137

) Yam

aguc

hi 1

975,

138

) Kor

ovch

insk

y 20

01, 1

39) W

atki

ns &

Bel

k 19

75, 1

40) E

l-R

ashi

dy &

Box

shal

l 200

1, 1

41) E

dmon

dson

195

4, 1

42) T

aked

a 19

80, 1

43) C

rosn

ier &

Ng

2003

, 144

) Ng

& M

cLay

200

3.N

otes

:1)

as H

. poo

rei C

roni

n in

Wei

ssle

ader

et a

l. (1

989)

, a n

omen

nud

um (T

. M. C

roni

n pe

rs. c

omm

. 200

1)2)

as L

. hen

ryi C

roni

n in

Wei

ssle

ader

et a

l. (1

989)

, a n

omen

nud

um (T

. M. C

roni

n pe

rs. c

omm

. 200

1)3)

as O

. ros

si C

roni

n in

Wei

ssle

ader

et a

l. (1

989)

, a n

omen

nud

um (T

. M. C

roni

n pe

rs. c

omm

. 200

1)4)

as T

. wei

ssle

ader

ae C

roni

n in

Wei

ssle

ader

et a

l. (1

989)

, a n

omen

nud

um (T

. M. C

roni

n pe

rs. c

omm

. 200

1)5)

as G

onod

acty

lus f

. in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee A

hyon

g &

Erd

man

n (2

003)

for g

ener

ic re

assi

gnm

ent

6) a

s G. c

hira

gra

var.

acut

us in

Edm

onds

on (1

921)

, see

Ahy

ong

& E

rdm

ann.

(200

3) fo

r rei

dent

ifica

tion

7) in

trodu

ced

to G

uam

in fi

sh p

onds

8) O

rigin

al sp

ellin

g as

A. d

iabi

lus i

n B

anne

r (19

56) a

s a re

sult

of ty

pogr

aphi

cal e

rror

, see

Cha

ce (1

988)

9) a

s A. e

dmon

dson

i in

Ban

ner (

1956

), se

e B

anne

r & B

anne

r (19

83) f

or sy

nony

my

10) a

s A. c

. ine

rmis

, A. c

. med

ius,

and

A. c

. pro

babi

lis in

Ban

ner (

1956

), se

e B

anne

r & B

anne

r (19

82) f

or sy

nony

my

11) a

s A. t

uthi

lli in

Ban

ner (

1956

), se

e B

anne

r & B

anne

r (19

82) f

or sy

nony

my

12) a

s A. g

raci

lis v

ar. s

impl

ex in

Ban

ner (

1956

), se

e B

anne

r & B

anne

r (19

82) f

or sy

nony

my

13) a

s Thu

nor i

. in

Ban

ner (

1956

), se

e B

anne

r & B

anne

r (19

82) f

or g

ener

ic re

assi

gnm

ent

14) a

s A. v

entr

osus

in B

anne

r (19

56) a

nd C

loud

(195

9), s

ee B

anne

r & B

anne

r (19

82) f

or sy

nony

my

15) a

s A. p

hyrg

ianu

s in

Ban

ner (

1956

) and

Clo

ud (1

959)

, see

Ban

ner &

Ban

ner (

1982

) for

syno

nym

y16

) as A

. p. b

enga

lens

is in

Ban

ner (

1956

) and

Clo

ud (1

959)

, see

Ban

ner &

Ban

ner (

1982

) for

syno

nym

y17

) mis

spel

led

as A

. pea

rcyi

in H

ayas

hi e

t al.

(199

4)18

) as T

huno

r sp.

1 (n

. sp.

) in

Ban

ner (

1956

), se

e B

anne

r & B

anne

r (19

82) f

or g

ener

ic re

assi

gnm

ent

19) a

s A. c

loud

i in

Ban

ner (

1956

), se

e C

hace

(198

8) fo

r syn

onym

y20

) as A

. dub

ius i

n B

anne

r (19

56),

see

Ban

ner &

Ban

ner (

1973

) for

syno

nym

y

21) a

s A. g

ardi

neri

in C

loud

(195

9) a

nd a

s A. j

ohns

oni i

n B

anne

r (19

56),

see

Ban

ner &

Ban

ner (

1973

) for

syno

nym

y22

) as A

lphe

us p

. in

Clo

ud (1

959)

, see

Ban

ner &

Ban

ner (

1982

) for

gen

eric

reas

sign

men

t23

) as A

. nan

us in

Ban

ner (

1956

) and

Clo

ud (1

959)

, see

Ban

ner &

Ban

ner (

1982

) for

syno

nym

y24

) as A

. c. o

bscu

rus i

n B

anne

r (19

56),

see

Ban

ner &

Ban

ner (

1975

) for

syno

nym

y25

) as A

. cou

tiere

i exi

lipes

in B

anne

r (19

56),

see

Ban

ner &

Ban

ner (

1975

) for

syno

nym

y26

) as D

. n. s

p. in

Clo

ud (1

959)

