02-plantanatomy
TRANSCRIPT
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Plant Anatomy
= the study of plant cell and tissue structure
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Plant cell structure
“Cell” - originally coined by Robert Hooke, in1600’s.
Cell theory:1) All life made of cells2) Cells arise from pre-existing cells3) Cells units of metabolism4) Cells contain complete DNA, hereditary
substance.
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Cell structure
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Ergastic substances
= by-products of metabolism
amyloplasts - starch grains= polymer of alpha-glucose
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2 forms ofglucose
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Ergastic substances
chromoplasts - pigmented bodies, composed ofcarotenoids (e.g., carotene); cause red/orangecoloration.
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carotene
when hydrolysed, yields two vitamin Amolecules:
carotene ---> vitamin A + vitamin A
function in plants is pigmentation, however
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carotene
2 vitamin A
+ 2 H2O
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Ergastic substances
crystals - calcium oxalate (e.g., raphides/druses) or silica - waste/metabolic products or protective
(deter herbivory)
raphides druse
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Ergastic substances
aleurone grains - protein (storage)tannins - phenol derivatives (deter herbivory,
deter infection)fats, oils, waxes - tri-/di-glyerides (storage,
secretion)
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Cell structure
Cell wallPrimary (1˚) cell wall
Formed during growth/expansion of cellsMade up mainly of cellulose
= polymer of beta-glucose unitsPlasmodesmata
Holes in 1˚ cell wall, allow for cell to cellcommunication
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cell wall:cellulosicplasmodesmata
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cellulose:beta formof glucose
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Cellulosic cell wall
made of cellulose microfibrils
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Cell structure
Cell wallSecondary (2˚) cell wall
Formed after growth/expansion of cells,inside original primary cell wall
Made up mainly of lignin
= polymer of phenolic unitshard subtance
Pits - holes in secondary cell wall
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lignin - secondary cell wall
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Secretory cells
laticifers - cells that secrete latex,containing isoprenes (e.g., rubber)
glandular trichomes, oil ducts -secrete oils
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Plant growthMeristems - actively growing regions of plant
Primary meristemsRoot apical meristemsShoot apical meristems
Meristems- cell division- cell elongation- cell differentiation
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root apical
meristem
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SHOOT
leaf primordium
apical
meristem
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cell division
cell elongation
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Cell differentiation
Results in different cell and tissue types
Tissue = 1 or more cell types having acommon origin or function
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Epidermal cell
- outer layer of all plant organs- in land plants, protected by outer cuticle
- cuticle helps to prevent dessication
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cuticle - protective layer on outside - helps prevent water loss
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parenchyma
• - living at maturity• - involved in metabolic reactions
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parenchyma
nucleus
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collenchyma
• - cells living, elongate• - cell walls unevenly thickened,
rich in pectins [pectin - complex polysaccaride,
mainly of galacturonic acid units]
• - functions in structural support(while stem is still elongating)
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collenchymapectic-rich
cell walls
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sclerenchyma
- cells dead at maturity- thick, secondary, lignified cell walls- functions in structural support
2 cell types:fibers - very long, thin, taperedsclereids - variable in shape, mostly isodiametric
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2˚ cell wall
2˚ cell wall
pits
a) fiber b) sclereids
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xylem
• -function: conduction of water and minerals• -consists of:
– parenchyma– fibers– tracheary elements (2 types: tracheids and vessels)
actual conductive cellsdead at maturityjoined end to end, form pipe-like conduitslignified secondary cell walls with pits
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vessel
perforation plate
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phloem
-function: conduction of sugars-consists of:
parenchymafiberssieve elements (2 types: sieve cells or sieve tube members)
- actual sugar-conductive cells- semi-live at maturity (lose nuclei, but have cytoplasm)- primary cell-wall only- have callose-lined pores (making up a sieve plate orsieve area), through which sugar sol. passes
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sieve plate
callose-lined
pore
sieve tube
member
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Plant organs
root - absorptive, anchoring, storage organshoot = stem + associated leaves
stem - conductive, supportive, storage organleaf - photosynthetic organ
bud = immature shoot, gen. arising from leafaxile; ---> lateral branch
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root caproot apical
meristemROOT (l.s.)
1) Protective root cap
2) Absorptive root hairs
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ROOT (c.s.)
3) Give rise to new roots endogenously(from within)
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Casparian StripFunction: forces fluids
from outside throughplasma membrane =selective absorption
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SPOROPHYTIC
SHOOT
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eustele
STEM - has discrete vascular bundles (xylem & phloem)
STEM (of Eudicot)
-bundles in a single ring
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STEM
fiber “bundle” or “cap”
phloem
xylem
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Monocotsstem with many scattered
bundles (atactostele)
vascular bundles
ground meristem
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Leaf anatomy
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stomate:controls
gasexchangeof CO2
& H2O
2
guard cells
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C3 photosynthesis:Normal type
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C4 photosynthesis:PEP (C3) + CO2 --> Malic acid (C4) [Mesophyll]
Malic acid --> CO2 + PEP (C3) [B.S.C.]
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Kranz anatomy (in C4 plants):enlarged Bundle Sheath Cells w/ large chloroplasts
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CAM photosynthesis: CO2 fixed at night(stomates open), released in day (stomates closed)