, des

crib

ed in

Hol

thui

s (19

81)

27) a

s M. l

atim

anus

in C

amac

ho 1

997,

reid

entif

ied

by C

ai &

Ng

(unp

ublis

hed

data

)28

) as P

. ves

tigia

lis in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee C

hace

& B

ruce

(199

3) fo

r gen

eric

reas

sign

men

t29

) as P

eric

limen

es s.

in H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Dur

is &

Bru

ce (1

995)

for g

ener

ic re

assi

gnm

ent

30) a

s Nik

oide

s nan

us in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee C

hace

& B

ruce

(199

3) fo

r syn

onym

y31

) Thi

s is

a th

ird c

olor

form

of A

xiop

sis

on G

uam

that

like

ly re

pres

ents

yet

ano

ther

sib

ling

spec

ies

in th

e A.

ser

ratif

rons

-com

plex

, so

sepa

rate

d by

Paul

ay32

) as A

xius

a. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Ken

sley

(200

3) fo

r gen

eric

reas

sign

men

t33

) err

oneo

usly

reco

rded

in W

oost

er (1

979)

; Woo

ster

’s A

nicu

lus c

orre

spon

ds to

A. u

rsus

, che

cked

and

cor

rect

ed b

y Pa

ulay

34) a

s A. a

nicu

lus i

n W

oost

er (1

979)

, che

cked

and

cor

rect

ed b

y Pa

ulay

35) a

s Cal

cinu

s im

peri

alis

in W

oost

er (1

982)

, see

Pou

pin

(199

7) fo

r rei

dent

ifica

tion

36)

as P

agur

us l

aevi

man

us i

n H

olth

uis

(195

3) a

nd C

loud

(19

59);

see

McL

augh

lin &

Hai

g (1

984)

for

gen

eric

rea

ssig

nmen

t. Id

entif

icat

ion

ofC

loud

’s sp

ecim

ens n

eed

to b

e ve

rifie

d, a

s Pag

urix

us is

a d

iver

se g

enus

in th

e M

aria

nas;

see

com

men

t 40.

McL

augh

lin’s

cur

rent

con

cept

of t

his

spec

ies i

s kno

wn

on G

uam

, see

pho

to a

nd sp

ecim

en v

ouch

ers.

37) a

s Cal

cinu

s sp.

1 in

Woo

ster

(198

2), c

heck

ed a

nd c

orre

cted

by

Paul

ay38

) as C

. gai

mar

di in

all

refs

. cite

d, se

para

ted

out b

y R

ahay

u &

For

est (

1999

); G

uam

mat

eria

l rei

dent

ified

by

McL

augh

lin39

) as O

rtho

pagu

rus h

arm

si in

Woo

ster

(197

9), s

ee M

cLau

ghlin

& L

emai

tre (1

993)

for d

escr

iptio

n of

new

spec

ies

40) a

s Pag

urix

us la

evim

anus

in M

cLau

ghlin

& H

aig

(198

4), r

eide

ntifi

ed b

y M

cLau

ghlin

41) a

s Pa

rapa

guru

s d.

in D

unn

et a

l. (1

980)

, see

Lem

aitre

(198

9) fo

r gen

eric

reas

sign

men

t; as

soci

ated

with

the

deep

wat

er s

hell-

form

ing

anem

one

Styl

obat

es a

eneu

s42

) as C

rypt

odor

mia

can

alic

ulat

a in

Tak

eda

et a

l. (1

994)

, see

McL

ay (1

993)

for s

ynon

ymy

43) a

s Tri

gono

thir

o. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Grif

fin &

Tra

nter

(198

6) fo

r syn

onym

y44

) ide

ntifi

catio

n of

Clo

ud’s

spec

imen

s not

che

cked

; pot

entia

lly T

iari

nia

corn

iger

a45

) as L

itoch

eira

ara

nea

in H

olth

uis (

1953

), C

loud

(195

9), g

ener

ic re

assi

gnm

ent b

y N

g (1

987)

46) a

s Act

aea

h. in

Clo

ud (1

959)

, see

Ser

ène

(198

4) fo

r gen

eric

reas

sign

men

t47

) lik

ely

corr

espo

nds t

o At

erga

tis su

bden

tatu

s rec

orde

d he

re48

) may

be

juve

nile

of A

. gra

nula

tus

49) a

s Act

aea

s. in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee S

erèn

e (1

984)

for g

ener

ic re

assi

gnm

ent

50) a

s Het

erop

ilum

nus i

. in

Hol

thui

s (19

53),

Clo

ud (1

959)

, gen

eric

reas

sign

men

t by

Ng

(198

7)

51) a

s L. a

. in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ubsp

ecifi

c id

entif

icat

ion

by K

ropp

(unp

ublis

hed

data

)52

) as P

ilodi

us h

. in

Take

da e

t al.

(199

4), s

ee C

lark

& G

alil,

199

3 fo

r gen

eric

reas

sign

men

t53

) as C

arpi

lode

s m. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Ser

ène

(198

4) fo

r gen

eric

reas

sign

men

t54

) as C

arpi

lode

s s. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Ser

ène

(198

4) fo

r gen

eric

reas

sign

men

t55

) as C

arpi

lode

s b. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Ser

ène

(198

4) fo

r gen

eric

reas

sign

men

t56

) as C

arpi

lode

s t. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Ser

ène

(198

4) fo

r gen

eric

reas

sign

men

t57

) spe

lling

cor

rect

ed fr

om P

. par

rus i

n C

loud

(195

9)58

) as C

hlor

odop

sis p

. in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee C

lark

& G

alil,

199

3 fo

r gen

eric

reas

sign

men

t59

) as C

hlor

odop

sis m

elan

odac

tyla

in H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Cla

rk &

Gal

il, 1

993

for s

ynon

ymy

and

gene

ric re

assi

gnm

ent

60) a

s Chl

orod

opsi

s p. i

n H

olth

uis (

1953

) and

Clo

ud (1

959)

, see

Cla

rk &

Gal

il, 1

993

for g

ener

ic re

assi

gnm

ent

61) c

onsi

dere

d a

syn.

of P

. gra

nula

tus b

y C

lark

& G

alil

(199

3), b

ut n

ot N

g62

) as A

ctae

a ca

vipe

s in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee S

erèn

e (1

984)

for g

ener

ic re

assi

gnm

ent

63) a

s Act

aea

l. in

Hol

thui

s (19

53) a

nd C

loud

(195

9), s

ee S

erèn

e (1

984)

for g

ener

ic re

assi

gnm

ent

64) a

s The

lphu

sa r.

in Q

uoy

& G

aim

ard

(192

4) a

nd a

s Car

diso

ma

r. in

Tak

eda

et a

l. (1

994)

, see

Ng

& G

uino

t (20

01) f

or g

ener

ic re

assi

gnm

ent

65) T

aked

a et

al.

(199

4) re

cord

P. t

uber

cula

ta, n

ow c

onsi

dere

d a

spec

ies c

ompl

ex o

f P. i

mm

acul

ata

and

P. sq

uam

osa

(Ng

et a

l. 20

00)

66) T

he E

umed

onid

ae h

as b

een

syno

nym

ised

with

the

Pilu

mni

dae

by N

g &

Cla

rk (2

002)

and

now

rega

rded

onl

y as

a su

bfam

ily w

ithin

it.

67) N

g &

Lia

o (2

002)

rece

ntly

reco

gnis

ed th

e Ps

eudo

ziid

ae a

s a d

istin

ct fa

mily

and

form

ally

tran

sfer

red

Eury

oziu

s to

this

taxo

n as

wel

l.68

) as S

aipa

nella

clo

udi i

n M

cKen

zie

(196

7), g

ener

ic re

assi

gnm

ent b

y M

cKen

zie

(197

0)69

) as P

enae

us s.

in E

ldre

dge

(199

4), g

ener

ic re

assi

gnm

ent b

y Pé

rez

Farf

ante

& K

ensl

ey (1

997)

70) a

s Tho

r mal

dive

nsis

in C

loud

(195

9) a

nd H

olth

uis (

1953

), ge

neric

reas

sign

men

t by

Bru

ce (1

997,

199

8)71

) des

crib

e as

Hyp

soph

rys w

illia

msi

, gen

eric

reas

sign

men

t by

Ng

(199

8b)

72)

This

spe

cies

is c

omm

on in

Ple

isto

cene

dep

osits

in A

pra

Har

bor

(Kes

ling

1958

), bu

t was

nev

er s

een

by u

s al

ive.

How

ever

a s

ingl

e, u

nlab

eled

spec

imen

(U

F 28

46)

of t

his

spec

ies

was

fou

nd i

n th

e U

OG

ML

teac

hing

col

lect

ion.

Thi

s sp

ecim

en li

kely

cam

e fr

om G

uam

, but

add

ition

alsp

ecim

ens a

re n

eede

d to

asc

erta

in th

is.

The

crus

tace

an fa

una

of A

pra

harb

or is

still

poo

rly sa

mpl

ed.

73) a

s Pag

urus

latr

o in

Quo

y &

Gai

mar

d (1

824)

, now

in m

onot

ypic

Bir

gus (

Am

esbu

ry 1

980)